398 lines
59 KiB
XML
398 lines
59 KiB
XML
<document id="3366B0F0CE1F56D5D054693C9E6BFD3D" ID-CLB-Dataset="45454" ID-DOI="10.11646/zootaxa.3815.4.6" ID-GBIF-Dataset="8c9e9a24-cc17-4abc-a27b-08a81ad9b982" ID-ISSN="1175-5326" ID-Zenodo-Dep="230822" ID-ZooBank="13A3E4D1-A4D4-4C6B-AB43-5268594EA221" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438438669" checkinUser="plazi" docAuthor="Husana, Daniel Edison M., Yamamuro, Masumi & Ng, Peter K. L." docDate="2014" docId="03DC8797FF83FFB4FF188694230F2D47" docLanguage="en" docName="zt03815p574.pdf" docOrigin="Zootaxa 3815 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sundathelphusa danae Husana, Yamamuro & Ng, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="569" masterDocId="FFE5FFEFFF82FFB0FF8F860322142B17" masterDocTitle="Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines" masterLastPageNumber="574" masterPageNumber="565" pageNumber="566" updateTime="1698332556256" updateUser="ExternalLinkService">
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<mods:title id="09B433D9E5CC6EEC416AD2D0B5C9744F">Two new species of freshwater crabs of the genus Sundathelphusa Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) from caves in Luzon, Philippines</mods:title>
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<mods:namePart id="F1B2198610BACBE0AE0EE8972AB9CDAB">Husana, Daniel Edison M.</mods:namePart>
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<mods:namePart id="1DEF2AB31DD9CF9897FA0D8807F64F92">Yamamuro, Masumi</mods:namePart>
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<mods:namePart id="CBACAC5DB486C4CF9C9CDB9D9316C39C">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B4A94181E8FF23A64894F9809F647FEE">Zootaxa</mods:title>
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<mods:part id="F33EDB58B391ECCBB864B31AFEC2921C">
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<mods:date id="787767B0EA7A776A0DAD13FE8CA47D1A">2014</mods:date>
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<mods:detail id="7A67382EB313BA69F6DD5A9A19F0D564" type="volume">
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<mods:number id="0CFADCF4DECFF4C2D92F177CACF09A7A">3815</mods:number>
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<treatment id="03DC8797FF83FFB4FF188694230F2D47" ID-DOI="http://doi.org/10.5281/zenodo.6132780" ID-GBIF-Taxon="119507213" ID-Zenodo-Dep="6132780" LSID="urn:lsid:plazi:treatment:03DC8797FF83FFB4FF188694230F2D47" httpUri="http://treatment.plazi.org/id/03DC8797FF83FFB4FF188694230F2D47" lastPageId="4" lastPageNumber="569" pageId="1" pageNumber="566">
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<subSubSection id="C36F650AFF83FFB1FF18869423772BC3" pageId="1" pageNumber="566" type="nomenclature">
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<paragraph id="8BCA3681FF83FFB1FF18869420152BA7" blockId="1.[151,513,151,213]" box="[151,513,151,176]" pageId="1" pageNumber="566">
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<heading id="D08281EDFF83FFB1FF18869420152BA7" bold="true" box="[151,513,151,176]" fontSize="10" level="2" pageId="1" pageNumber="566" reason="2">
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<emphasis id="B901EA93FF83FFB1FF18869420152BA7" bold="true" box="[151,513,151,176]" pageId="1" pageNumber="566">
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<taxonomicName id="4C754D02FF83FFB1FF18869423B42BA7" ID-CoL="53GBJ" box="[151,416,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
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<emphasis id="B901EA93FF83FFB1FF18869423B42BA7" bold="true" box="[151,416,151,176]" italics="true" pageId="1" pageNumber="566">Sundathelphusa danae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23257E8FF83FFB1FE28869420152BA7" box="[423,513,151,176]" pageId="1" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA3681FF83FFB1FF1886BF23772BC3" blockId="1.[151,513,151,213]" box="[151,355,188,213]" pageId="1" pageNumber="566">
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(
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<figureCitation id="134E2A04FF83FFB1FF1086BF231E2BC3" box="[159,266,188,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1717,1739]" captionStartId-1="2.[151,250,1765,1787]" captionStartId-2="3.[151,250,1639,1661]" captionTargetBox-0="[313,1273,697,1695]" captionTargetBox-1="[163,1417,526,1734]" captionTargetBox-2="[313,1273,181,1631]" captionTargetId-0="figure@1.[313,1273,697,1696]" captionTargetId-1="figure@2.[151,1436,517,1744]" captionTargetId-2="figure@3.[313,1273,181,1632]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." captionText-1="FIGURE 2. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen; B, right second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 5.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." captionText-2="FIGURE 3. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, thoracic sternum and abdomen. Scale bars, A, B = 5.0 mm." httpUri-0="https://zenodo.org/record/230823/files/figure.png" httpUri-1="https://zenodo.org/record/230824/files/figure.png" httpUri-2="https://zenodo.org/record/230825/files/figure.png" pageId="1" pageNumber="566">Figs. 1–3</figureCitation>
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,
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<figureCitation id="134E2A04FF83FFB1FE9986BF234D2BC3" box="[278,345,188,212]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="1" pageNumber="566">7A, B</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36F650AFF83FFB1FF18870720F02993" pageId="1" pageNumber="566" type="materials_examined">
