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<document id="F5DEE28C279325C21A55FE2899A105BC" ID-CLB-Dataset="29210" ID-DOI="10.11646/zootaxa.4497.2.7" ID-GBIF-Dataset="aef8fd78-8592-4090-bcff-353eaebf47a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1452195" ID-ZooBank="13627E62-F717-4873-B752-33A0EF17516F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539065916076" checkinUser="plazi" docAuthor="Fomichev, Alexander A. &amp; Marusik, Yuri M." docDate="2018" docId="03A3EA5A994170572FCBFDB4936B0E71" docLanguage="en" docName="zootaxa.4497.2.7.pdf" docOrigin="Zootaxa 4497 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantholycosa vahterae Fomichev &amp; Marusik 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="273" masterDocId="FF9A9222994070552F5CFFB8914F0D36" masterDocTitle="Five new species of the Acantholycosa - complex (Araneae: Lycosidae) from Mongolia" masterLastPageNumber="284" masterPageNumber="271" pageNumber="272" updateTime="1698568488534" updateUser="ExternalLinkService">
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<mods:title id="56526C410E46413C62F8529C7CA05006">Five new species of the Acantholycosa - complex (Araneae: Lycosidae) from Mongolia</mods:title>
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<mods:namePart id="0F741544D37F36BDDF5328983E32DBBC">Fomichev, Alexander A.</mods:namePart>
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<mods:namePart id="A57AC74E69FD85EA986BBFAF357D61AD">Marusik, Yuri M.</mods:namePart>
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<mods:date id="300540A8DA749A62B71C5B6B0A4E02D9">2018</mods:date>
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<mods:number id="8B875EE89224B10DB69E346EEA78ED78">2018-10-09</mods:number>
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<treatment id="03A3EA5A994170572FCBFDB4936B0E71" ID-DOI="http://doi.org/10.5281/zenodo.5978167" ID-GBIF-Taxon="148657230" ID-Zenodo-Dep="5978167" LSID="urn:lsid:plazi:treatment:03A3EA5A994170572FCBFDB4936B0E71" httpUri="http://treatment.plazi.org/id/03A3EA5A994170572FCBFDB4936B0E71" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272">
<subSubSection id="C31008C7994170542FCBFDB493E90F71" pageId="1" pageNumber="272" type="nomenclature">
<paragraph id="8BB55B4C994170542FCBFDB4934E0F10" blockId="1.[151,678,524,583]" box="[151,513,524,550]" pageId="1" pageNumber="272">
<heading id="D0FDEC20994170542FCBFDB4934E0F10" bold="true" box="[151,513,524,550]" fontSize="11" level="1" pageId="1" pageNumber="272" reason="1">
<emphasis id="B97E875E994170542FCBFDB4934E0F10" bold="true" box="[151,513,524,550]" pageId="1" pageNumber="272">
<taxonomicName id="4C0A20CF994170542FCBFDB490F70F10" ID-CoL="8YFK" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[151,440,524,550]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542FCBFDB490F70F10" bold="true" box="[151,440,524,550]" italics="true" pageId="1" pageNumber="272">Acantholycosa vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542EE3FDB4934E0F10" box="[447,513,524,550]" pageId="1" pageNumber="272" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB55B4C994170542FCBFD9693E90F71" blockId="1.[151,678,524,583]" box="[151,678,558,583]" pageId="1" pageNumber="272">
<figureCitation id="133147C9994170542FCBFD9690480F71" box="[151,263,558,583]" captionStart="FIGURE 1" captionStartId="2.[151,250,1972,1994]" captionTargetBox="[152,1436,902,1951]" captionTargetId="figure@2.[150,1436,892,1951]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and male palp of Acantholycosa vahterae sp. n. (a–d) and Mongolicosa azarkinae sp. n. (e–g). a, e male habitus, dorsal; b female habitus, dorsal; c, g palp, retrolateral; d, f palp, ventral. Scale: a–b, e = 2 mm; c–d, f–g = 0.2 mm." httpUri="https://zenodo.org/record/1452197/files/figure.png" pageId="1" pageNumber="272">Figs 1a‒d</figureCitation>
,
<figureCitation id="133147C9994170542E4FFD9790050F71" box="[275,330,559,583]" captionStart="FIGURE 2" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[154,1431,327,1881]" captionTargetId="figure@4.[151,1436,318,1881]" captionTargetPageId="4" captionText="FIGURE 2. Male palp of Acantholycosa vahterae sp. n. (a–e) and A. kronestedti sp. n. (f–j). a, f bulb, ventral; b, g bulb, anterior; c, h bulb, retrolateral; d, i tegulum, ventral; e, j tegulum, anterior. Scale = 0.2 mm. Abbreviations: Es basal spine of embolus, Pa palea, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452199/files/figure.png" pageId="1" pageNumber="272">2a‒e</figureCitation>
,
<figureCitation id="133147C9994170542E0AFD9790C20F71" box="[342,397,559,583]" captionStart="FIGURE 3" captionStartId="5.[151,250,1892,1914]" captionTargetBox="[151,1436,223,1853]" captionTargetId="figure@5.[151,1436,223,1853]" captionTargetPageId="5" captionText="FIGURE 3. Bulb of Acantholycosa vahterae sp. n. (a–c), A. kronestedti sp. n. (d–f) and Mongolicosa azarkinae sp. n. (g–i, embolic spine broken). a, d, g ventral; b, e, h retrolateral; c, f, i anterior. Scale = 0.1 mm (if indicated). Abbreviations: Em embolus, Es basal spine of embolus, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452201/files/figure.png" pageId="1" pageNumber="272">3a‒c</figureCitation>
,
<figureCitation id="133147C9994170542EC4FD97909F0F71" box="[408,464,559,583]" captionStart="FIGURE 4" captionStartId="6.[151,250,1172,1194]" captionTargetBox="[151,1436,208,1138]" captionTargetId="figure@6.[151,1436,208,1138]" captionTargetPageId="6" captionText="FIGURE 4. Embolic division of Acantholycosa vahterae sp. n. (a–c), A. kronestedti sp. n. (d–f) and Mongolicosa azarkinae sp. n. (g–h, embolic spine broken). a, d, g ventral; b, e, h anterior; c, f posterior. Scale = 0.1 mm." httpUri="https://zenodo.org/record/1452203/files/figure.png" pageId="1" pageNumber="272">4a‒c</figureCitation>
,
