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<document id="0958029FD7A16B43F0AC3398D0724245" ID-CLB-Dataset="25362" ID-DOI="10.11646/zootaxa.4674.5.6" ID-GBIF-Dataset="534da4e0-2064-4098-93c2-1f614aef9d2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3465101" ID-ZooBank="250CE8E7-B7B5-4503-873C-5E79827F6D3C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569830604473" checkinUser="plazi" docAuthor="Shear, William A. &amp; Crawford, Rodney L." docDate="2019" docId="03D17B73FFDA6D77D9CD688886A84872" docLanguage="en" docName="zootaxa.4674.5.6.pdf" docOrigin="Zootaxa 4674 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Taiyutyla caseophila Shear &amp; Crawford 2019, new species" docType="treatment" docVersion="11" lastPageNumber="574" masterDocId="FFE8030BFFDB6D74D95A6D1684364C0D" masterDocTitle="Cave millipedes of the United States. XVI. Two new species from Oregon Caves National Monument, Oregon (Chordeumatida, Conotylidae and Caseyidae)" masterLastPageNumber="580" masterPageNumber="571" pageNumber="572" updateTime="1698756535735" updateUser="ExternalLinkService">
<mods:mods id="0DDEB0962E2E3DD052D6168FE7E5C947" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0F4DF024C7769F536CF38DC9648CACB2">
<mods:title id="D20CDFD9BB0B8C587C3C5D05E815F235">Cave millipedes of the United States. XVI. Two new species from Oregon Caves National Monument, Oregon (Chordeumatida, Conotylidae and Caseyidae)</mods:title>
</mods:titleInfo>
<mods:name id="908762EEFC86D8B1C264B971F765B6C6" type="personal">
<mods:role id="DC07A37E69AF334C7A52A64DFE37E47D">
<mods:roleTerm id="68F761CFD72F826E3C1037800F2B3F18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="257B4F98875090710268D428D83EDF61">Shear, William A.</mods:namePart>
</mods:name>
<mods:name id="3A67FE44B969235619EE1F477AFF3C04" type="personal">
<mods:role id="BFF6F78A6BDEC3ECBE6F0111D70C6C91">
<mods:roleTerm id="00DF6FF2FDE6AB4D10FC67735C9869A6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F98A30A197C87EE5E0EB8E2AC0DAB97">Crawford, Rodney L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D2E5517031230D8F607DA08A1F14AEF9">text</mods:typeOfResource>
<mods:relatedItem id="B81A08ECE86D8DC04B42CF69D68494ED" type="host">
<mods:titleInfo id="6A8CD74BF16B28CFE94189EC10094B1C">
<mods:title id="292DC8A8447181400F955E15559265AF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="54571A551AD43E43F40F2A598FC01BBC">
<mods:date id="77255BA8C09813B2B6F1B19723F0A4F7">2019</mods:date>
<mods:detail id="E0F8CE352427788D941354FA0DA58D9D" type="pubDate">
<mods:number id="0CFC3760068CAF05BCF53091525673DC">2019-09-27</mods:number>
</mods:detail>
<mods:detail id="36D4FD3BD1C84389D26BC3F145F3084B" type="volume">
<mods:number id="84ABD91F6648B26C51F18442D5A21912">4674</mods:number>
</mods:detail>
<mods:detail id="206CA278A38576020FC831677589E5EF" type="issue">
<mods:number id="72D2B4AC0B4E5969046D8C3F5DA6F931">5</mods:number>
</mods:detail>
<mods:extent id="748C3B1DA2648E484918182478FE4848" unit="page">
<mods:start id="B2B93EA61C43AE351C3FE626C9B66C9F">571</mods:start>
<mods:end id="355AA81A735C82A92C5504A80BDB5BC3">580</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EFA28BA24A0B288B3443125B0FDF241E">journal article</mods:classification>
<mods:identifier id="C32AF901D40CE6D52A579FC79D1AD9BF" type="CLB-Dataset">25362</mods:identifier>
<mods:identifier id="68703D179351D9DADF987ECAABDB0CFE" type="DOI">10.11646/zootaxa.4674.5.6</mods:identifier>
<mods:identifier id="C036F932E65B671DF2C7DBCE6DCBDAEF" type="GBIF-Dataset">534da4e0-2064-4098-93c2-1f614aef9d2e</mods:identifier>
<mods:identifier id="2AA1B504418B47DB677C00A4EAB5E5F4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="82239845F013DA9D071EED26EEF54AE4" type="Zenodo-Dep">3465101</mods:identifier>