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<paragraph id="8BCA3681FF83FFB1FF188707273F2A72" blockId="1.[151,1436,260,645]" pageId="1" pageNumber="566">
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<emphasis id="B901EA93FF83FFB1FF18870723922A0A" bold="true" box="[151,390,260,285]" pageId="1" pageNumber="566">Material examined.</emphasis>
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||
<typeStatus id="54CE8823FF83FFB1FE1D870723EB2A0A" box="[402,511,260,285]" pageId="1" pageNumber="566" type="holotype">Holotype</typeStatus>
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||
: male (25.9 ×
|
||
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) (
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||
<collectionCode id="ED64AE44FF83FFB1FCB48707219B2A0B" box="[827,911,260,284]" pageId="1" pageNumber="566">NMCR</collectionCode>
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||
39076), Bantakay Cave, Atimonan, Quezon Province, Luzon,
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||
<collectingCountry id="F3627611FF83FFB1FED1872423CB2A57" box="[350,479,295,320]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
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, 14˚
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121˚
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<geoCoordinate id="EE415046FF83FFB1FD28872421182A57" box="[679,780,295,320]" direction="east" orientation="longitude" pageId="1" pageNumber="566" precision="55" value="47.197">47.197E</geoCoordinate>
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||
, coll. D.E.M. Husana,
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||
<date id="FFCB1041FF83FFB1FB82872B26DE2A57" box="[1037,1226,296,320]" pageId="1" pageNumber="566" value="2010-01-13">13 January 2010</date>
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.
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||
<typeStatus id="54CE8823FF83FFB1FB5A872B27522A57" box="[1237,1350,296,320]" pageId="1" pageNumber="566" type="paratype">Paratypes</typeStatus>
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:
|
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<specimenCount id="9D73FD08FF83FFB1FADE872B27882A57" box="[1361,1436,295,320]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
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||
(21.7 ×
|
||
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||
) (
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||
<collectionCode id="ED64AE44FF83FFB1FEE5874F23B62A73" box="[362,418,332,356]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="566">ZRC</collectionCode>
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2013.0274),
|
||
<specimenCount id="9D73FD08FF83FFB1FDB7874F208D2A72" box="[568,665,332,357]" pageId="1" pageNumber="566" type="female">1 female</specimenCount>
|
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(20.4 ×
|
||
<quantity id="4C8D9B64FF83FFB1FD77874F214A2A73" box="[760,862,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="1" pageNumber="566" unit="mm" value="16.8">16.8 mm</quantity>
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||
) (
|
||
<collectionCode id="ED64AE44FF83FFB1FCFC874F21D32A73" box="[883,967,332,356]" pageId="1" pageNumber="566">NMCR</collectionCode>
|
||
39093), same data as
|
||
<typeStatus id="54CE8823FF83FFB1FB4E874F27332A72" box="[1217,1319,332,357]" pageId="1" pageNumber="566" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF83FFB1FF48876C20F02993" blockId="1.[151,1436,260,645]" pageId="1" pageNumber="566">
|
||
<emphasis id="B901EA93FF83FFB1FF48876C23F42A9F" bold="true" box="[199,480,367,392]" pageId="1" pageNumber="566">Comparative material.</emphasis>
|
||
<taxonomicName id="4C754D02FF83FFB1FE7E876C21882A9F" authority="Ng, 2010" authorityName="Ng" authorityYear="2010" box="[497,924,367,392]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="holthuisi">
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||
<emphasis id="B901EA93FF83FFB1FE7E876C21032A9F" box="[497,791,367,392]" italics="true" pageId="1" pageNumber="566">Sundathelphusa holthuisi</emphasis>
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||
<bibRefCitation id="EFE44B70FF83FFB1FCA8877321882A9F" author="Ng" box="[807,924,368,392]" pageId="1" pageNumber="566" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng, 2010</bibRefCitation>
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||
</taxonomicName>
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||
,
|
||
<typeStatus id="54CE8823FF83FFB1FC3F876C26062A9F" box="[944,1042,367,392]" pageId="1" pageNumber="566" type="holotype">holotype</typeStatus>
|
||
male (24.3 ×
|
||
<quantity id="4C8D9B64FF83FFB1FB5A877327502A9F" box="[1237,1348,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="1" pageNumber="566" unit="mm" value="19.4">19.4 mm</quantity>
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) (
|
||
<collectionCode id="ED64AE44FF83FFB1FAEB877327882A9F" box="[1380,1436,368,392]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="566">ZRC</collectionCode>
|
||
1989.2169), Bantakay Cave, Atimonan, Quezon Province, Luzon,
|
||
<collectingCountry id="F3627611FF83FFB1FC3D879726272ABA" box="[946,1075,404,429]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
|
||
, ca. 14˚00N 121˚52E Quezon Province, Luzon,
|
||
<collectingCountry id="F3627611FF83FFB1FEE987B423F32AC7" box="[358,487,439,464]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
|
||
, coll. D.S. Balete,
|
||
<date id="FFCB1041FF83FFB1FD4587BB21782AC7" box="[714,876,440,464]" pageId="1" pageNumber="566" value="1987-05-22">22 May 1987</date>