<figureCitation id="133147C9994170542E87FD96935C0F71" box="[475,531,558,583]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[155,1428,1290,1929]" captionTargetId="figure@6.[151,1436,1279,1929]" captionTargetPageId="6" captionText="FIGURE 5. Embolic division of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d anterior; b, e retrolateral; c, f ventral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/1452205/files/figure.png" pageId="1" pageNumber="272">5a‒c</figureCitation>
,
<figureCitation id="133147C9994170542D42FD96931A0F71" box="[542,597,558,583]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="1" pageNumber="272">6a‒c</figureCitation>
,
<figureCitation id="133147C9994170542D3DFD9793310F71" box="[609,638,559,583]" captionStart="FIGURE 9" captionStartId="11.[151,250,1734,1756]" captionTargetBox="[150,1436,267,1717]" captionTargetId="figure@11.[150,1436,267,1717]" captionTargetPageId="11" captionText="FIGURE 9. Habitats of Acantholycosa vahterae sp. n. (a), A. kronestedti sp. n. (b), Mongolicosa azarkinae sp. n. (c), M. ozkutuki sp. n. (d) and M. cherepanovi sp. n. (e–f)." httpUri="https://zenodo.org/record/1452213/files/figure.png" pageId="1" pageNumber="272">9a</figureCitation>
,
<figureCitation id="133147C9994170542DD6FD9793E90F71" box="[650,678,559,583]" captionStart="FIGURE 10" captionStartId="12.[151,250,1631,1653]" captionTargetBox="[153,1437,1254,1608]" captionTargetId="figure@12.[151,1437,1254,1609]" captionTargetPageId="12" captionText="FIGURE 10. Collecting localities of Acantholycosa vahterae sp. n. (5), A. kronestedti sp. n. (4), Mongolicosa azarkinae sp. n. (3), M. cherepanovi sp. n. (1) and M. ozkutuki sp. n. (2)." httpUri="https://zenodo.org/record/1452215/files/figure.png" pageId="1" pageNumber="272">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31008C7994170542FCBFDCE94D20FE1" pageId="1" pageNumber="272" type="description">
<paragraph id="8BB55B4C994170542FCBFDCE94D20FE1" blockId="1.[151,1437,630,1447]" pageId="1" pageNumber="272">
<materialsCitation id="3B625111994170542FCBFDCE93A10FE1" ID-GBIF-Occurrence="1915261933" collectionCode="ZMMU" collectorName="Mt. Range &amp; A. A. Fomichev" country="Mongolia" elevation="3480" latitude="45.4" location="Mt. Pass" longLatPrecision="1130" longitude="97.11667" municipality="Dotiyn-Davaa Mt." pageId="1" pageNumber="272" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altai" typeStatus="holotype">
<emphasis id="B97E875E994170542FCBFDCE91AC0FB9" bold="true" box="[151,227,630,655]" pageId="1" pageNumber="272">
<typeStatus id="54B1E5EE994170542FCBFDCE91900FB9" box="[151,223,630,655]" pageId="1" pageNumber="272">Types</typeStatus>
.
</emphasis>
<typeStatus id="54B1E5EE994170542FB7FDCE901C0FB9" box="[235,339,630,655]" pageId="1" pageNumber="272" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0C90C5994170542E00FDCF903E0FB8" box="[348,369,631,654]" pageId="1" pageNumber="272" type="male">♂</specimenCount>
(
<collectionCode id="ED1BC389994170542EDCFDCF90940FB9" box="[384,475,631,655]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="272">ZMMU</collectionCode>
)
<collectingCountry id="F31D1BDC994170542EB7FDCF93C90FB9" box="[491,646,631,655]" name="Mongolia" pageId="1" pageNumber="272">MONGOLIA</collectingCountry>
,
<emphasis id="B97E875E994170542DCEFDCE92120FB9" bold="true" box="[658,861,630,655]" italics="true" pageId="1" pageNumber="272">
<collectingRegion id="49CE95AE994170542DCEFDCE92460FB9" box="[658,777,630,655]" country="Mongolia" name="Govi-Altay" pageId="1" pageNumber="272">Govi-Altai</collectingRegion>
Aimag
</emphasis>
, Cakhir-Khaalgyn-Nuruu
<collectorName id="26FF3E9A994170542BDBFDCF944D0FB9" box="[1159,1282,631,655]" pageId="1" pageNumber="272">Mt. Range</collectorName>
,
<quantity id="4CF2F6A9994170542A52FDCE942C0FB9" box="[1294,1379,630,655]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="272" unit="km" value="3.5" valueMax="5.0" valueMin="2.0">2‒5 km</quantity>
SSE from
<collectingMunicipality id="6BD1C136994170542F88FD2290ED0F84" box="[212,418,666,691]" pageId="1" pageNumber="272">
Dotiyn-Davaa
<location id="8ED50D97994170542E25FD2390ED0F84" LSID="urn:lsid:plazi:treatment:03A3EA5A994170572FCBFDB4936B0E71:8ED50D97994170542E25FD2390ED0F84" box="[377,418,667,690]" country="Mongolia" latitude="45.4" longLatPrecision="1130" longitude="97.11667" municipality="Dotiyn-Davaa Mt." name="Mt. Pass" pageId="1" pageNumber="272" stateProvince="Govi-Altai">Mt.</location>
</collectingMunicipality>
Pass (
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,
<geoCoordinate id="EE3E3D8B994170542D08FD2293E10F85" box="[596,686,666,691]" direction="east" orientation="longitude" pageId="1" pageNumber="272" precision="925" value="97.083336">97°05'E</geoCoordinate>
‒
<geoCoordinate id="EE3E3D8B994170542D95FD2292680F85" box="[713,807,666,691]" direction="north" orientation="latitude" pageId="1" pageNumber="272" precision="925" value="45.4">45°24'N</geoCoordinate>
,
<geoCoordinate id="EE3E3D8B994170542C6FFD2392C00F85" box="[819,911,667,691]" direction="east" orientation="longitude" pageId="1" pageNumber="272" precision="925" value="97.11667">97°07'E</geoCoordinate>
), mountain stony tundra with screes and rocks,
<elevation id="0027BC7F994170542FCBFD06907B0FE0" box="[151,308,702,727]" metricMagnitude="3" metricUnit="m" metricValue="3.4799999999999995" metricValueMax="3.66" metricValueMin="3.3" pageId="1" pageNumber="272" unit="m" value="3480.0" valueMax="3660.0" valueMin="3300.0">
<quantity id="4CF2F6A9994170542FCBFD06907B0FE0" box="[151,308,702,727]" metricMagnitude="3" metricUnit="m" metricValue="3.4799999999999995" metricValueMax="3.66" metricValueMin="3.3" pageId="1" pageNumber="272" unit="m" value="3480.0" valueMax="3660.0" valueMin="3300.0">3300‒3660 m</quantity>
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, 4.07.2017 (
<collectorName id="26FF3E9A994170542EE3FD0793200FE1" box="[447,623,702,727]" pageId="1" pageNumber="272">A.A. Fomichev</collectorName>
)
<typeStatus id="54B1E5EE994170542D21FD0793A10FE1" box="[637,750,703,727]" pageId="1" pageNumber="272" type="paratype">Paratypes</typeStatus>
</materialsCitation>
: 2♂ 3♀ (ISEA), 1♂ 4♀ (ZMMU) together with the holotype.