<mods:identifier id="419FC96164695689EEB5862EA81EF593" type="ZooBank">250CE8E7-B7B5-4503-873C-5E79827F6D3C</mods:identifier>
</mods:mods>
<treatment id="03D17B73FFDA6D77D9CD688886A84872" ID-DOI="http://doi.org/10.5281/zenodo.5940775" ID-GBIF-Taxon="159609129" ID-Zenodo-Dep="5940775" LSID="urn:lsid:plazi:treatment:03D17B73FFDA6D77D9CD688886A84872" httpUri="http://treatment.plazi.org/id/03D17B73FFDA6D77D9CD688886A84872" lastPageId="3" lastPageNumber="574" pageId="1" pageNumber="572">
<subSubSection id="C36299EEFFDA6D75D9CD6888871649B5" box="[151,800,1438,1465]" pageId="1" pageNumber="572" type="nomenclature">
<paragraph id="8BC7CA65FFDA6D75D9CD6888871649B5" blockId="1.[151,800,1438,1500]" box="[151,800,1438,1465]" pageId="1" pageNumber="572">
<heading id="D08F7D09FFDA6D75D9CD6888871649B5" bold="true" box="[151,800,1438,1465]" fontSize="11" level="1" pageId="1" pageNumber="572" reason="1">
<emphasis id="B90C1677FFDA6D75D9CD6888871649B5" bold="true" box="[151,800,1438,1465]" pageId="1" pageNumber="572">
<taxonomicName id="4C78B1E6FFDA6D75D9CD688886B449B5" authority="Shear &amp; Crawford" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[151,642,1438,1465]" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="caseophila" status="sp. nov.">
<emphasis id="B90C1677FFDA6D75D9CD688885B949B4" bold="true" box="[151,399,1438,1465]" italics="true" pageId="1" pageNumber="572">Taiyutyla caseophila</emphasis>
Shear &amp; Crawford
</taxonomicName>
,
<taxonomicNameLabel id="A23FAB0CFFDA6D75DBCA6889871649B5" box="[656,800,1438,1465]" pageId="1" pageNumber="572" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDA6D75D9CD68D484C949D1" box="[151,255,1473,1500]" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC7CA65FFDA6D75D9CD68D484C949D1" blockId="1.[151,800,1438,1500]" box="[151,255,1473,1500]" pageId="1" pageNumber="572">
<figureCitation id="1343D6E0FFDA6D75D9CD68D484C949D1" box="[151,255,1473,1500]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4, 5" captionStartId-0="2.[151,264,1925,1950]" captionStartId-1="3.[151,264,1811,1836]" captionTargetBox-0="[151,1432,167,1905]" captionTargetBox-1="[151,1436,1165,1792]" captionTargetId-0="figure@2.[151,1432,166,1905]" captionTargetId-1="figure@3.[151,1436,1165,1792]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–3. Taiyutyla caseophila, new species. Fig. 1. Anterior and posterior gonopods, anterior view; agp, anterior gonopod; pf9, prefemur of posterior gonopod; s9, sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view; agp, anterior gonopod; cx9, coxite of posterior gonopod; T, limb of T-shaped process. Fig. 3. Left leg 7, anterior view." captionText-1="FIGURES 4, 5. Taiyutyla caseophila, new species. Fig. 4. Tip of posterior gonopod coxite, posterior view; f, fimbriate region; h, hook-like process; T, limb of T-shaped process. Fig. 5. Pits on femoral knob of leg 7, anterior view." figureDoi-0="http://doi.org/10.5281/zenodo.3467968" figureDoi-1="http://doi.org/10.5281/zenodo.3467970" httpUri-0="https://zenodo.org/record/3467968/files/figure.png" httpUri-1="https://zenodo.org/record/3467970/files/figure.png" pageId="1" pageNumber="572">Figs. 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDA6D75D9CD6B1F85F24AD4" pageId="1" pageNumber="572" type="materials_examined">
<paragraph id="8BC7CA65FFDA6D75D9CD6B1F85F24AD4" blockId="1.[151,1437,1545,2005]" pageId="1" pageNumber="572">
<emphasis id="B90C1677FFDA6D75D9CD6B1F854E4A29" bold="true" box="[151,376,1545,1572]" pageId="1" pageNumber="572">Material examined</emphasis>
:
<typeStatus id="54C374C7FFDA6D75D8DF6B1D858B4A28" box="[389,445,1547,1573]" pageId="1" pageNumber="572">Type</typeStatus>
<specimenCount id="9D7E01ECFFDA6D75D8996B1C860D4A29" box="[451,571,1546,1572]" pageId="1" pageNumber="572" type="generic">specimens</specimenCount>
.