|
||
;
|
||
<specimenCount id="9D73FD08FF83FFB1FCF487BB21F52AC7" box="[891,993,439,464]" pageId="1" pageNumber="566" type="female">1 female</specimenCount>
|
||
(23.1 ×
|
||
<quantity id="4C8D9B64FF83FFB1FBC287B426A32AC7" box="[1101,1207,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="1" pageNumber="566" unit="mm" value="18.6">18.6 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED64AE44FF83FFB1FB5E87BB271D2AC7" box="[1233,1289,440,464]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="566">ZRC</collectionCode>
|
||
1989.2168), station C-054, National Botanic Gardens, Real, Quezon Province, Luzon,
|
||
<collectingCountry id="F3627611FF83FFB1FB9387DF26892AE2" box="[1052,1181,476,501]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
|
||
, coll. D.S. Balete,
|
||
<date id="FFCB1041FF83FFB1FA0187DF2340290F" pageId="1" pageNumber="566" value="1987-09-09">9 September 1987</date>
|
||
;
|
||
<specimenCount id="9D73FD08FF83FFB1FEEC840323A6290F" box="[355,434,511,536]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
|
||
(18.5 × 15.9) (NSMT-Cr 22316) (
|
||
<collectionCode id="ED64AE44FF83FFB1FCCA8403218D290F" box="[837,921,512,536]" pageId="1" pageNumber="566">NMCR</collectionCode>
|
||
39093), Bantakay Cave, Atimonan, Quezon Province, Luzon,
|
||
<collectingCountry id="F3627611FF83FFB1FEEC842723F0292A" box="[355,484,548,573]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
|
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, 14˚
|
||
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121˚
|
||
<geoCoordinate id="EE415046FF83FFB1FD3F84272102292A" box="[688,790,548,573]" direction="east" orientation="longitude" pageId="1" pageNumber="566" precision="55" value="47.197">47.197E</geoCoordinate>
|
||
, coll. D.E.M. Husana,
|
||
<date id="FFCB1041FF83FFB1FBAF842726F7292B" box="[1056,1251,548,573]" pageId="1" pageNumber="566" value="2010-01-13">13 January 2010</date>
|
||
;
|
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<specimenCount id="9D73FD08FF83FFB1FB7E8427272A292A" box="[1265,1342,548,573]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
|
||
(15.6 ×
|
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<quantity id="4C8D9B64FF83FFB1FF18844B22EB2977" box="[151,255,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="1" pageNumber="566" unit="mm" value="13.7">13.7 mm</quantity>
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) (
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<collectionCode id="ED64AE44FF83FFB1FE96844B23452977" box="[281,337,584,608]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="566">ZRC</collectionCode>
|
||
2013.0275), Nilulubugan Cave, Atimonan, Quezon Province, Luzon,
|
||
<collectingCountry id="F3627611FF83FFB1FBF0844427142977" box="[1151,1280,583,608]" name="Philippines" pageId="1" pageNumber="566">Philippines</collectingCountry>
|
||
, 14˚
|
||
<geoCoordinate id="EE415046FF83FFB1FAA08444278C2977" box="[1327,1432,583,608]" direction="north" orientation="latitude" pageId="1" pageNumber="566" precision="55" value="1.0">01.000N</geoCoordinate>
|
||
, 121˚
|
||
<geoCoordinate id="EE415046FF83FFB1FF4A846F233E2992" box="[197,298,620,645]" direction="east" orientation="longitude" pageId="1" pageNumber="566" precision="55" value="47.775">47.775E</geoCoordinate>
|
||
, coll. D.E.M. Husana,
|
||
<date id="FFCB1041FF83FFB1FDA5846F20F52993" box="[554,737,620,645]" pageId="1" pageNumber="566" value="2009-12">December 2009</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0A6609FF83FFB1FF1880B627602DFD" httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="1" pageNumber="566" targetBox="[313,1273,697,1695]" targetPageId="1">
|
||
<paragraph id="8BCA3681FF83FFB1FF1880B627602DFD" blockId="1.[151,1436,1717,1770]" pageId="1" pageNumber="566">
|
||
<emphasis id="B901EA93FF83FFB1FF1880B623012DDC" bold="true" box="[151,277,1717,1739]" pageId="1" pageNumber="566">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C754D02FF83FFB1FE9180B620132DDC" box="[286,519,1717,1739]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF83FFB1FE9180B620132DDC" box="[286,519,1717,1739]" italics="true" pageId="1" pageNumber="566">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF83FFB1FD9F80B520772DDC" box="[528,611,1718,1739]" pageId="1" pageNumber="566" rank="species">
|
||
<emphasis id="B901EA93FF83FFB1FD9F80B520772DDC" bold="true" box="[528,611,1718,1739]" pageId="1" pageNumber="566">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36F650AFF83FFB4FF488112276C2802" lastPageId="4" lastPageNumber="569" pageId="1" pageNumber="566" type="description">
|
||
<paragraph id="8BCA3681FF83FFB2FF48811226082AC7" blockId="1.[151,1437,1809,2014]" lastBlockId="2.[151,1436,151,464]" lastPageId="2" lastPageNumber="567" pageId="1" pageNumber="566">
|
||
<emphasis id="B901EA93FF83FFB1FF48811223452C3D" bold="true" box="[199,337,1809,1834]" pageId="1" pageNumber="566">Description</emphasis>
|
||
. Carapace quadrate to trapezoidal (
|
||
<figureCitation id="134E2A04FF83FFB1FD6B811221372C3D" box="[740,803,1809,1834]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="1" pageNumber="566">Fig. 1</figureCitation>
|
||
A), widest at anterior quarter, broader than long, dorsoventrally compressed, dorsal surface slightly convex longitudinally, covered with short setae, regions distinct; gastric regions covered with distinct oblique striae; cervical groove prominent; H-shaped gastric groove deep. Epigastric cristae distinct, edges sharp, separated by distinct median furrow; postorbital cristae sharp; epigastric, postorbital cristae confluent; epibranchial teeth, postorbital cristae not confluent, separated by gaps (