</paragraph>
</subSubSection>
<subSubSection id="C31008C7994170542F9BFD5A93B50E29" pageId="1" pageNumber="272" type="etymology">
<paragraph id="8BB55B4C994170542F9BFD5A93B50E29" blockId="1.[151,1437,630,1447]" pageId="1" pageNumber="272">
<emphasis id="B97E875E994170542F9BFD5A90090FCD" bold="true" box="[199,326,738,763]" pageId="1" pageNumber="272">Etymology</emphasis>
. The specific name is a patronym in honor of our colleague, Varpu Vahtera (Zoological Museum, University of Turku,
<collectingCountry id="F31D1BDC994170542ED8FCBE90910E29" box="[388,478,774,799]" name="Finland" pageId="1" pageNumber="272">Finland</collectingCountry>
), an expert in centipedes.
</paragraph>
</subSubSection>
<subSubSection id="C31008C7994170542F9BFC9293BE0821" pageId="1" pageNumber="272" type="diagnosis">
<paragraph id="8BB55B4C994170542F9BFC9293BE0821" blockId="1.[151,1437,630,1447]" pageId="1" pageNumber="272">
<emphasis id="B97E875E994170542F9BFC9290750E75" bold="true" box="[199,314,810,835]" pageId="1" pageNumber="272">Diagnosis</emphasis>
.
<taxonomicName id="4C0A20CF994170542E17FC9293120E75" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[331,605,810,835]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542E17FC9293120E75" box="[331,605,810,835]" italics="true" pageId="1" pageNumber="272">Acantholycosa vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542D3BFC9293E50E75" box="[615,682,810,835]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542D3BFC9293E50E75" bold="true" box="[615,682,810,835]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C0A20CF994170542CD7FC9391800E51" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="katunensis">
<emphasis id="B97E875E994170542CD7FC93956C0E75" box="[907,1059,810,835]" italics="true" pageId="1" pageNumber="272">A. katunensis</emphasis>
<bibRefCitation id="EF9B26BD994170542B72FC9291800E51" pageId="1" pageNumber="272" refId="ref5098" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article">
Marusik, Azarkina &amp; Koponen,
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</bibRefCitation>
</taxonomicName>
in having the almost identical shape of the paleal apophysis (
<emphasis id="B97E875E994170542CEAFCF7929A0E50" box="[950,981,847,870]" italics="true" pageId="1" pageNumber="272">Pa</emphasis>
), the widened embolic tip, the tegular apophysis lacking an apical arm, the epigynal septum with parallel margins and not widened posteriorly. The new species readily differs from the related species in having a large triangle-shaped spine (
<emphasis id="B97E875E994170542BD0FC2F95E70E98" box="[1164,1192,919,942]" italics="true" pageId="1" pageNumber="272">Es</emphasis>
) at the embolic base (
<figureCitation id="133147C9994170542FC3FC0291B60EE5" box="[159,249,954,979]" captionStart="FIGURE 2" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[154,1431,327,1881]" captionTargetId="figure@4.[151,1436,318,1881]" captionTargetPageId="4" captionText="FIGURE 2. Male palp of Acantholycosa vahterae sp. n. (a–e) and A. kronestedti sp. n. (f–j). a, f bulb, ventral; b, g bulb, anterior; c, h bulb, retrolateral; d, i tegulum, ventral; e, j tegulum, anterior. Scale = 0.2 mm. Abbreviations: Es basal spine of embolus, Pa palea, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452199/files/figure.png" pageId="1" pageNumber="272">Figs 2b</figureCitation>
,
<figureCitation id="133147C9994170542E54FC0290680EE5" box="[264,295,954,979]" captionStart="FIGURE 4" captionStartId="6.[151,250,1172,1194]" captionTargetBox="[151,1436,208,1138]" captionTargetId="figure@6.[151,1436,208,1138]" captionTargetPageId="6" captionText="FIGURE 4. Embolic division of Acantholycosa vahterae sp. n. (a–c), A. kronestedti sp. n. (d–f) and Mongolicosa azarkinae sp. n. (g–h, embolic spine broken). a, d, g ventral; b, e, h anterior; c, f posterior. Scale = 0.1 mm." httpUri="https://zenodo.org/record/1452203/files/figure.png" pageId="1" pageNumber="272">4b</figureCitation>
), which is absent from
<taxonomicName id="4C0A20CF994170542D61FC0393990EE5" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[573,726,954,979]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="katunensis">
<emphasis id="B97E875E994170542D61FC0393990EE5" box="[573,726,954,979]" italics="true" pageId="1" pageNumber="272">A. katunensis</emphasis>
</taxonomicName>
. The female of the new species has the arch-shaped anterior pocket not subdivided into 2 parts and receptacles not reaching the anterior margin of fovea (
<figureCitation id="133147C9994170542B8AFC6694080EC1" box="[1238,1351,990,1015]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="1" pageNumber="272">Figs 6a‒c</figureCitation>
<emphasis id="B97E875E994170542A0DFC6794270EC0" box="[1361,1384,991,1014]" italics="true" pageId="1" pageNumber="272">vs</emphasis>
. the anterior pocket being clearly subdivided into 2 parts, receptacles long, reaching the anterior margin of fovea in
<taxonomicName id="4C0A20CF994170542AD8FBBB90420909" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="katunensis">
<emphasis id="B97E875E994170542AD8FBBB90420909" italics="true" pageId="1" pageNumber="272">A. katunensis</emphasis>
</taxonomicName>
; cf. figs.
<quantity id="4CF2F6A9994170542E23FB9F90900909" box="[383,479,1062,1087]" metricMagnitude="0" metricUnit="m" metricValue="2.0955" metricValueMax="2.1082" metricValueMin="2.0827999999999998" pageId="1" pageNumber="272" unit="in" value="82.5" valueMax="83.0" valueMin="82.0">82-83 in</quantity>
<bibRefCitation id="EF9B26BD994170542EB5FB9E939E0909" box="[489,721,1062,1087]" pageId="1" pageNumber="272" refId="ref5098" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article">
Marusik
<emphasis id="B97E875E994170542D0CFB9F93CC0909" box="[592,643,1062,1087]" italics="true" pageId="1" pageNumber="272">et al</emphasis>
. 2004
</bibRefCitation>
).
<taxonomicName id="4C0A20CF994170542DB9FB9F92260909" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[741,873,1062,1087]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542DB9FB9F92260909" box="[741,873,1062,1087]" italics="true" pageId="1" pageNumber="272">A. vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542C2FFB9E92FA0909" box="[883,949,1062,1087]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542C2FFB9E92FA0909" bold="true" box="[883,949,1062,1087]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0A20CF994170542B13FB9F95A20909" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[1103,1261,1062,1087]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="kronestedti" status="sp. nov.">
<emphasis id="B97E875E994170542B13FB9F95A20909" box="[1103,1261,1062,1087]" italics="true" pageId="1" pageNumber="272">A. kronestedti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542BAAFB9E94770909" box="[1270,1336,1062,1087]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542BAAFB9E94770909" bold="true" box="[1270,1336,1062,1087]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
, another species occurring in
<collectingRegion id="49CE95AE994170542EDBFBF2934F0955" box="[391,512,1098,1123]" country="Mongolia" name="Govi-Altay" pageId="1" pageNumber="272">Govi-Altai</collectingRegion>
Aimag, in having the paleal apophysis with a triangular anterior outgrowth, no small spine (
<emphasis id="B97E875E994170542E75FBD7900E09B0" box="[297,321,1135,1158]" italics="true" pageId="1" pageNumber="272">Ss</emphasis>
) on the anterior embolic margin and the shorter posterior arm of tegular apophysis (cf.