<materialsCitation id="3B10C038FFDA6D75DB1E6B1C80894A44" ID-GBIF-Occurrence="2423596447" collectionCode="UWBM" county="Josephine Co." latitude="42.0974" location="Oregon" municipality="Caves National Monument" pageId="1" pageNumber="572" specimenCount="14" specimenCount-female="1" specimenCount-male="13" stateProvince="Oregon" typeStatus="holotype">
<specimenCount id="9D7E01ECFFDA6D75DB1E6B1C86484A29" box="[580,638,1546,1572]" pageId="1" pageNumber="572" type="male">Male</specimenCount>
<typeStatus id="54C374C7FFDA6D75DBD96B1C86D34A29" box="[643,741,1546,1572]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED6952A0FFDA6D75DBA96B1C87674A29" box="[755,849,1546,1572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="1" pageNumber="572" type="Museum">UWBM</collectionCode>
) and
<specimenCount id="9D7E01ECFFDA6D75DAD66B1C87D54A29" box="[908,995,1546,1572]" pageId="1" pageNumber="572" type="male">12 male</specimenCount>
and five
<specimenCount id="9D7E01ECFFDA6D75DD136B1C80A34A29" box="[1097,1173,1546,1572]" pageId="1" pageNumber="572" type="female">female</specimenCount>
paratypes from Oregon Cave,
<collectingRegion id="49BC0487FFDA6D75D9856B3885054A45" box="[223,307,1582,1608]" country="United States of America" name="Oregon" pageId="1" pageNumber="572">Oregon</collectingRegion>
<collectingMunicipality id="6BA3501FFFDA6D75D8616B38865A4A44" box="[315,620,1582,1609]" county="Josephine Co." latitude="42.0974" longLatPrecision="6" longitude="-123.4099" municipality="Cave" name="Caves National Monument" pageId="1" pageNumber="572" stateProvince="Oregon">Caves National Monument</collectingMunicipality>
,
<collectingCounty id="62A6B2E9FFDA6D75DB226B38872E4A45" box="[632,792,1582,1608]" pageId="1" pageNumber="572">Josephine Co.</collectingCounty>
,
<location id="8EA79CBEFFDA6D75DA7E6B38874E4A45" LSID="urn:lsid:plazi:treatment:03D17B73FFDA6D77D9CD688886A84872:8EA79CBEFFDA6D75DA7E6B38874E4A45" box="[804,888,1582,1608]" county="Josephine Co." latitude="42.0974" municipality="Caves National Monument" name="Oregon" pageId="1" pageNumber="572" stateProvince="Oregon">Oregon</location>
(
<geoCoordinate id="EE4CACA2FFDA6D75DADD6B38803D4A44" box="[903,1035,1582,1609]" degrees="42.0974" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="42.0974">42.0974º N</geoCoordinate>
,
<geoCoordinate id="EE4CACA2FFDA6D75DD4D6B3880814A44" box="[1047,1207,1582,1609]" pageId="1" pageNumber="572">-123.4099º W</geoCoordinate>
)
</materialsCitation>
,
<collectingMethod id="5239B272FFDA6D75DD916B3884CF4A60" pageId="1" pageNumber="572">collected from pitfall traps</collectingMethod>
set at different in-cave sites from
<date id="FFC6ECA5FFDA6D75DB2E6B44877E4A61" box="[628,840,1618,1644]" pageId="1" pageNumber="572" value="1992-12-24">
<collectingDate id="EF82154DFFDA6D75DB2E6B44877E4A61" box="[628,840,1618,1644]" pageId="1" pageNumber="572" value="1992-12-24">24 December 1992</collectingDate>
</date>
to
<date id="FFC6ECA5FFDA6D75DA316B44802D4A61" box="[875,1051,1618,1644]" pageId="1" pageNumber="572" value="1993-08-21">
<collectingDate id="EF82154DFFDA6D75DA316B44802D4A61" box="[875,1051,1618,1644]" pageId="1" pageNumber="572" value="1993-08-21">21 August 1993</collectingDate>
</date>
by
<collectorName id="268DAFB3FFDA6D75DD1E6B4580A84A61" box="[1092,1182,1618,1645]" pageId="1" pageNumber="572">R. Reed</collectorName>
and
<collectorName id="268DAFB3FFDA6D75DD896B4581174A61" box="[1235,1313,1618,1645]" pageId="1" pageNumber="572">J. Roth</collectorName>
(
<collectionCode id="ED6952A0FFDA6D75DC6A6B4481A74A61" box="[1328,1425,1618,1644]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="1" pageNumber="572" type="Museum">UWBM</collectionCode>
).