|
||
<figureCitation id="134E2A04FF83FFB1FABD81A227622CAD" box="[1330,1398,1953,1978]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="1" pageNumber="566">Fig. 1</figureCitation>
|
||
B). Frontal margin (
|
||
<figureCitation id="134E2A04FF83FFB1FEC081C6238F2CC9" box="[335,411,1989,2014]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="1" pageNumber="566">Figs. 1</figureCitation>
|
||
B, 2C) protruded, sinuous, deflexed. External orbital tooth triangular, produced anteriorly; outer margin concave, granular, longer than inner margin; epibranchial tooth distinct, triangular, well separated from external orbital tooth by deep U-shaped notch, tapering anteriorly; anterolateral margin straight to gently convex, upswept, serrated, clearly demarcated from posterolateral margin; posterolateral margin almost straight to gently concave, with oblique striae converging gradually towards posterior margin of carapace. Frontal median triangle distinct (
|
||
<figureCitation id="134E2A04FF80FFB2FEDB8724238A2A57" box="[340,414,295,320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="2" pageNumber="567">Figs. 1</figureCitation>
|
||
B, 2C), complete, with sharp, protruded dorsal margin, confluent with lateral margins; orbit well demarcated; supraorbital margin smooth; infraorbital margin lined with distinct granules; outer edge reaching, fused with anterolateral margin; suborbital, subbranchial regions covered with scattered oblique striae of nearly uniform size; pterygostomial region with oblique ridges on anterior dorsolateral part. Posterior margin of epistome (
|
||
<figureCitation id="134E2A04FF80FFB2FF1087B422FD2AC7" box="[159,233,439,464]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="2" pageNumber="567">Figs. 1</figureCitation>
|
||
B, 2D) with 3 lobes; median lobe triangular; lateral lobes sinuous, wider.
|
||
</paragraph>
|
||
<caption id="DF0A6609FF80FFB2FF1880E6203A2C61" httpUri="https://zenodo.org/record/230824/files/figure.png" pageId="2" pageNumber="567" targetBox="[163,1417,526,1734]" targetPageId="2">
|
||
<paragraph id="8BCA3681FF80FFB2FF1880E6203A2C61" blockId="2.[151,1436,1765,1910]" pageId="2" pageNumber="567">
|
||
<emphasis id="B901EA93FF80FFB2FF1880E623012DEC" bold="true" box="[151,277,1765,1787]" pageId="2" pageNumber="567">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C754D02FF80FFB2FE9180E620132DEC" box="[286,519,1765,1787]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF80FFB2FE9180E620132DEC" box="[286,519,1765,1787]" italics="true" pageId="2" pageNumber="567">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF80FFB2FD9F80E520772DEC" box="[528,611,1766,1787]" pageId="2" pageNumber="567" rank="species">
|
||
<emphasis id="B901EA93FF80FFB2FD9F80E520772DEC" bold="true" box="[528,611,1766,1787]" pageId="2" pageNumber="567">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen; B, right second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G1, ventral view; F, left G1, dorsal view; G, left G2; H, terminal segment of left G1, ventral view; I, terminal segment of left G1, dorsal view; J, flagellum of left G2. Scale bars, A, B = 5.0 mm; C, D = 2.0 mm; E‒G = 1.0 mm; H‒J = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0A6609FF81FFB3FF18806423142DAD" httpUri="https://zenodo.org/record/230825/files/figure.png" pageId="3" pageNumber="568" targetBox="[313,1273,181,1631]" targetPageId="3">
|
||
<paragraph id="8BCA3681FF81FFB3FF18806423142DAD" blockId="3.[151,1436,1639,1722]" pageId="3" pageNumber="568">
|
||
<emphasis id="B901EA93FF81FFB3FF18806423012D6B" bold="true" box="[151,277,1639,1661]" pageId="3" pageNumber="568">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C754D02FF81FFB3FE91806420132D6A" box="[286,519,1639,1661]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF81FFB3FE91806420132D6A" box="[286,519,1639,1661]" italics="true" pageId="3" pageNumber="568">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901EA93FF81FFB3FD9F806420772D6B" bold="true" box="[528,611,1639,1660]" pageId="3" pageNumber="568">
|
||
<taxonomicNameLabel id="A23257E8FF81FFB3FD9F806420772D6B" box="[528,611,1639,1660]" pageId="3" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, thoracic sternum and abdomen. Scale bars, A, B = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA3681FF81FFB3FF4880E223152C9F" blockId="3.[151,1437,1760,2038]" pageId="3" pageNumber="568">
|
||
Eyes well developed (
|
||
<figureCitation id="134E2A04FF81FFB3FE3080E323EB2DEE" box="[447,511,1760,1785]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="3" pageNumber="568">Fig. 1</figureCitation>
|
||
B), well pigmented, occupying almost entire orbit. Third maxilliped ischium (
|
||
<figureCitation id="134E2A04FF81FFB3FAE080E322B02C0A" captionStart="FIGURE 3" captionStartId="3.[151,250,1639,1661]" captionTargetBox="[313,1273,181,1631]" captionTargetId="figure@3.[313,1273,181,1632]" captionTargetPageId="3" captionText="FIGURE 3. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, buccal field and thoracic sternum; B, thoracic sternum and abdomen. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230825/files/figure.png" pageId="3" pageNumber="568">Fig. 3</figureCitation>
|
||
A) rectangular, much longer than broad, with distinct oblique submedian sulcus closer to mesial margin, surfaces covered with scattered short setae; merus quadrate, antero-external angle convex, upswept; tip of exopod not reaching midpoint of outer margin of merus, with short flagellum barely reaching edge of upswept mesial margin of merus.