<figureCitation id="133147C9994170542A72FBD6904D099D" captionStart="FIGURE 2" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[154,1431,327,1881]" captionTargetId="figure@4.[151,1436,318,1881]" captionTargetPageId="4" captionText="FIGURE 2. Male palp of Acantholycosa vahterae sp. n. (a–e) and A. kronestedti sp. n. (f–j). a, f bulb, ventral; b, g bulb, anterior; c, h bulb, retrolateral; d, i tegulum, ventral; e, j tegulum, anterior. Scale = 0.2 mm. Abbreviations: Es basal spine of embolus, Pa palea, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452199/files/figure.png" pageId="1" pageNumber="272">Figs 2a‒e and 2 f‒j</figureCitation>
). The females of the two species occurring in
<collectingRegion id="49CE95AE994170542C48FB2A92C2099D" box="[788,909,1170,1195]" country="Mongolia" name="Govi-Altay" pageId="1" pageNumber="272">Govi-Altai</collectingRegion>
Aimag can be distinguished by the shape and size of septal lobes (
<emphasis id="B97E875E994170542E27FB0E90DE09F9" box="[379,401,1206,1231]" italics="true" pageId="1" pageNumber="272">Sl</emphasis>
), which are longer than wide in
<taxonomicName id="4C0A20CF994170542DA7FB0F923309F9" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[763,892,1206,1231]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542DA7FB0F924309F8" box="[763,780,1207,1230]" italics="true" pageId="1" pageNumber="272">A</emphasis>
.
<emphasis id="B97E875E994170542C46FB0E923309F9" box="[794,892,1206,1231]" italics="true" pageId="1" pageNumber="272">vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542CDFFB0E928D09F9" box="[899,962,1206,1231]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542CDFFB0E928D09F9" bold="true" box="[899,962,1206,1231]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
and almost triangular in
<taxonomicName id="4C0A20CF994170542B85FB0F943C09F9" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[1241,1395,1206,1231]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="kronestedti" status="sp. nov.">
<emphasis id="B97E875E994170542B85FB0F943C09F9" box="[1241,1395,1206,1231]" italics="true" pageId="1" pageNumber="272">A. kronestedti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542A26FB0E91E109C5" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542A26FB0E91E109C5" bold="true" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
(cf.
<figureCitation id="133147C9994170542FBFFB6290F309C5" box="[227,444,1242,1267]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="1" pageNumber="272">Figs 6a‒c and 6d‒f</figureCitation>
) and by direction of receptacles (
<emphasis id="B97E875E994170542C6BFB64921B09C5" box="[823,852,1244,1267]" italics="true" pageId="1" pageNumber="272">Re</emphasis>
), which directed inwardly in
<taxonomicName id="4C0A20CF994170542BFFFB63946B09C5" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[1187,1316,1242,1267]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542BFFFB63946B09C5" box="[1187,1316,1242,1267]" italics="true" pageId="1" pageNumber="272">A. vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542A70FB62942309C5" box="[1324,1388,1242,1267]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542A70FB62942309C5" bold="true" box="[1324,1388,1242,1267]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
and outwardly in
<taxonomicName id="4C0A20CF994170542E70FB4790890821" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[300,454,1278,1303]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="kronestedti" status="sp. nov.">
<emphasis id="B97E875E994170542E70FB4790890821" box="[300,454,1278,1303]" italics="true" pageId="1" pageNumber="272">A. kronestedti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542E91FB4693420821" box="[461,525,1278,1303]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542E91FB4693420821" bold="true" box="[461,525,1278,1303]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
(cf.
<figureCitation id="133147C9994170542D1DFB4693AC0821" box="[577,739,1278,1303]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="1" pageNumber="272">Figs 6c and 6f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31008C7994170572F9BFA9A928B0FC9" lastPageId="2" lastPageNumber="273" pageId="1" pageNumber="272" type="description">
<paragraph id="8BB55B4C994170542F9BFA9A903F0891" blockId="1.[151,1437,630,1447]" pageId="1" pageNumber="272">
<emphasis id="B97E875E994170542F9BFA9A9016080D" bold="true" box="[199,345,1314,1339]" pageId="1" pageNumber="272">Description.</emphasis>
Male (
<typeStatus id="54B1E5EE994170542EF0FA9A935B080D" box="[428,532,1314,1339]" pageId="1" pageNumber="272" type="holotype">holotype</typeStatus>
). General appearance as in
<figureCitation id="133147C9994170542C0CFA9A92DF080D" box="[848,912,1314,1339]" captionStart="FIGURE 1" captionStartId="2.[151,250,1972,1994]" captionTargetBox="[152,1436,902,1951]" captionTargetId="figure@2.[150,1436,892,1951]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and male palp of Acantholycosa vahterae sp. n. (a–d) and Mongolicosa azarkinae sp. n. (e–g). a, e male habitus, dorsal; b female habitus, dorsal; c, g palp, retrolateral; d, f palp, ventral. Scale: a–b, e = 2 mm; c–d, f–g = 0.2 mm." httpUri="https://zenodo.org/record/1452197/files/figure.png" pageId="1" pageNumber="272">Fig. 1</figureCitation>
a-b. Total length 6.3. Carapace 3.2 long, 2.55 wide. Coloration: prosoma, chelicerae and palps black. Legs: Fe black, Pt‒Ta yellow-gray. Leg IV darker than others, almost black. Abdomen black. Spination of leg I: Fe d1-1-1 p0-0-2 r0-1-1; Ti p1-0-0 r1-0-0
<date id="FFB47D8C994170542A55FAD3947008B5" box="[1289,1343,1387,1411]" pageId="1" pageNumber="272">v2-2</date>
-2-2; Mt p1-1-0 r1-1-0
<date id="FFB47D8C994170542E69FA37903F0891" box="[309,368,1423,1447]" pageId="1" pageNumber="272">v2-2.</date>
</paragraph>
<caption id="DF750BC4994170542FCBFA6E92E408DB" box="[151,939,1493,1518]" pageId="1" pageNumber="272">
<paragraph id="8BB55B4C994170542FCBFA6E92E408DB" blockId="1.[151,939,1493,1518]" box="[151,939,1493,1518]" pageId="1" pageNumber="272">
<emphasis id="B97E875E994170542FCBFA6E904508DB" bold="true" box="[151,266,1494,1517]" pageId="1" pageNumber="272">TABLE 1.</emphasis>
Leg measurements of the