<materialsCitation id="3B10C038FFDA6D75D9CD6B60803C4A9C" ID-GBIF-Occurrence="2423596446" box="[151,1034,1654,1681]" collectorName="The" location="Ghost Room" pageId="1" pageNumber="572" specimenCount="2" typeStatus="holotype">
<collectorName id="268DAFB3FFDA6D75D9CD6B6084F54A9D" box="[151,195,1654,1680]" pageId="1" pageNumber="572">The</collectorName>
<typeStatus id="54C374C7FFDA6D75D9936B60851D4A9D" box="[201,299,1654,1680]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
and several
<typeStatus id="54C374C7FFDA6D75D8EE6B6186294A9C" box="[436,543,1655,1681]" pageId="1" pageNumber="572" type="paratype">paratypes</typeStatus>
were collected at station #11 (
<location id="8EA79CBEFFDA6D75DA346B6080374A9C" LSID="urn:lsid:plazi:treatment:03D17B73FFDA6D77D9CD688886A84872:8EA79CBEFFDA6D75DA346B6080374A9C" box="[878,1025,1654,1681]" name="Ghost Room" pageId="1" pageNumber="572">Ghost Room</location>
)
</materialsCitation>
;
<materialsCitation id="3B10C038FFDA6D75DD4E6B6085F24AD4" ID-GBIF-Occurrence="2423596448" location="Wedding Cake Room" municipality="Imagination Room" pageId="1" pageNumber="572" specimenCount="8" specimenCount-juvenile="2" specimenCount-male="6" typeStatus="paratype">
additional
<typeStatus id="54C374C7FFDA6D75DDD06B6180C34A9C" box="[1162,1269,1655,1681]" pageId="1" pageNumber="572" type="paratype">paratypes</typeStatus>
were collected at stations #3 (connecting tunnel), #5 (
<collectingMunicipality id="6BA3501FFFDA6D75DB1A6B8C87214AB8" box="[576,791,1690,1717]" pageId="1" pageNumber="572">Imagination Room</collectingMunicipality>
;
<specimenCount id="9D7E01ECFFDA6D75DA7B6B8C875A4AB9" box="[801,876,1690,1716]" pageId="1" pageNumber="572" type="male">2 male</specimenCount>
<typeStatus id="54C374C7FFDA6D75DA286B8D87E94AB8" box="[882,991,1691,1717]" pageId="1" pageNumber="572" type="paratype">paratypes</typeStatus>
) and #13 (
<location id="8EA79CBEFFDA6D75DD096B8C81714AB8" LSID="urn:lsid:plazi:treatment:03D17B73FFDA6D77D9CD688886A84872:8EA79CBEFFDA6D75DD096B8C81714AB8" box="[1107,1351,1690,1717]" municipality="Imagination Room" name="Wedding Cake Room" pageId="1" pageNumber="572">Wedding Cake Room</location>
;
<specimenCount id="9D7E01ECFFDA6D75DC086B8C81AA4AB9" box="[1362,1436,1690,1716]" pageId="1" pageNumber="572" type="male">4 male</specimenCount>
<typeStatus id="54C374C7FFDA6D75D9CD6BA985344AD4" box="[151,258,1727,1753]" pageId="1" pageNumber="572" type="paratype">paratypes</typeStatus>
and
<specimenCount id="9D7E01ECFFDA6D75D8636BA8858E4AD4" box="[313,440,1726,1753]" pageId="1" pageNumber="572" type="juvenile">2 juveniles</specimenCount>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDA6D75D99D6BF786394B49" pageId="1" pageNumber="572" type="diagnosis">
<paragraph id="8BC7CA65FFDA6D75D99D6BF786394B49" blockId="1.[151,1437,1545,2005]" pageId="1" pageNumber="572">
<emphasis id="B90C1677FFDA6D75D99D6BF7850C4AF1" bold="true" box="[199,314,1761,1788]" pageId="1" pageNumber="572">Diagnosis</emphasis>
. Distinct from other
<collectingRegion id="49BC0487FFDA6D75DB796BF486414AF1" box="[547,631,1762,1788]" country="United States of America" name="Oregon" pageId="1" pageNumber="572">Oregon</collectingRegion>
species of
<taxonomicName id="4C78B1E6FFDA6D75DBA96BF4876C4AF1" authorityName="Chamberlin" authorityYear="1952" box="[755,858,1762,1788]" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="572" phylum="Arthropoda" rank="genus">
<emphasis id="B90C1677FFDA6D75DBA96BF4876C4AF1" box="[755,858,1762,1788]" italics="true" pageId="1" pageNumber="572">Taiyutyla</emphasis>
</taxonomicName>
in the shape of the gonopods and in having only the pregonopodal legpair 7 with a modified femur, which is flattened and broadened distally and with a distinctly curved distomesal apophysis (
