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<mods:title id="90283F5429575BB56E58423C395C6873">A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex</mods:title>
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<treatment id="038413308A1DFFEC4FA896241431FD72" ID-DOI="http://doi.org/10.5281/zenodo.6054065" ID-GBIF-Taxon="116973314" ID-Zenodo-Dep="6054065" LSID="urn:lsid:plazi:treatment:038413308A1DFFEC4FA896241431FD72" httpUri="http://treatment.plazi.org/id/038413308A1DFFEC4FA896241431FD72" lastPageNumber="348" pageId="5" pageNumber="348">
<subSubSection id="C337F1AD8A1DFFEC4FA8962414A3FDA6" pageId="5" pageNumber="348" type="nomenclature">
<paragraph id="8B92A2268A1DFFEC4FA8962415C7FE3E" blockId="5.[151,685,368,394]" box="[151,685,368,394]" pageId="5" pageNumber="348">
<heading id="D0DA154A8A1DFFEC4FA8962415C7FE3E" bold="true" box="[151,685,368,394]" fontSize="11" level="1" pageId="5" pageNumber="348" reason="1">
<emphasis id="B9597E348A1DFFEC4FA8962415C7FE3E" bold="true" box="[151,685,368,394]" pageId="5" pageNumber="348">
Tribe
<taxonomicName id="4C2DD9A58A1DFFEC4FDD96241557FE3E" ID-CoL="KZX" authority="Forel 1912" authorityName="Forel" authorityYear="1912" box="[226,573,368,394]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Myrmelachistini">
Myrmelachistini
<bibRefCitation id="EFBCDFD78A1DFFEC4E8796241557FE3E" author="Forel" box="[440,573,368,394]" pageId="5" pageNumber="348" refString="Forel, A. (1912) Formicides neotropiques. Part VI. 5 me sous-famille Camponotinae Forel. Memoires de la Societe Entomologique de Belgique, 20, 59 - 92." type="journal article" year="1912">Forel 1912</bibRefCitation>
</taxonomicName>
stat. rev.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B92A2268A1DFFEC4FA896E31547FE78" blockId="5.[151,557,438,460]" box="[151,557,438,460]" pageId="5" pageNumber="348">
=
<taxonomicName id="4C2DD9A58A1DFFEC4F9396E2168BFE78" authority="Emery 1925" authorityName="Emery" authorityYear="1925" box="[172,481,438,460]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Brachymyrmicini">
Brachymyrmicini
<bibRefCitation id="EFBCDFD78A1DFFEC4E5B96E3168BFE78" author="Emery" box="[356,481,438,460]" pageId="5" pageNumber="348" refString="Emery, C. (1925) Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum, 183, 1 - 302." type="journal article" year="1925">Emery 1925</bibRefCitation>
</taxonomicName>
<emphasis id="B9597E348A1DFFEC4ED896E31547FE78" bold="true" box="[487,557,439,460]" pageId="5" pageNumber="348">syn. n.</emphasis>
</paragraph>
<paragraph id="8B92A2268A1DFFEC4FA896AD14A3FDA6" blockId="5.[151,1436,505,710]" box="[151,969,505,530]" pageId="5" pageNumber="348">
<emphasis id="B9597E348A1DFFEC4FA896AD1793FDA6" bold="true" box="[151,249,505,530]" pageId="5" pageNumber="348">Genera:</emphasis>
<taxonomicName id="4C2DD9A58A1DFFEC4E3F96AD16C0FDA6" box="[256,426,505,530]" class="Insecta" family="Formicidae" genus="Brachymyrmex" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B9597E348A1DFFEC4E3F96AD16C0FDA6" box="[256,426,505,530]" italics="true" pageId="5" pageNumber="348">Brachymyrmex</emphasis>
</taxonomicName>
(tribal transfer),
<taxonomicName id="4C2DD9A58A1DFFEC4D5796AD147AFDA6" box="[616,784,505,530]" class="Insecta" family="Formicidae" genus="Myrmelachista" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B9597E348A1DFFEC4D5796AD147AFDA6" box="[616,784,505,530]" italics="true" pageId="5" pageNumber="348">Myrmelachista</emphasis>
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(tribal transfer).
</paragraph>
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<subSubSection id="C337F1AD8A1DFFEC4FF895491431FD72" pageId="5" pageNumber="348" type="discussion">
<paragraph id="8B92A2268A1DFFEC4FF895491431FD72" blockId="5.[151,1436,505,710]" pageId="5" pageNumber="348">
<emphasis id="B9597E348A1DFFEC4FF895491625FD82" bold="true" box="[199,335,541,566]" pageId="5" pageNumber="348">Comments.</emphasis>
These two genera form a robustly supported clade that is sister to all other
<taxonomicName id="4C2DD9A58A1DFFEC4BA295491277FD82" box="[1181,1309,541,566]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
(
<figureCitation id="1316BEA38A1DFFEC4A13954912E1FD82" box="[1324,1419,541,566]" captionStart="FIGURE 1" captionStartId="3.[151,250,1698,1720]" captionTargetBox="[164,1419,191,1667]" captionTargetId="figure@3.[156,1432,185,1676]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of the ant subfamily Formicinae based on 959 UCE (ultraconserved element) loci (Blaimer et al. 2015). Support values are maximum likelihood bootstrap percentages. The tree depicts six major clades, here treated as tribes, and five isolated genera subtended by long branches (Gesomyrmex, Gigantiops, Myrmoteras, Oecophylla, and Santschiella, in bold font). Note that species assigned to Camponotus occur as three separate lineages within the tribe Camponotini, and that Phasmomyrmex and Forelophilus are embedded within Camponotus (sensu stricto)." httpUri="https://zenodo.org/record/264544/files/figure.png" pageId="5" pageNumber="348">Figure 1</figureCitation>
). Shared worker characters include a reduced antennal count (910 segments), five mandibular teeth, petiole inclined anteriorly and/or with long posterior peduncle, and anterior tergosternal fusion of the third abdominal segment (
<bibRefCitation id="EFBCDFD78A1DFFEC4FA195DD1658FD16" author="Bolton" box="[158,306,649,674]" pageId="5" pageNumber="348" refString="Bolton, B. (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute, 71, 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
). Both genera have the plesiomorphic palp formula 6,4 but this has been reduced to five maxillary palp segments in some
<taxonomicName id="4C2DD9A58A1DFFEC4EA395F9152EFD72" box="[412,580,685,710]" class="Insecta" family="Formicidae" genus="Myrmelachista" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B9597E348A1DFFEC4EA395F9152EFD72" box="[412,580,685,710]" italics="true" pageId="5" pageNumber="348">Myrmelachista</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFBCDFD78A1DFFEC4D9595F91425FD72" author="Longino" box="[682,847,685,710]" pageId="5" pageNumber="348" refString="Longino, J. T. (2006) A taxonomic review of the genus Myrmelachista (Hymenoptera: Formicidae) in Costa Rica. Zootaxa, 1141, 1 - 54." type="journal article" year="2006">Longino 2006</bibRefCitation>
).
</paragraph>
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</treatment>
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