<taxonomicName id="4C0A20CF994170542D78FA6D926B08DB" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[548,804,1493,1517]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994170542D78FA6D926B08DB" box="[548,804,1493,1517]" italics="true" pageId="1" pageNumber="272">Acantholycosa vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994170542C77FA6E922808DB" box="[811,871,1494,1517]" pageId="1" pageNumber="272" rank="species">
<emphasis id="B97E875E994170542C77FA6E922808DB" bold="true" box="[811,871,1494,1517]" pageId="1" pageNumber="272">sp. n.</emphasis>
</taxonomicNameLabel>
male:
</paragraph>
</caption>
<paragraph id="8BB55B4C994170542E6BF9B295B00BF2" pageId="1" pageNumber="272">
<table id="F90AA9EC99418FAA2FC3F9B195B00BF2" box="[159,1279,1545,1732]" gridcols="7" gridrows="5" pageId="1" pageNumber="272" rowsContinueIn="2.[159,1287,378,566]">
<tr id="353A590E99418FAA2FC3F9B195B00B29" box="[159,1279,1545,1567]" gridrow="0" pageId="1" pageNumber="272" rowspan-0="1">
<th id="76EB307299418FAA2E6BF9B1902C0B29" box="[311,355,1545,1567]" gridcol="1" gridrow="0" pageId="1" pageNumber="272">Fe</th>
<th id="76EB307299418FAA2EB1F9B193420B29" box="[493,525,1545,1567]" gridcol="2" gridrow="0" pageId="1" pageNumber="272">Pt</th>
<th id="76EB307299418FAA2DDCF9B193E30B29" box="[640,684,1545,1567]" gridcol="3" gridrow="0" pageId="1" pageNumber="272">Ti</th>
<th id="76EB307299418FAA2C19F9B1923E0B29" box="[837,881,1545,1567]" gridcol="4" gridrow="0" pageId="1" pageNumber="272">Mt</th>
<th id="76EB307299418FAA2B4CF9B195730B29" box="[1040,1084,1545,1567]" gridcol="5" gridrow="0" pageId="1" pageNumber="272">Ta</th>
<th id="76EB307299418FAA2B9AF9B195B00B29" box="[1222,1279,1545,1567]" gridcol="6" gridrow="0" pageId="1" pageNumber="272">Total</th>
</tr>
<tr id="353A590E99418FAA2FC3F98A95B00B7E" box="[159,1279,1586,1608]" gridrow="1" pageId="1" pageNumber="272">
<th id="76EB307299418FAA2FC3F98A91F50B7E" box="[159,186,1586,1608]" gridcol="0" gridrow="1" pageId="1" pageNumber="272">I</th>
<td id="76EB307299418FAA2E6BF98A902C0B7E" box="[311,355,1586,1608]" gridcol="1" gridrow="1" pageId="1" pageNumber="272">3.1</td>
<td id="76EB307299418FAA2EB1F98A93420B7E" box="[493,525,1586,1608]" gridcol="2" gridrow="1" pageId="1" pageNumber="272">1.3</td>
<td id="76EB307299418FAA2DDCF98A93E30B7E" box="[640,684,1586,1608]" gridcol="3" gridrow="1" pageId="1" pageNumber="272">3.0</td>
<td id="76EB307299418FAA2C19F98A923E0B7E" box="[837,881,1586,1608]" gridcol="4" gridrow="1" pageId="1" pageNumber="272">3.0</td>
<td id="76EB307299418FAA2B4CF98A95730B7E" box="[1040,1084,1586,1608]" gridcol="5" gridrow="1" pageId="1" pageNumber="272">1.5</td>
<td id="76EB307299418FAA2B9AF98A95B00B7E" box="[1222,1279,1586,1608]" gridcol="6" gridrow="1" pageId="1" pageNumber="272">11.9</td>
</tr>
<tr id="353A590E99418FAA2FC3F9E495B00B44" box="[159,1279,1628,1650]" gridrow="2" pageId="1" pageNumber="272">
<th id="76EB307299418FAA2FC3F9E491F50B44" box="[159,186,1628,1650]" gridcol="0" gridrow="2" pageId="1" pageNumber="272">II</th>
<td id="76EB307299418FAA2E6BF9E4902C0B44" box="[311,355,1628,1650]" gridcol="1" gridrow="2" pageId="1" pageNumber="272">3.05</td>
<td id="76EB307299418FAA2EB1F9E493420B44" box="[493,525,1628,1650]" gridcol="2" gridrow="2" pageId="1" pageNumber="272">1.3</td>
<td id="76EB307299418FAA2DDCF9E493E30B44" box="[640,684,1628,1650]" gridcol="3" gridrow="2" pageId="1" pageNumber="272">2.75</td>
<td id="76EB307299418FAA2C19F9E4923E0B44" box="[837,881,1628,1650]" gridcol="4" gridrow="2" pageId="1" pageNumber="272">3.05</td>
<td id="76EB307299418FAA2B4CF9E495730B44" box="[1040,1084,1628,1650]" gridcol="5" gridrow="2" pageId="1" pageNumber="272">1.4</td>
<td id="76EB307299418FAA2B9AF9E495B00B44" box="[1222,1279,1628,1650]" gridcol="6" gridrow="2" pageId="1" pageNumber="272">11.55</td>
</tr>
<tr id="353A590E99418FAA2FC3F93D95B00BAD" box="[159,1279,1669,1691]" gridrow="3" pageId="1" pageNumber="272">
<th id="76EB307299418FAA2FC3F93D91F50BAD" box="[159,186,1669,1691]" gridcol="0" gridrow="3" pageId="1" pageNumber="272">III</th>
<td id="76EB307299418FAA2E6BF93D902C0BAD" box="[311,355,1669,1691]" gridcol="1" gridrow="3" pageId="1" pageNumber="272">2.95</td>
<td id="76EB307299418FAA2EB1F93D93420BAD" box="[493,525,1669,1691]" gridcol="2" gridrow="3" pageId="1" pageNumber="272">1.2</td>
<td id="76EB307299418FAA2DDCF93D93E30BAD" box="[640,684,1669,1691]" gridcol="3" gridrow="3" pageId="1" pageNumber="272">2.45</td>
<td id="76EB307299418FAA2C19F93D923E0BAD" box="[837,881,1669,1691]" gridcol="4" gridrow="3" pageId="1" pageNumber="272">3.45</td>
<td id="76EB307299418FAA2B4CF93D95730BAD" box="[1040,1084,1669,1691]" gridcol="5" gridrow="3" pageId="1" pageNumber="272">1.35</td>
<td id="76EB307299418FAA2B9AF93D95B00BAD" box="[1222,1279,1669,1691]" gridcol="6" gridrow="3" pageId="1" pageNumber="272">11.4</td>
</tr>
<tr id="353A590E99418FAA2FC3F91695B00BF2" box="[159,1279,1710,1732]" gridrow="4" pageId="1" pageNumber="272">
<th id="76EB307299418FAA2FC3F91691F50BF2" box="[159,186,1710,1732]" gridcol="0" gridrow="4" pageId="1" pageNumber="272">IV</th>
<td id="76EB307299418FAA2E6BF916902C0BF2" box="[311,355,1710,1732]" gridcol="1" gridrow="4" pageId="1" pageNumber="272">3.65</td>
<td id="76EB307299418FAA2EB1F91693420BF2" box="[493,525,1710,1732]" gridcol="2" gridrow="4" pageId="1" pageNumber="272">1.3</td>
<td id="76EB307299418FAA2DDCF91693E30BF2" box="[640,684,1710,1732]" gridcol="3" gridrow="4" pageId="1" pageNumber="272">3.25</td>
<td id="76EB307299418FAA2C19F916923E0BF2" box="[837,881,1710,1732]" gridcol="4" gridrow="4" pageId="1" pageNumber="272">4.95</td>
<td id="76EB307299418FAA2B4CF91695730BF2" box="[1040,1084,1710,1732]" gridcol="5" gridrow="4" pageId="1" pageNumber="272">1.8</td>
<td id="76EB307299418FAA2B9AF91695B00BF2" box="[1222,1279,1710,1732]" gridcol="6" gridrow="4" pageId="1" pageNumber="272">14.95</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB55B4C994170542F9BF8BD938A0ABC" blockId="1.[151,1436,1797,2038]" pageId="1" pageNumber="272">
Palp as in