<figureCitation id="1343D6E0FFDA6D75D8CF6A3C85D24B49" box="[405,484,1834,1860]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1925,1950]" captionTargetBox="[151,1432,167,1905]" captionTargetId="figure@2.[151,1432,166,1905]" captionTargetPageId="2" captionText="FIGURES 1–3. Taiyutyla caseophila, new species. Fig. 1. Anterior and posterior gonopods, anterior view; agp, anterior gonopod; pf9, prefemur of posterior gonopod; s9, sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view; agp, anterior gonopod; cx9, coxite of posterior gonopod; T, limb of T-shaped process. Fig. 3. Left leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467968" httpUri="https://zenodo.org/record/3467968/files/figure.png" pageId="1" pageNumber="572">Figs. 3</figureCitation>
,
<figureCitation id="1343D6E0FFDA6D75D8AB6A3F86374B49" box="[497,513,1833,1860]" captionStart="FIGURES 4, 5" captionStartId="3.[151,264,1811,1836]" captionTargetBox="[151,1436,1165,1792]" captionTargetId="figure@3.[151,1436,1165,1792]" captionTargetPageId="3" captionText="FIGURES 4, 5. Taiyutyla caseophila, new species. Fig. 4. Tip of posterior gonopod coxite, posterior view; f, fimbriate region; h, hook-like process; T, limb of T-shaped process. Fig. 5. Pits on femoral knob of leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467970" httpUri="https://zenodo.org/record/3467970/files/figure.png" pageId="1" pageNumber="572">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFDA6D77D99D6A5B804E4DA2" lastPageId="3" lastPageNumber="574" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC7CA65FFDA6D77D99D6A5B86324D86" blockId="1.[151,1437,1545,2005]" lastBlockId="3.[151,1437,153,1151]" lastPageId="3" lastPageNumber="574" pageId="1" pageNumber="572">
<emphasis id="B90C1677FFDA6D75D99D6A5B85674B65" bold="true" box="[199,337,1869,1896]" pageId="1" pageNumber="572">Description</emphasis>
. Male
<typeStatus id="54C374C7FFDA6D75D8F86A5986304B64" box="[418,518,1871,1897]" pageId="1" pageNumber="572" type="paratype">paratype</typeStatus>
. Length,
<quantity id="4C806780FFDA6D75DB2A6A5886F64B64" box="[624,704,1870,1897]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="572" unit="mm" value="11.0">11 mm</quantity>
, greatest width, 1.0 mm. Subtriangular eyepatch of 21 or 22 distinct but unpigmented ocelli (in preserved specimens, ocelli pigmented in life—see below). Metazonite shoulders weakly developed. Segmental setae about equal in length to half metazonite width. Color (after&gt; 20 years of preservation) pale yellow. Pregonopodal legs enlarged, but pairs one to six usually without femoral modifications (the sixth legs of some males have slightly thickened femora with a small swelling on the mesal side). Legpair seven with femora distally broadened, flattened, with prominent distomesal apophysis (
<figureCitation id="1343D6E0FFD86D77DD646DAB80A64CDA" box="[1086,1168,189,215]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1925,1950]" captionTargetBox="[151,1432,167,1905]" captionTargetId="figure@2.[151,1432,166,1905]" captionTargetPageId="2" captionText="FIGURES 1–3. Taiyutyla caseophila, new species. Fig. 1. Anterior and posterior gonopods, anterior view; agp, anterior gonopod; pf9, prefemur of posterior gonopod; s9, sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view; agp, anterior gonopod; cx9, coxite of posterior gonopod; T, limb of T-shaped process. Fig. 3. Left leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467968" httpUri="https://zenodo.org/record/3467968/files/figure.png" pageId="3" pageNumber="574">Figs. 3</figureCitation>
,
<figureCitation id="1343D6E0FFD86D77DDFA6DAA80864CDA" box="[1184,1200,188,215]" captionStart="FIGURES 4, 5" captionStartId="3.[151,264,1811,1836]" captionTargetBox="[151,1436,1165,1792]" captionTargetId="figure@3.[151,1436,1165,1792]" captionTargetPageId="3" captionText="FIGURES 4, 5. Taiyutyla caseophila, new species. Fig. 4. Tip of posterior gonopod coxite, posterior view; f, fimbriate region; h, hook-like process; T, limb of T-shaped process. Fig. 5. Pits on femoral knob of leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467970" httpUri="https://zenodo.org/record/3467970/files/figure.png" pageId="3" pageNumber="574">5</figureCitation>
). Anterior gonopods long, slender, curving posteriorly, abruptly narrowed distally, narrowly acuminate (
<figureCitation id="1343D6E0FFD86D77DD636DF780934CF6" box="[1081,1189,225,251]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1925,1950]" captionTargetBox="[151,1432,167,1905]" captionTargetId="figure@2.[151,1432,166,1905]" captionTargetPageId="2" captionText="FIGURES 1–3. Taiyutyla caseophila, new species. Fig. 1. Anterior and posterior gonopods, anterior view; agp, anterior gonopod; pf9, prefemur of posterior gonopod; s9, sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view; agp, anterior gonopod; cx9, coxite of posterior gonopod; T, limb of T-shaped process. Fig. 3. Left leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467968" httpUri="https://zenodo.org/record/3467968/files/figure.png" pageId="3" pageNumber="574">Figs. 1, 2</figureCitation>
). Posterior gonopodal coxites stout, broad, with mesal T-shaped branch short. Proximal part of posterior gonopod acuminate, with small basal tooth, distal part complexly fimbriate. Body of posterior gonopod coxite with subterminal fimbriate area, terminating in distal hook-like structure (
<figureCitation id="1343D6E0FFD86D77DB306C5B868D4D6A" box="[618,699,333,359]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1925,1950]" captionTargetBox="[151,1432,167,1905]" captionTargetId="figure@2.[151,1432,166,1905]" captionTargetPageId="2" captionText="FIGURES 1–3. Taiyutyla caseophila, new species. Fig. 1. Anterior and posterior gonopods, anterior view; agp, anterior gonopod; pf9, prefemur of posterior gonopod; s9, sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view; agp, anterior gonopod; cx9, coxite of posterior gonopod; T, limb of T-shaped process. Fig. 3. Left leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467968" httpUri="https://zenodo.org/record/3467968/files/figure.png" pageId="3" pageNumber="574">Figs. 2</figureCitation>
,
<figureCitation id="1343D6E0FFD86D77DB926C5B86EF4D6A" box="[712,729,333,359]" captionStart="FIGURES 4, 5" captionStartId="3.[151,264,1811,1836]" captionTargetBox="[151,1436,1165,1792]" captionTargetId="figure@3.[151,1436,1165,1792]" captionTargetPageId="3" captionText="FIGURES 4, 5. Taiyutyla caseophila, new species. Fig. 4. Tip of posterior gonopod coxite, posterior view; f, fimbriate region; h, hook-like process; T, limb of T-shaped process. Fig. 5. Pits on femoral knob of leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467970" httpUri="https://zenodo.org/record/3467970/files/figure.png" pageId="3" pageNumber="574">4</figureCitation>
). Tenth legpair with coxal glands, eleventh legpair with basal, curved apophyses on prefemora.
</paragraph>
<caption id="DF079AEDFFD96D76D9CD6A9381B04BEB" ID-DOI="http://doi.org/10.5281/zenodo.3467968" ID-Zenodo-Dep="3467968" httpUri="https://zenodo.org/record/3467968/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,264,1925,1950]" targetBox="[151,1432,167,1905]" targetPageId="2">
<paragraph id="8BC7CA65FFD96D76D9CD6A9381B04BEB" blockId="2.[151,1436,1925,2022]" pageId="2" pageNumber="573">
<emphasis id="B90C1677FFD96D76D9CD6A93850D4B93" bold="true" box="[151,315,1925,1950]" pageId="2" pageNumber="573">FIGURES 1–3.</emphasis>
<emphasis id="B90C1677FFD96D76D81B6A93862F4B90" box="[321,537,1925,1949]" italics="true" pageId="2" pageNumber="573">
<taxonomicName id="4C78B1E6FFD96D76D81B6A9386234B90" authority="Shear &amp; Crawford, 2019" authorityName="Shear &amp; Crawford" authorityYear="2019" box="[321,533,1925,1949]" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="caseophila" status="sp. nov.">Taiyutyla caseophila</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23FAB0CFFD96D76DB456A9086AA4B93" box="[543,668,1925,1950]" pageId="2" pageNumber="573" rank="species">new species</taxonomicNameLabel>
. Fig. 1. Anterior and posterior gonopods, anterior view;
<emphasis id="B90C1677FFD96D76DD826A9080C94B93" bold="true" box="[1240,1279,1926,1950]" pageId="2" pageNumber="573">agp</emphasis>
, anterior gonopod;
<emphasis id="B90C1677FFD96D76D9916ABF84C24BCF" bold="true" box="[203,244,1961,1986]" pageId="2" pageNumber="573">pf9,</emphasis>
prefemur of posterior gonopod;
<emphasis id="B90C1677FFD96D76DB656ABF866B4BCF" bold="true" box="[575,605,1961,1986]" pageId="2" pageNumber="573">s9,</emphasis>
sternum of posterior gonopods. Fig. 2. Anterior and posterior gonopods, posterior view;
<emphasis id="B90C1677FFD96D76D98C6AD885324BEB" bold="true" box="[214,260,1998,2022]" pageId="2" pageNumber="573">agp,</emphasis>
anterior gonopod;
<emphasis id="B90C1677FFD96D76D89F6ADB85C64BEB" bold="true" box="[453,496,1997,2022]" pageId="2" pageNumber="573">cx9,</emphasis>
coxite of posterior gonopod;
<emphasis id="B90C1677FFD96D76DA466AD887044BEB" bold="true" box="[796,818,1998,2022]" pageId="2" pageNumber="573">T,</emphasis>
limb of T-shaped process. Fig. 3. Left leg 7, anterior view.
</paragraph>
</caption>
<paragraph id="8BC7CA65FFD86D77D99D6C83804E4DA2" blockId="3.[151,1437,153,1151]" box="[199,1144,405,431]" pageId="3" pageNumber="574">
Female
<typeStatus id="54C374C7FFD86D77D87B6C8385B34DA2" box="[289,389,405,431]" pageId="3" pageNumber="574" type="paratype">paratype</typeStatus>
. Length,
<quantity id="4C806780FFD86D77D8B66C8386674DA2" box="[492,593,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="574" unit="mm" value="11.4">11.4 mm</quantity>
, width
<quantity id="4C806780FFD86D77DBFE6C8386CB4DA2" box="[676,765,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="574" unit="mm" value="1.1">1.1 mm</quantity>
. Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFD86D77D99D6CAE868B483A" pageId="3" pageNumber="574" type="distribution">
<paragraph id="8BC7CA65FFD86D77D99D6CAE808F4DDE" blockId="3.[151,1437,153,1151]" box="[199,1209,440,467]" pageId="3" pageNumber="574">
<emphasis id="B90C1677FFD86D77D99D6CAE856F4DDE" bold="true" box="[199,345,440,467]" pageId="3" pageNumber="574">Distribution</emphasis>
. Known only from the
<typeStatus id="54C374C7FFD86D77DB3A6CAF86A64DDE" box="[608,656,441,467]" pageId="3" pageNumber="574">type</typeStatus>
locality and other locations within
<collectingRegion id="49BC0487FFD86D77DD476CAF80474DDE" box="[1053,1137,441,467]" country="United States of America" name="Oregon" pageId="3" pageNumber="574">Oregon</collectingRegion>
Cave.
</paragraph>
<paragraph id="8BC7CA65FFD86D77D99D6CCB87404EA6" blockId="3.[151,1437,153,1151]" pageId="3" pageNumber="574">
<emphasis id="B90C1677FFD86D77D99D6CCB853C4DFA" bold="true" box="[199,266,477,503]" pageId="3" pageNumber="574">Notes</emphasis>
. The femoral knobs of the seventh legs bear a mesal group of about 20 small pits. Under high magnification, at least some of these pits have numerous small pores within them (
<figureCitation id="1343D6E0FFD86D77DAE56F1780354E16" box="[959,1027,512,539]" captionStart="FIGURES 4, 5" captionStartId="3.[151,264,1811,1836]" captionTargetBox="[151,1436,1165,1792]" captionTargetId="figure@3.[151,1436,1165,1792]" captionTargetPageId="3" captionText="FIGURES 4, 5. Taiyutyla caseophila, new species. Fig. 4. Tip of posterior gonopod coxite, posterior view; f, fimbriate region; h, hook-like process; T, limb of T-shaped process. Fig. 5. Pits on femoral knob of leg 7, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3467970" httpUri="https://zenodo.org/record/3467970/files/figure.png" pageId="3" pageNumber="574">Fig. 5</figureCitation>
). These pits and pores have not been found on the femoral knobs of any other
<taxonomicName id="4C78B1E6FFD86D77DB296F3386EC4E32" authorityName="Chamberlin" authorityYear="1952" box="[627,730,549,575]" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B90C1677FFD86D77DB296F3386EC4E32" box="[627,730,549,575]" italics="true" pageId="3" pageNumber="574">Taiyutyla</emphasis>
</taxonomicName>
species so far described, and a scanning electron microscopy study of several species for a forthcoming monograph on northwestern North American conotylids did not show the pits on any examined species. The pores within the pits are only about
<quantity id="4C806780FFD86D77DA826F7B80194E8A" box="[984,1071,621,647]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="574" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 µm</quantity>
in diameter and so are likely the outlets of unicellular glands, but their function remains unknown.
</paragraph>
<paragraph id="8BC7CA65FFD86D77D99D6FA380F44FE2" blockId="3.[151,1437,153,1151]" pageId="3" pageNumber="574">
Specimens were collected throughout the dark zone of the cave (but included sites on the commercial tour route, lighted during tours), during each month from
<date id="FFC6ECA5FFD86D77DBB26FCF874E4EFE" box="[744,888,729,755]" pageId="3" pageNumber="574" value="1992-08">August 1992</date>
to
<date id="FFC6ECA5FFD86D77DAC06FCF801B4EFE" box="[922,1069,729,755]" pageId="3" pageNumber="574" value="1993-08">August 1993</date>
. In all, over
<specimenCount id="9D7E01ECFFD86D77DDE36FCF81694EFE" box="[1209,1375,729,755]" pageId="3" pageNumber="574" type="generic">200 specimens</specimenCount>
were collected, all but 5 by cheese-baited pitfall traps. The traps, which were baited with spreadable Limburger cheese smeared on the underside of a rock covering the trap, were left in place for periods ranging from 2 to 4 weeks. No specimens were found at the cave entrance or twilight sites. Of the five hand-collected specimens, one was found on the cave wall, one under a half-imbedded rock, and two associated with mammal scats (one next to a moldy
<taxonomicName id="4C78B1E6FFD86D77D9CD6E9884CA4FAA" authorityName="Say &amp; Ord" authorityYear="1825" box="[151,252,910,935]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B90C1677FFD86D77D9CD6E9884CA4FAA" box="[151,252,910,935]" italics="true" pageId="3" pageNumber="574">Neotoma</emphasis>
</taxonomicName>
scat and one apparently feeding on a
<taxonomicName id="4C78B1E6FFD86D77DBC66E9887144FAA" authorityName="Gloger" authorityYear="1841" box="[668,802,910,935]" class="Mammalia" family="Muridae" genus="Peromyscus" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="574" phylum="Chordata" rank="genus">
<emphasis id="B90C1677FFD86D77DBC66E9887144FAA" box="[668,802,910,935]" italics="true" pageId="3" pageNumber="574">Peromyscus</emphasis>
</taxonomicName>
scat, or its microflora). Observations of freshly collected specimens indicated pale yellowish cuticular pigmentation and black, pigmented ocelli. Epigean species of
<taxonomicName id="4C78B1E6FFD86D77DC0C6EA484F74FE2" authorityName="Chamberlin" authorityYear="1952" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B90C1677FFD86D77DC0C6EA484F74FE2" italics="true" pageId="3" pageNumber="574">Taiyutyla</emphasis>
</taxonomicName>
are usually well pigmented, tan to brown, and with dark purplish brown cuticular mottling.
</paragraph>
<paragraph id="8BC7CA65FFD86D77D99D6EEF868B483A" blockId="3.[151,1437,153,1151]" pageId="3" pageNumber="574">
Shear (2011) mentioned a conotylid collected in the Marble Mountain Wilderness caves of Siskiyou Co.,
<collectingRegion id="49BC0487FFD86D77DC3E6EEF84EB483A" country="United States of America" name="California" pageId="3" pageNumber="574">California</collectingRegion>
, but only females were available for study.
</paragraph>
</subSubSection>
<subSubSection id="C36299EEFFD86D77D99D695786A84872" pageId="3" pageNumber="574" type="etymology">
<paragraph id="8BC7CA65FFD86D77D99D695786A84872" blockId="3.[151,1437,153,1151]" pageId="3" pageNumber="574">
<emphasis id="B90C1677FFD86D77D99D6957857E4856" bold="true" box="[199,328,1089,1115]" pageId="3" pageNumber="574">Etymology</emphasis>
. The species epithet (Latin) is an adjective meaning “cheese-loving” and refers to the capture of most known specimens in cheese-baited traps.
</paragraph>
</subSubSection>
</treatment>
</document>
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