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF81FFB3FF488196279F2CE1" blockId="3.[151,1437,1760,2038]" pageId="3" pageNumber="568">Chelipeds not noticeably elongated, subequal, stronger in males; margins of merus serrated, dorsal margin with small subdistal tooth; carpus armed with strong distal sharply pointed inner tooth, dorso-ventrally flattened; palm with smooth outer surface; fingers robust, cutting edges armed with numerous medium-sized to large sharp teeth.</paragraph>
|
||
<paragraph id="8BCA3681FF86FFB4FF48869420912A0A" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
Ambulatory legs long (
|
||
<figureCitation id="134E2A04FF86FFB4FE43869420022BA7" box="[460,534,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
|
||
A, 2B), slender, second leg longest, anterior margins of meri distinctly serrated, without subdistal tooth or spine, posterior margins smooth on all legs; second, third meri slightly shorter than carapace length; carpi short, with longitudinal submedian ridge on dorsal surface except for fourth leg, widened distally; propodi, dactyli subequal in length.
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF86FFB4FF48872422E82992" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
Male abdominal cavity reaching imaginary level demarcated by proximal quarter of coxae of chelipeds. Male abdomen (
|
||
<figureCitation id="134E2A04FF86FFB4FE82874F234F2A73" box="[269,347,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1765,1787]" captionTargetBox="[163,1417,526,1734]" captionTargetId="figure@2.[151,1436,517,1744]" captionTargetPageId="2" captionText="FIGURE 2. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen; B, right second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 5.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230824/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
|
||
A, 3B) narrow, T-shaped; somite 1 short, somite 6 longer than broad; proximal, distal margins of somites 3–5 sinuous; lateral margins of somite 2 convex; lateral margins of somite 3 convex; lateral margins of somite 4 straight, narrows gradually to distal end; somites 3–5 narrow gradually; lateral margins of somite 5 concave, proximally wide distally narrow; lateral margins of somite 6 longer than broad, straight; telson subtriangular, longer than broad, lateral margin slightly concave proximally, rounded distally. G1 slender (
|
||
<figureCitation id="134E2A04FF86FFB4FAB887DF27912AE3" box="[1335,1413,476,501]" captionStart="FIGURE 2" captionStartId="2.[151,250,1765,1787]" captionTargetBox="[163,1417,526,1734]" captionTargetId="figure@2.[151,1436,517,1744]" captionTargetPageId="2" captionText="FIGURE 2. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen; B, right second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 5.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230824/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
|
||
E, F, H, I), subterminal segment gradually tapering towards terminal segment, proximal two-thirds straight, distal onethird slightly bent outward, segments separated by distinct suture; terminal segment straight, tapering, cylindrical, obliquely bent outwards from suture. G2 (
|
||
<figureCitation id="134E2A04FF86FFB4FDE3844420AC2977" box="[620,696,583,608]" captionStart="FIGURE 2" captionStartId="2.[151,250,1765,1787]" captionTargetBox="[163,1417,526,1734]" captionTargetId="figure@2.[151,1436,517,1744]" captionTargetPageId="2" captionText="FIGURE 2. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines. A, male abdomen; B, right second ambulatory leg, ventral view; C, frontal margins, frontal view; D, posterior margin of epistome; E, left G 1, ventral view; F, left G 1, dorsal view; G, left G 2; H, terminal segment of left G 1, ventral view; I, terminal segment of left G 1, dorsal view; J, flagellum of left G 2. Scale bars, A, B = 5.0 mm; C, D = 2.0 mm; E ‒ G = 1.0 mm; H ‒ J = 0.5 mm." httpUri="https://zenodo.org/record/230824/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
|
||
G, J) longer than G1, flagellum long, about half the length of basal segment.