<figureCitation id="133147C9994170542E62F8BD90E10A28" box="[318,430,1797,1822]" captionStart="FIGURE 1" captionStartId="2.[151,250,1972,1994]" captionTargetBox="[152,1436,902,1951]" captionTargetId="figure@2.[150,1436,892,1951]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and male palp of Acantholycosa vahterae sp. n. (a–d) and Mongolicosa azarkinae sp. n. (e–g). a, e male habitus, dorsal; b female habitus, dorsal; c, g palp, retrolateral; d, f palp, ventral. Scale: a–b, e = 2 mm; c–d, f–g = 0.2 mm." httpUri="https://zenodo.org/record/1452197/files/figure.png" pageId="1" pageNumber="272">Figs 1c‒d</figureCitation>
,
<figureCitation id="133147C9994170542EE7F8BE90BC0A28" box="[443,499,1798,1822]" captionStart="FIGURE 2" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[154,1431,327,1881]" captionTargetId="figure@4.[151,1436,318,1881]" captionTargetPageId="4" captionText="FIGURE 2. Male palp of Acantholycosa vahterae sp. n. (a–e) and A. kronestedti sp. n. (f–j). a, f bulb, ventral; b, g bulb, anterior; c, h bulb, retrolateral; d, i tegulum, ventral; e, j tegulum, anterior. Scale = 0.2 mm. Abbreviations: Es basal spine of embolus, Pa palea, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452199/files/figure.png" pageId="1" pageNumber="272">2a‒e</figureCitation>
,
<figureCitation id="133147C9994170542EA3F8BE93780A28" box="[511,567,1798,1822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1892,1914]" captionTargetBox="[151,1436,223,1853]" captionTargetId="figure@5.[151,1436,223,1853]" captionTargetPageId="5" captionText="FIGURE 3. Bulb of Acantholycosa vahterae sp. n. (a–c), A. kronestedti sp. n. (d–f) and Mongolicosa azarkinae sp. n. (g–i, embolic spine broken). a, d, g ventral; b, e, h retrolateral; c, f, i anterior. Scale = 0.1 mm (if indicated). Abbreviations: Em embolus, Es basal spine of embolus, Sp spine-like process of terminal apophysis, Ss small spine in mid part of embolus, Tp triangle outgrowth of paleal apophysis." httpUri="https://zenodo.org/record/1452201/files/figure.png" pageId="1" pageNumber="272">3a‒c</figureCitation>
,
<figureCitation id="133147C9994170542D1FF8BE93350A28" box="[579,634,1798,1822]" captionStart="FIGURE 4" captionStartId="6.[151,250,1172,1194]" captionTargetBox="[151,1436,208,1138]" captionTargetId="figure@6.[151,1436,208,1138]" captionTargetPageId="6" captionText="FIGURE 4. Embolic division of Acantholycosa vahterae sp. n. (a–c), A. kronestedti sp. n. (d–f) and Mongolicosa azarkinae sp. n. (g–h, embolic spine broken). a, d, g ventral; b, e, h anterior; c, f posterior. Scale = 0.1 mm." httpUri="https://zenodo.org/record/1452203/files/figure.png" pageId="1" pageNumber="272">4a‒c</figureCitation>
,
<figureCitation id="133147C9994170542DDBF8BD93F00A28" box="[647,703,1797,1822]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[155,1428,1290,1929]" captionTargetId="figure@6.[151,1436,1279,1929]" captionTargetPageId="6" captionText="FIGURE 5. Embolic division of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d anterior; b, e retrolateral; c, f ventral. Scale = 0.2 mm." httpUri="https://zenodo.org/record/1452205/files/figure.png" pageId="1" pageNumber="272">5a‒c</figureCitation>
. Tibia 2 times longer than wide. Cymbium 1.5 times longer than tibia. Tegular apophysis without anterior arm. Embolus broad, with a large basal spine (
<emphasis id="B97E875E994170542BFFF89295F00A77" box="[1187,1215,1834,1857]" italics="true" pageId="1" pageNumber="272">Es</emphasis>
) similar to a large triangular lamina, embolic tip slightly widened. Paleal apophysis large, with an anterior triangular outgrowth (
<emphasis id="B97E875E994170542A31F8F694C50A53" box="[1389,1418,1870,1893]" italics="true" pageId="1" pageNumber="272">Tp</emphasis>
); terminal apophysis with a spine-like process (
<emphasis id="B97E875E994170542DC4F8CA93FB0ABF" box="[664,692,1906,1929]" italics="true" pageId="1" pageNumber="272">Sp</emphasis>
).
</paragraph>
<paragraph id="8BB55B4C994170542F9BF82D91B10AC0" blockId="1.[151,1436,1797,2038]" pageId="1" pageNumber="272">
Female. Total length 7.2. Carapace 3.7 long, 2.9 wide. Coloration: prosoma, palps and legs black. Chelicerae dark brown. Abdomen black. Spination of leg I: Fe d1-1-1 p0-0-2 r1-0-1; Ti p1-0-0 r1-0-0
<date id="FFB47D8C994170542BCDF80295880AE4" box="[1169,1223,1978,2002]" pageId="1" pageNumber="272">v2-2</date>
-2-2; Mt p0-1-0 r0- 1-0
<date id="FFB47D8C994170542F9FF86691B10AC0" box="[195,254,2014,2038]" pageId="1" pageNumber="272">v2-2.</date>
</paragraph>
<paragraph id="8BB55B4C994270572F9BFF2F92CE0C2B" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="273">
Epigyne as in
<figureCitation id="133147C9994270572E31FF2F90930D86" box="[365,476,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="2" pageNumber="273">Fig. 6a‒c</figureCitation>
. Fovea large, diamond-shaped (
<figureCitation id="133147C9994270572C14FF2F92D50D86" box="[840,922,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="2" pageNumber="273">Fig. 6a</figureCitation>
) or triangular (
<figureCitation id="133147C9994270572B17FF2F95EE0D86" box="[1099,1185,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1374,1396]" captionTargetBox="[151,1436,483,1351]" captionTargetId="figure@8.[151,1436,483,1352]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne of Acantholycosa vahterae sp. n. (a–c) and A. kronestedti sp. n. (d–f). a, d intact, ventral; b, e macerated, ventral; c, f macerated, dorsal. Scale = 0.2 mm. Abbreviations: Ap anterior pocket, Cd copulatory ducts, Re receptacle, Se septum, Sl septal lobes." httpUri="https://zenodo.org/record/1452207/files/figure.png" pageId="2" pageNumber="273">Fig. 6b</figureCitation>
), not entirely covered with septum. Septum long with parallel lateral margins, not widened in posterior part, with large lateral lobes (
<emphasis id="B97E875E994270572A2AFF0494C30DE3" box="[1398,1420,188,213]" italics="true" pageId="2" pageNumber="273">Sl</emphasis>
), lobes wider than septum itself. Anterior pocket (
<emphasis id="B97E875E994270572D98FF5893AC0DC1" box="[708,739,224,247]" italics="true" pageId="2" pageNumber="273">Ap</emphasis>
) single, arch-shaped. Receptacles (
<emphasis id="B97E875E994270572B24FF5995DA0DCE" box="[1144,1173,225,248]" italics="true" pageId="2" pageNumber="273">Re</emphasis>
) as long as copulatory ducts (
<emphasis id="B97E875E994270572FBEFEBC904C0C2B" box="[226,259,260,285]" italics="true" pageId="2" pageNumber="273">Cd</emphasis>
), reaching the middle part of epigynal plate, converging.
</paragraph>
<paragraph id="8BB55B4C994270572FCBFEF095DF0F81" blockId="2.[151,959,328,350]" lastBlockId="2.[151,1436,634,839]" pageId="2" pageNumber="273">
<emphasis id="B97E875E994270572FCBFEF090460C69" bold="true" box="[151,265,328,351]" pageId="2" pageNumber="273">TABLE 2.</emphasis>
Leg measurements of the
<taxonomicName id="4C0A20CF994270572D78FEFF926B0C69" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[548,804,327,351]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="273" phylum="Arthropoda" rank="species" species="vahterae" status="sp. nov.">
<emphasis id="B97E875E994270572D78FEFF926B0C69" box="[548,804,327,351]" italics="true" pageId="2" pageNumber="273">Acantholycosa vahterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D3A25994270572C77FEF092280C69" box="[811,871,328,351]" pageId="2" pageNumber="273" rank="species">
<emphasis id="B97E875E994270572C77FEF092280C69" bold="true" box="[811,871,328,351]" pageId="2" pageNumber="273">sp. n.</emphasis>
</taxonomicNameLabel>
female:
<emphasis id="B97E875E994270572F9BFDC290390FA5" bold="true" box="[199,374,634,659]" pageId="2" pageNumber="273">Size variation.</emphasis>
Males vary from
<quantity id="4CF2F6A9994270572D0CFDC293AC0FA5" box="[592,739,634,659]" metricMagnitude="-1" metricUnit="m" metricValue="1.6637" metricValueMax="1.7526" metricValueMin="1.5748" pageId="2" pageNumber="273" unit="in" value="6.55" valueMax="6.9" valueMin="6.2">6.2 to 6.9 in</quantity>
body length, carapace 3.2‒3.6 long, 2.55‒2.85 wide (n=3). Females vary from
<quantity id="4CF2F6A9994270572E2DFD2690B70F81" box="[369,504,670,695]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="1.8796" metricValueMin="1.6764" pageId="2" pageNumber="273" unit="in" value="7.0" valueMax="7.4" valueMin="6.6">6.6 to 7.4 in</quantity>
body length, carapace 3.3‒3.7 long, 2.65‒2.95 wide (n=3).
</paragraph>
<paragraph id="8BB55B4C994270572E79FEC394480F00" pageId="2" pageNumber="273">
<table id="F90AA9EC99428FAA2FC3FEC294480F00" box="[159,1287,378,566]" gridcols="7" gridrows="5" pageId="2" pageNumber="273" rowsContinueFrom="1.[159,1279,1545,1732]">
<tr id="353A590E99428FAA2FC3FEC294480CA6" box="[159,1287,378,400]" gridrow="0" pageId="2" pageNumber="273" rowspan-0="1">
<th id="76EB307299428FAA2E79FEC2901E0CA6" box="[293,337,378,400]" gridcol="1" gridrow="0" pageId="2" pageNumber="273">Fe</th>
<th id="76EB307299428FAA2EBCFEC293430CA6" box="[480,524,378,400]" gridcol="2" gridrow="0" pageId="2" pageNumber="273">Pt</th>
<th id="76EB307299428FAA2DCAFEC2938D0CA6" box="[662,706,378,400]" gridcol="3" gridrow="0" pageId="2" pageNumber="273">Ti</th>
<th id="76EB307299428FAA2C3DFEC292C20CA6" box="[865,909,378,400]" gridcol="4" gridrow="0" pageId="2" pageNumber="273">Mt</th>
<th id="76EB307299428FAA2B4BFEC2950C0CA6" box="[1047,1091,378,400]" gridcol="5" gridrow="0" pageId="2" pageNumber="273">Ta</th>
<th id="76EB307299428FAA2B92FEC294480CA6" box="[1230,1287,378,400]" gridcol="6" gridrow="0" pageId="2" pageNumber="273">Total</th>
</tr>
<tr id="353A590E99428FAA2FC3FE1C94480C8C" box="[159,1287,420,442]" gridrow="1" pageId="2" pageNumber="273">
<th id="76EB307299428FAA2FC3FE1C91F50C8C" box="[159,186,420,442]" gridcol="0" gridrow="1" pageId="2" pageNumber="273">I</th>
<td id="76EB307299428FAA2E79FE1C901E0C8C" box="[293,337,420,442]" gridcol="1" gridrow="1" pageId="2" pageNumber="273">3.25</td>
<td id="76EB307299428FAA2EBCFE1C93430C8C" box="[480,524,420,442]" gridcol="2" gridrow="1" pageId="2" pageNumber="273">1.45</td>
<td id="76EB307299428FAA2DCAFE1C938D0C8C" box="[662,706,420,442]" gridcol="3" gridrow="1" pageId="2" pageNumber="273">2.9</td>
<td id="76EB307299428FAA2C3DFE1C92C20C8C" box="[865,909,420,442]" gridcol="4" gridrow="1" pageId="2" pageNumber="273">2.65</td>
<td id="76EB307299428FAA2B4BFE1C950C0C8C" box="[1047,1091,420,442]" gridcol="5" gridrow="1" pageId="2" pageNumber="273">1.4</td>
<td id="76EB307299428FAA2B92FE1C94480C8C" box="[1230,1287,420,442]" gridcol="6" gridrow="1" pageId="2" pageNumber="273">11.65</td>
</tr>
<tr id="353A590E99428FAA2FC3FE7594480CD5" box="[159,1287,461,483]" gridrow="2" pageId="2" pageNumber="273">
<th id="76EB307299428FAA2FC3FE7591F50CD5" box="[159,186,461,483]" gridcol="0" gridrow="2" pageId="2" pageNumber="273">II</th>
<td id="76EB307299428FAA2E79FE75901E0CD5" box="[293,337,461,483]" gridcol="1" gridrow="2" pageId="2" pageNumber="273">3.25</td>
<td id="76EB307299428FAA2EBCFE7593430CD5" box="[480,524,461,483]" gridcol="2" gridrow="2" pageId="2" pageNumber="273">1.45</td>
<td id="76EB307299428FAA2DCAFE75938D0CD5" box="[662,706,461,483]" gridcol="3" gridrow="2" pageId="2" pageNumber="273">2.75</td>
<td id="76EB307299428FAA2C3DFE7592C20CD5" box="[865,909,461,483]" gridcol="4" gridrow="2" pageId="2" pageNumber="273">2.7</td>
<td id="76EB307299428FAA2B4BFE75950C0CD5" box="[1047,1091,461,483]" gridcol="5" gridrow="2" pageId="2" pageNumber="273">1.45</td>