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF86FFB4FF48848C21A529E7" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
<emphasis id="B901EA93FF86FFB4FF48848C230A29BF" bold="true" box="[199,286,655,680]" pageId="4" pageNumber="569">Female</emphasis>
|
||
. Female chelipeds subequal, neither inflated; abdomen rounded, covering entire thoracic sternum, all somites and telson freely articulating; telson broadly triangular with convex lateral margins (
|
||
<figureCitation id="134E2A04FF86FFB4FB5884B7272129DB" box="[1239,1333,692,717]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="4" pageNumber="569">Fig. 7A</figureCitation>
|
||
). Vulvae large, round, without operculum; on median part of somite 6 (
|
||
<figureCitation id="134E2A04FF86FFB4FCC484D421B029E7" box="[843,932,727,752]" captionStart="FIGURE 7. A, B" captionStartId="8.[151,250,1219,1241]" captionTargetBox="[151,1436,303,1196]" captionTargetId="figure@8.[151,1436,303,1196]" captionTargetPageId="8" captionText="FIGURE 7. A, B, Sundathelphusa danae sp. nov., female (20.4 × 16.8 mm) (NMCR 39093), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines; C, D, Sundathelphusa vienae sp. nov., paratype female (19.8 × 16.5 mm) (NSMT-Cr 22317), Nilulubugan Cave, Atimonan, Quezon Province, Luzon, Philippines. A, C, female abdomens; B, D, thoracic sternums showing vulvae." httpUri="https://zenodo.org/record/230829/files/figure.png" pageId="4" pageNumber="569">Fig. 7B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF86FFB4FF4884FF276C2802" blockId="4.[151,1437,151,1616]" box="[199,1400,764,789]" pageId="4" pageNumber="569">
|
||
<emphasis id="B901EA93FF86FFB4FF4884FF235C2802" bold="true" box="[199,328,764,789]" pageId="4" pageNumber="569">Coloration</emphasis>
|
||
. Dorsal surface of carapace is purple to maroon in life, cheliped is yellowish to reddish-orange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F650AFF86FFB4FF48851C231F284A" pageId="4" pageNumber="569" type="etymology">
|
||
<paragraph id="8BCA3681FF86FFB4FF48851C231F284A" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
<emphasis id="B901EA93FF86FFB4FF48851C2352282F" bold="true" box="[199,326,799,824]" pageId="4" pageNumber="569">Etymology</emphasis>
|
||
. Named after a daughter of the first author, Dana Vi Husana, who accompanied him in some of his fieldwork.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F650AFF86FFB4FF48856420C32E27" pageId="4" pageNumber="569" type="discussion">
|
||
<paragraph id="8BCA3681FF86FFB4FF48856420C32E27" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
<emphasis id="B901EA93FF86FFB4FF48856423202897" bold="true" box="[199,308,871,896]" pageId="4" pageNumber="569">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C754D02FF86FFB4FECB856420512897" box="[324,581,871,896]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF86FFB4FECB856420512897" box="[324,581,871,896]" italics="true" pageId="4" pageNumber="569">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901EA93FF86FFB4FDC2856420BD2897" bold="true" box="[589,681,871,896]" pageId="4" pageNumber="569">
|
||
<taxonomicNameLabel id="A23257E8FF86FFB4FDC2856420BD2897" box="[589,681,871,896]" pageId="4" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was collected from Bantakay Cave, Atimonan, Quezon Province, Luzon, the
|
||
<typeStatus id="54CE8823FF86FFB4FE95858E235E28B2" box="[282,330,909,933]" pageId="4" pageNumber="569">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C754D02FF86FFB4FE5D858E20DE28B3" authority="Ng, 2010" authorityName="Ng" authorityYear="2010" box="[466,714,908,933]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FE5D858E204628B2" box="[466,594,908,933]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
<bibRefCitation id="EFE44B70FF86FFB4FDD4858E20DE28B3" author="Ng" box="[603,714,908,933]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
. Although
|
||
<taxonomicName id="4C754D02FF86FFB4FCC6858F265E28B2" box="[841,1098,908,933]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF86FFB4FCC6858F265E28B2" box="[841,1098,908,933]" italics="true" pageId="4" pageNumber="569">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF86FFB4FBDC858F26BB28B2" box="[1107,1199,908,933]" pageId="4" pageNumber="569" rank="species">
|
||
<emphasis id="B901EA93FF86FFB4FBDC858F26BB28B2" bold="true" box="[1107,1199,908,933]" pageId="4" pageNumber="569">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is sympatric with
|
||
<taxonomicName id="4C754D02FF86FFB4FA08858E22ED28DF" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FA08858E22ED28DF" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
</taxonomicName>
|
||
(and the two species were collected together, see comparative material), and both possess a relatively slender G1 with a simple cylindrical terminal segment, they differ markedly in several other aspects.
|
||
<taxonomicName id="4C754D02FF86FFB4FF1885F4238F2F07" box="[151,411,1015,1040]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF86FFB4FF1885F4238F2F07" box="[151,411,1015,1040]" italics="true" pageId="4" pageNumber="569">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF86FFB4FE2885F420122F07" box="[423,518,1015,1040]" pageId="4" pageNumber="569" rank="species">
|
||
<emphasis id="B901EA93FF86FFB4FE2885F420122F07" bold="true" box="[423,518,1015,1040]" pageId="4" pageNumber="569">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can easily be distinguished from
|
||
<taxonomicName id="4C754D02FF86FFB4FC2E85FB26302F07" box="[929,1060,1015,1040]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FC2E85FB26302F07" box="[929,1060,1015,1040]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
</taxonomicName>
|
||
as follows: 1) the anterolateral margins are prominently more convex (
|
||
<figureCitation id="134E2A04FF86FFB4FDD8821F20832F23" box="[599,663,1052,1077]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
|
||
A) (margins gently convex in
|
||
<taxonomicName id="4C754D02FF86FFB4FC7E821E26F22F23" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[1009,1254,1052,1077]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FC7E821E26642F22" box="[1009,1136,1052,1077]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FBF0821E26F22F23" author="Ng" box="[1151,1254,1052,1077]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 2B); 2) the branchial regions are proportionately more inflated (
|
||
<figureCitation id="134E2A04FF86FFB4FD7E823C21272F4F" box="[753,819,1087,1112]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
|
||
B) (only gently swollen in
|
||
<taxonomicName id="4C754D02FF86FFB4FBE08243277D2F4F" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[1135,1385,1087,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FBE0824326E42F4F" box="[1135,1264,1087,1112]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FA8E8243277D2F4F" author="Ng" box="[1281,1385,1088,1112]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 2C); 3) the median triangular lobe on the posterior margin of the epistome is sharper and relatively broader (
|
||
<figureCitation id="134E2A04FF86FFB4FAE0826722B62FB7" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
|
||
B) (wider and the tip rounded in
|
||
<taxonomicName id="4C754D02FF86FFB4FD95828B21192FB7" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[538,781,1159,1185]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FD95828B208C2FB7" box="[538,664,1159,1184]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FD28828A21192FB7" author="Ng" box="[679,781,1160,1185]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 3B); 4) the frontal median triangle is complete (
|
||
<figureCitation id="134E2A04FF86FFB4FAEB828422B62FD3" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
|
||
B, 2C) (incomplete in
|
||
<taxonomicName id="4C754D02FF86FFB4FE3682AE20C82FD3" authority="Ng 2010: 3" authorityName="Ng" authorityPageNumber="3" authorityYear="2010" box="[441,732,1196,1221]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FE3682AE20292FD2" box="[441,573,1196,1221]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FDDD82AE20A92FD3" author="Ng" box="[594,701,1196,1221]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: 3
|
||
</taxonomicName>
|
||
B); 5) the ambulatory legs are relatively more slender and elongated, especially with regard to the merus and propodus (
|
||
<figureCitation id="134E2A04FF86FFB4FCC682CC21862FFF" box="[841,914,1231,1256]" captionStart="FIGURE 1" captionStartId="1.[151,250,1717,1739]" captionTargetBox="[313,1273,697,1695]" captionTargetId="figure@1.[313,1273,697,1696]" captionTargetPageId="1" captionText="FIGURE 1. Sundathelphusa danae sp. nov., holotype male (25.9 × 19.8 mm) (NMCR 39076), Bantakay Cave, Atimonan, Quezon Province, Luzon, Philippines: A, habitus, dorsal view; B, cephalothorax, anterior view. Scale bars, A, B = 5.0 mm." httpUri="https://zenodo.org/record/230823/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
|
||
A, 2B) (merus and propodus relatively shorter and stouter in
|
||
<taxonomicName id="4C754D02FF86FFB4FEBA82F620322E1B" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[309,550,1268,1293]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FEBA82F623A72E1A" box="[309,435,1268,1293]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FE4E82F620322E1B" author="Ng" box="[449,550,1268,1293]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 2B); and 6) the G1 terminal segment is more strongly bent (Fig. E, F, H, I) (gently curved in
|
||
<taxonomicName id="4C754D02FF86FFB4FED2831B205A2E27" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[349,590,1303,1329]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FED2831B23CF2E27" box="[349,475,1303,1328]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FE66831A205A2E27" author="Ng" box="[489,590,1304,1329]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: fig. 4J–M).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F650AFF86FFB4FF48833F230F2D47" pageId="4" pageNumber="569" type="reference_group">
|
||
<paragraph id="8BCA3681FF86FFB4FF48833F22FD2EF3" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
<taxonomicName id="4C754D02FF86FFB4FF48833F23D32E42" ID-CoL="53GBJ" box="[199,455,1340,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="danae" status="sp. nov.">
|
||
<emphasis id="B901EA93FF86FFB4FF48833F23D32E42" box="[199,455,1340,1365]" italics="true" pageId="4" pageNumber="569">Sundathelphusa danae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257E8FF86FFB4FE41833F203C2E42" box="[462,552,1340,1365]" pageId="4" pageNumber="569" rank="species">
|
||
<emphasis id="B901EA93FF86FFB4FE41833F203C2E42" bold="true" box="[462,552,1340,1365]" pageId="4" pageNumber="569">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can immediately be separated from
|
||
<taxonomicName id="4C754D02FF86FFB4FC49833E269A2E43" ID-CoL="93759" authority="Ng, 1991" authorityName="Ng" authorityYear="1991" box="[966,1166,1340,1365]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="celer">
|
||
<emphasis id="B901EA93FF86FFB4FC49833E260F2E42" box="[966,1051,1340,1365]" italics="true" pageId="4" pageNumber="569">S. celer</emphasis>
|
||
Ng, 1991
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C754D02FF86FFB4FB44833E22CC2E6F" ID-CoL="53GBR" authority="Balss, 1937" authorityName="Balss" authorityYear="1937" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B901EA93FF86FFB4FB44833E27532E42" box="[1227,1351,1340,1365]" italics="true" pageId="4" pageNumber="569">S. longipes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B70FF86FFB4FAD8833F22C42E6F" author="Balss" pageId="4" pageNumber="569" refString="Balss, H. (1937) Potamoniden (Dekapoda Brachyura) der Philippinen und des Malayischen Archipels. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34, 143 - 197. http: // dx. doi. org / 10.1002 / iroh. 19370340111" type="journal article" year="1937">Balss, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(both from Luzon) by its more strongly convex anterolateral margins (cf
|
||
<bibRefCitation id="EFE44B70FF86FFB4FBFE836226CA2E6F" author="Ng" box="[1137,1246,1376,1401]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: figs. 1A, 2A), proportionately longer ambulatory legs (cf.