<td id="76EB307299428FAA2B92FE7594480CD5" box="[1230,1287,461,483]" gridcol="6" gridrow="2" pageId="2" pageNumber="273">11.6</td>
</tr>
<tr id="353A590E99428FAA2FC3FE4E94480F3A" box="[159,1287,502,524]" gridrow="3" pageId="2" pageNumber="273">
<th id="76EB307299428FAA2FC3FE4E91F50F3A" box="[159,186,502,524]" gridcol="0" gridrow="3" pageId="2" pageNumber="273">III</th>
<td id="76EB307299428FAA2E79FE4E901E0F3A" box="[293,337,502,524]" gridcol="1" gridrow="3" pageId="2" pageNumber="273">3.15</td>
<td id="76EB307299428FAA2EBCFE4E93430F3A" box="[480,524,502,524]" gridcol="2" gridrow="3" pageId="2" pageNumber="273">1.3</td>
<td id="76EB307299428FAA2DCAFE4E938D0F3A" box="[662,706,502,524]" gridcol="3" gridrow="3" pageId="2" pageNumber="273">2.55</td>
<td id="76EB307299428FAA2C3DFE4E92C20F3A" box="[865,909,502,524]" gridcol="4" gridrow="3" pageId="2" pageNumber="273">3.2</td>
<td id="76EB307299428FAA2B4BFE4E950C0F3A" box="[1047,1091,502,524]" gridcol="5" gridrow="3" pageId="2" pageNumber="273">1.45</td>
<td id="76EB307299428FAA2B92FE4E94480F3A" box="[1230,1287,502,524]" gridcol="6" gridrow="3" pageId="2" pageNumber="273">11.65</td>
</tr>
<tr id="353A590E99428FAA2FC3FD9894480F00" box="[159,1287,544,566]" gridrow="4" pageId="2" pageNumber="273">
<th id="76EB307299428FAA2FC3FD9891F50F00" box="[159,186,544,566]" gridcol="0" gridrow="4" pageId="2" pageNumber="273">IV</th>
<td id="76EB307299428FAA2E79FD98901E0F00" box="[293,337,544,566]" gridcol="1" gridrow="4" pageId="2" pageNumber="273">4.1</td>
<td id="76EB307299428FAA2EBCFD9893430F00" box="[480,524,544,566]" gridcol="2" gridrow="4" pageId="2" pageNumber="273">1.5</td>
<td id="76EB307299428FAA2DCAFD98938D0F00" box="[662,706,544,566]" gridcol="3" gridrow="4" pageId="2" pageNumber="273">3.55</td>
<td id="76EB307299428FAA2C3DFD9892C20F00" box="[865,909,544,566]" gridcol="4" gridrow="4" pageId="2" pageNumber="273">5.1</td>
<td id="76EB307299428FAA2B4BFD98950C0F00" box="[1047,1091,544,566]" gridcol="5" gridrow="4" pageId="2" pageNumber="273">3.9</td>
<td id="76EB307299428FAA2B92FD9894480F00" box="[1230,1287,544,566]" gridcol="6" gridrow="4" pageId="2" pageNumber="273">18.15</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB55B4C994270572F9BFD7A928B0FC9" blockId="2.[151,1436,634,839]" pageId="2" pageNumber="273">
<emphasis id="B97E875E994270572F9BFD7A906C0FED" bold="true" box="[199,291,706,731]" pageId="2" pageNumber="273">Habitat</emphasis>
. Xeromorphic mountain stony tundra with screes and rocks, situated on the flat top of Cakhir- Khaalgyn-Nuruu Mt. Range, at the elevations of
<quantity id="4CF2F6A9994270572DE7FD5E92190FC8" box="[699,854,742,767]" metricMagnitude="3" metricUnit="m" metricValue="3.4799999999999995" metricValueMax="3.66" metricValueMin="3.3" pageId="2" pageNumber="273" unit="m" value="3480.0" valueMax="3660.0" valueMin="3300.0">3300‒3660 m</quantity>
(
<figureCitation id="133147C9994270572C39FD5E92F80FC9" box="[869,951,742,767]" captionStart="FIGURE 9" captionStartId="11.[151,250,1734,1756]" captionTargetBox="[150,1436,267,1717]" captionTargetId="figure@11.[150,1436,267,1717]" captionTargetPageId="11" captionText="FIGURE 9. Habitats of Acantholycosa vahterae sp. n. (a), A. kronestedti sp. n. (b), Mongolicosa azarkinae sp. n. (c), M. ozkutuki sp. n. (d) and M. cherepanovi sp. n. (e–f)." httpUri="https://zenodo.org/record/1452213/files/figure.png" pageId="2" pageNumber="273">Fig. 9a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31008C7994270572F9BFCB2936B0E71" pageId="2" pageNumber="273" type="distribution">
<paragraph id="8BB55B4C994270572F9BFCB2936B0E71" blockId="2.[151,1436,634,839]" pageId="2" pageNumber="273">
<emphasis id="B97E875E994270572F9BFCB290160E15" bold="true" box="[199,345,778,803]" pageId="2" pageNumber="273">Distribution</emphasis>
. Known from the
<typeStatus id="54B1E5EE994270572D6AFCB393290E15" box="[566,614,779,803]" pageId="2" pageNumber="273">type</typeStatus>
locality only. Cakhir-Khaalgyn-Nuruu Mt. Range belongs to the main range of Mongolian
<collectingRegion id="49CE95AE994270572E20FC9690FB0E71" box="[380,436,814,839]" country="Russia" name="Altay" pageId="2" pageNumber="273">Altai</collectingRegion>
(
<figureCitation id="133147C9994270572E98FC9693580E71" box="[452,535,814,839]" captionStart="FIGURE 10" captionStartId="12.[151,250,1631,1653]" captionTargetBox="[153,1437,1254,1608]" captionTargetId="figure@12.[151,1437,1254,1609]" captionTargetPageId="12" captionText="FIGURE 10. Collecting localities of Acantholycosa vahterae sp. n. (5), A. kronestedti sp. n. (4), Mongolicosa azarkinae sp. n. (3), M. cherepanovi sp. n. (1) and M. ozkutuki sp. n. (2)." httpUri="https://zenodo.org/record/1452215/files/figure.png" pageId="2" pageNumber="273">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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