|
||
<bibRefCitation id="EFE44B70FF86FFB4FD04838620E72E8B" author="Ng" box="[651,755,1412,1437]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
: figs. 1E, 2A) and more slender G1 which has the terminal segment slender and cylindrical (relatively stouter and shorter in
|
||
<taxonomicName id="4C754D02FF86FFB4FC0C83AB21CD2ED7" ID-CoL="93759" box="[899,985,1447,1472]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="celer">
|
||
<emphasis id="B901EA93FF86FFB4FC0C83AB21CD2ED7" box="[899,985,1447,1472]" italics="true" pageId="4" pageNumber="569">S. celer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754D02FF86FFB4FB9B83AB271E2ED7" ID-CoL="53GBR" authority="Ng 2010" authorityName="Ng" authorityYear="2010" box="[1044,1290,1447,1473]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B901EA93FF86FFB4FB9B83AB26862ED7" box="[1044,1170,1447,1472]" italics="true" pageId="4" pageNumber="569">S. longipes</emphasis>
|
||
,
|
||
<bibRefCitation id="EFE44B70FF86FFB4FB2D83AA271E2ED7" author="Ng" box="[1186,1290,1448,1473]" pageId="4" pageNumber="569" refString="Ng, P. K. L. (2010) On the identity of Para - Bary - thelphusa grapsoides longipes Balss, 1937, with description of a new species from the Philippines (Brachyura, Gecarcinucidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Fransen, C. H. J M., S. De Grave & P. K. L. Ng (Eds.). Crustaceana Monographs, 14, 561 - 571. http: // dx. doi. org / 10.1163 / 9789047427759 _ 039" type="journal article" year="2010">Ng 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
: figs. 1H–K, 4E–H).
|
||
</paragraph>
|
||
<paragraph id="8BCA3681FF86FFB4FF4883EC230F2D47" blockId="4.[151,1437,151,1616]" pageId="4" pageNumber="569">
|
||
The spelling of the name of the
|
||
<typeStatus id="54CE8823FF86FFB4FDBB83F220702D1E" box="[564,612,1521,1545]" pageId="4" pageNumber="569">type</typeStatus>
|
||
locality for
|
||
<taxonomicName id="4C754D02FF86FFB4FD7E83F321642D1F" ID-CoL="53GBN" box="[753,880,1519,1544]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="holthuisi">
|
||
<emphasis id="B901EA93FF86FFB4FD7E83F321642D1F" box="[753,880,1519,1544]" italics="true" pageId="4" pageNumber="569">S. holthuisi</emphasis>
|
||
</taxonomicName>
|
||
– “Bautakay Cave, Station
|
||
<taxonomicName id="4C754D02FF86FFB4FB2283EC27862D1F" ID-CoL="8XK99" authority="Atomonau" authorityName="Atomonau" box="[1197,1426,1519,1544]" class="Insecta" family="Staphylinidae" genus="Catalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="genus">Catalina, Atomonau</taxonomicName>
|
||
” (see also Ng 1991: 15 as
|
||
<taxonomicName id="4C754D02FF86FFB4FE5B8017210E2D3A" box="[468,794,1556,1581]" class="Malacostraca" family="Gecarcinucidae" genus="Archipelothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B901EA93FF86FFB4FE5B8017210E2D3A" box="[468,794,1556,1581]" italics="true" pageId="4" pageNumber="569">Archipelothelphusa longipes</emphasis>
|
||
</taxonomicName>
|
||
) is incorrect. It should be “Bantakay, Santa
|
||
<taxonomicName id="4C754D02FF86FFB4FAB7801723192D47" ID-CoL="8XK99" authority="Atimonan" authorityName="Atimonan" class="Insecta" family="Staphylinidae" genus="Catalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="569" phylum="Arthropoda" rank="genus">Catalina, Atimonan</taxonomicName>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |