406 lines
51 KiB
XML
406 lines
51 KiB
XML
<document id="83EC37A986BAF099082795DA553AAAB3" ID-DOI="10.11646/zootaxa.3734.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5275595" ID-ZooBank="36ED88E6-2CEB-4071-8429-A39901B8B9BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629926016628" checkinUser="felipe" docAuthor="Simaiakis, Stylianos Michail & Edgecombe, Gregory D." docDate="2013" docId="03B68458FFB8FFFDFF56A8FBFEE8FC66" docLanguage="en" docName="zt03734p198.pdf" docOrigin="Zootaxa 3734 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus (Parotostigmus) coltellus Simaiakis & Edgecombe 2013, n. sp." docType="treatment" docVersion="3" lastPageNumber="179" masterDocId="FF8FFC20FFB0FFF7FFC1AA2DFF95FFA1" masterDocTitle="Scolopendromorph centipedes (Chilopoda: Scolopendromorpha) in the Natural History Museum (London): A review of the hitherto unidentified species collected in Africa, with remarks on taxonomy and distribution, and a new species of Otostigmus (Parotostigmus)" masterLastPageNumber="198" masterPageNumber="169" pageNumber="177" updateTime="1699128448735" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5F44C4548268E98D846FCDB7326054F6">
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<mods:title id="CC5E38342FF8DF2DC824F50250FEEED8">Scolopendromorph centipedes (Chilopoda: Scolopendromorpha) in the Natural History Museum (London): A review of the hitherto unidentified species collected in Africa, with remarks on taxonomy and distribution, and a new species of Otostigmus (Parotostigmus)</mods:title>
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</mods:titleInfo>
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<mods:name id="8D63792354619DAB2C2E5A9A69FEBA85" type="personal">
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<mods:role id="D825088F56211694858B9B9A226B4717">
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<mods:roleTerm id="ED33264D14F3E266D6F08F8E09DAED50">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A90C1E90B393FDB9371EE5C970B79CFC">Simaiakis, Stylianos Michail</mods:namePart>
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<mods:affiliation id="C343F52AA482C864EC9D1E33F5CA6BB5">Natural History Museum of Crete, University of Crete, Knossos Av., Herakleion 71409, Crete, Greece.</mods:affiliation>
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<mods:nameIdentifier id="A54C751FC1923A833B4012C7F67B0405" type="email">ssimaiakis@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="7202C9C4976B2B64C321091A42854017" type="personal">
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<mods:roleTerm id="F9AD640512B444640B1B2C0E3A988308">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EFC07BD5FDCAF8758D17F47C278F60CA">Edgecombe, Gregory D.</mods:namePart>
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<mods:affiliation id="94F77191768E5711F06C9B35BA64E8E8">Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:typeOfResource id="AF9ABD5C7846BAFBE865F040B6F34F30">text</mods:typeOfResource>
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<mods:titleInfo id="D00BF5693DCCBF85EDC9FB3F91608E9B">
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<mods:title id="72390930687F84E2C2512BACCD672489">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FC52D60FE12E106E4B887A511C9541F2">
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<mods:date id="C57AF0A2231CA7460FCC7BF535B2FE28">2013</mods:date>
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<mods:number id="211D4E436529FB8AB94FA4E8396B7E5B">2013-11-05</mods:number>
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<mods:number id="BC564C47C67A346174790EF6A7F2FA53">3734</mods:number>
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<mods:detail id="16420AF1C7E6B0DC4D7F3014AD10021A" type="issue">
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<mods:number id="276C6312D392544C90CFEE89872C1ECA">2</mods:number>
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<mods:start id="FD465B8A84F0D5DE0FFC9F6BBD0F4A40">169</mods:start>
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<mods:classification id="59B2CA0F6C37B74E9A0EDF0B78443AC1">journal article</mods:classification>
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<mods:identifier id="0492586A9DC9C89263ED87B399AC2096" type="DOI">10.11646/zootaxa.3734.2.5</mods:identifier>
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<mods:identifier id="A95E7313B428C24232A366AA44E0F833" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B68458FFB8FFFDFF56A8FBFEE8FC66" LSID="urn:lsid:plazi:treatment:03B68458FFB8FFFDFF56A8FBFEE8FC66" httpUri="http://treatment.plazi.org/id/03B68458FFB8FFFDFF56A8FBFEE8FC66" lastPageId="10" lastPageNumber="179" pageId="8" pageNumber="177">
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<subSubSection id="C30566C5FFB8FFFFFF56A8FBFD36FD4E" box="[151,675,726,751]" pageId="8" pageNumber="177" type="nomenclature">
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<paragraph id="8BA0354EFFB8FFFFFF56A8FBFD36FD4E" blockId="8.[151,675,726,787]" box="[151,675,726,751]" pageId="8" pageNumber="177">
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<heading id="D0E88222FFB8FFFFFF56A8FBFD36FD4E" bold="true" box="[151,675,726,751]" fontSize="10" level="2" pageId="8" pageNumber="177" reason="2">
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<emphasis id="B96BE95CFFB8FFFFFF56A8FBFD36FD4E" bold="true" box="[151,675,726,751]" pageId="8" pageNumber="177">
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8.
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<taxonomicName id="4C1F4ECDFFB8FFFFFF72A8FBFDC8FD4E" authority="Simaiakis & Edgecombe, 2013" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[179,605,726,751]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="coltellus" status="sp. nov." subGenus="Parotostigmus">
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<emphasis id="B96BE95CFFB8FFFFFF72A8FBFEA3FD4E" bold="true" box="[179,310,726,751]" italics="true" pageId="8" pageNumber="177">Otostigmus</emphasis>
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(
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<emphasis id="B96BE95CFFB8FFFFFE87A8FBFE79FD4E" bold="true" box="[326,492,726,751]" italics="true" pageId="8" pageNumber="177">Parotostigmus</emphasis>
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)
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<emphasis id="B96BE95CFFB8FFFFFE3DA8FBFDC8FD4E" bold="true" box="[508,605,726,751]" italics="true" pageId="8" pageNumber="177">coltellus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2585427FFB8FFFFFDA5A8FBFD36FD4E" box="[612,675,726,751]" pageId="8" pageNumber="177" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30566C5FFB8FFFFFF56A8D7FE8DFCB2" box="[151,280,762,787]" pageId="8" pageNumber="177" type="description">
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<paragraph id="8BA0354EFFB8FFFFFF56A8D7FE8DFCB2" blockId="8.[151,675,726,787]" box="[151,280,762,787]" pageId="8" pageNumber="177">
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||
(
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<figureCitation id="132429CBFFB8FFFFFF5EA8D7FE85FCB2" box="[159,272,762,787]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 9–14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30566C5FFB8FFFFFF56A96FFDCFFCD6" pageId="8" pageNumber="177" type="reference_group">
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<paragraph id="8BA0354EFFB8FFFFFF56A96FFD6EFCF9" blockId="8.[151,763,834,887]" box="[151,763,834,856]" pageId="8" pageNumber="177">
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<taxonomicName id="4C1F4ECDFFB8FFFFFF56A96FFE7FFCF9" ID-CoL="4B7XS" authority="Karsch, 1884" authorityName="Karsch" authorityYear="1884" box="[151,490,834,856]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="inermis">
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<emphasis id="B96BE95CFFB8FFFFFF56A96FFEC9FCF9" box="[151,348,834,856]" italics="true" pageId="8" pageNumber="177">Otostigmus inermis</emphasis>
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<bibRefCitation id="EF8E48BFFFB8FFFFFEA3A96FFE7FFCF9" author="Karsch, F." box="[354,490,834,856]" pageId="8" pageNumber="177" pagination="55 - 59" refId="ref17220" refString="Karsch, F. (1884) Myriapoda. In: Greef, R. Ueber die Fauna der Guinea-Inseln S. Thome und Rolas. Sitzungsberichte der Gesellschaft zur Beforderung der gesammten Naturwissenschaften zu Marburg, 2, 55 - 59." type="journal article" year="1884">Karsch, 1884</bibRefCitation>
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||
</taxonomicName>
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||
[non
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||
<bibRefCitation id="EF8E48BFFFB8FFFFFDE5A96EFD0EFCF9" author="Porat, C. O. von" box="[548,667,834,856]" pageId="8" pageNumber="177" pagination="1 - 48" refId="ref19054" refString="Porat, C. O. von (1876) Om nagra exotiska myriopoder. Bihang till Kongliga Svenska Vetenskaps-Akademien Handlingar, 4, 7, 1 - 48." type="journal article" year="1876">Porat, 1876</bibRefCitation>
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] [partim]
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</paragraph>
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||
<paragraph id="8BA0354EFFB8FFFFFF56A94CFDCFFCD6" blockId="8.[151,763,834,887]" box="[151,602,865,887]" pageId="8" pageNumber="177">
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||
<taxonomicName id="4C1F4ECDFFB8FFFFFF56A94CFD94FCD6" authority="Karsch, 1888" authorityName="Karsch" authorityYear="1888" box="[151,513,865,887]" class="Chilopoda" family="Scolopendridae" genus="Otostigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="productum">
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<emphasis id="B96BE95CFFB8FFFFFF56A94CFEE6FCD6" box="[151,371,865,887]" italics="true" pageId="8" pageNumber="177">Otostigma productum</emphasis>
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||
<bibRefCitation id="EF8E48BFFFB8FFFFFEBBA94CFD94FCD6" author="Karsch, F." box="[378,513,865,887]" pageId="8" pageNumber="177" pagination="29 - 31" refId="ref17263" refString="Karsch, F. (1888) Zwei neue Myriopoden von Ecuador. Berliner Entomologische Zeitschrift, 32, 29 - 31." type="journal article" year="1888">Karsch, 1888</bibRefCitation>
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</taxonomicName>
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||
[partim]
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30566C5FFB8FFFFFF56A989FD57FBA4" pageId="8" pageNumber="177" type="diagnosis">
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||
<paragraph id="8BA0354EFFB8FFFFFF56A989FD57FBA4" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
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||
<emphasis id="B96BE95CFFB8FFFFFF56A989FE84FC1C" bold="true" box="[151,273,932,957]" inLineHeading="true" pageId="8" pageNumber="177" reason="1">Diagnosis.</emphasis>
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||
<taxonomicName id="4C1F4ECDFFB8FFFFFED8A989FDF5FC1C" authority="" box="[281,608,932,957]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Parostostigmus">
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<emphasis id="B96BE95CFFB8FFFFFED8A989FE0CFC1A" box="[281,409,932,955]" italics="true" pageId="8" pageNumber="177">Otostigmus</emphasis>
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||
(
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<emphasis id="B96BE95CFFB8FFFFFE6BA988FDC2FC1D" box="[426,599,933,956]" italics="true" pageId="8" pageNumber="177">Parostostigmus</emphasis>
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)
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</taxonomicName>
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with four glabrous antennal articles; short, blade-like tooth plate lacking teeth, base of tooth plate set off by transverse suture; paramedian sutures complete from T3 or usually T4; short coxopleural process with two small apical spines.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30566C5FFB8FFFFFF06AE22FB25FB34" pageId="8" pageNumber="177" type="materials_examined">
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||
<paragraph id="8BA0354EFFB8FFFFFF06AE22FB25FB34" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
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||
<emphasis id="B96BE95CFFB8FFFFFF06AE22FEE1FB89" bold="true" box="[199,372,1039,1064]" inLineHeading="true" pageId="8" pageNumber="177" reason="1">Type material.</emphasis>
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||
<materialsCitation id="3B773F13FFB8FFFFFEBAAE22FE18FBD1" collectingDate="1949-09-13" collectionCode="BMNH" collectorName="I. L. J. Galbraith" country="Guinea-Bissau" location="Gulf" municipality="Zampalma" pageId="8" pageNumber="177" specimenCount="2" typeStatus="holotype">
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||
<typeStatus id="54A48BECFFB8FFFFFEBAAE22FE75FB89" box="[379,480,1039,1064]" pageId="8" pageNumber="177" type="holotype">Holotype</typeStatus>
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||
(
|
||
<figureCitation id="132429CBFFB8FFFFFE2EAE22FD22FB89" box="[495,695,1039,1064]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 9, 10, 12, 13</figureCitation>
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||
):
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<location id="8EC06395FFB8FFFFFD0AAE22FD6AFB89" LSID="urn:lsid:plazi:treatment:03B68458FFB8FFFDFF56A8FBFEE8FC66:8EC06395FFB8FFFFFD0AAE22FD6AFB89" box="[715,767,1039,1064]" country="Guinea-Bissau" municipality="Zampalma" name="Gulf" pageId="8" pageNumber="177">Gulf</location>
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of
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||
<collectingCountry id="F30875DEFFB8FFFFFCE5AE22FCECFB89" box="[804,889,1039,1064]" name="Guinea-Bissau" pageId="8" pageNumber="177">Guinea</collectingCountry>
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||
:
|
||
<collectingMunicipality id="6BC4AF34FFB8FFFFFC42AE22FC6BFB89" box="[899,1022,1039,1064]" pageId="8" pageNumber="177">Zampalma</collectingMunicipality>
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,
|
||
<collectingCountry id="F30875DEFFB8FFFFFBC8AE22FBECFB89" box="[1033,1145,1039,1064]" name="Sao Tome and Principe" pageId="8" pageNumber="177">São Tomé</collectingCountry>
|
||
Isl. [
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||
<collectingCountry id="F30875DEFFB8FFFFFB70AE22FE56FBEC" name="Sao Tome and Principe" pageId="8" pageNumber="177">Democratic Republic of São Tomé and Principe</collectingCountry>
|
||
],
|
||
<date id="FFA1138EFFB8FFFFFE16AE19FDDEFBEC" box="[471,587,1076,1101]" pageId="8" pageNumber="177" value="1949-09-13">
|
||
<collectingDate id="EFE5EA66FFB8FFFFFE16AE19FDDEFBEC" box="[471,587,1076,1101]" pageId="8" pageNumber="177" value="1949-09-13">13/9/1949</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EA5098FFB8FFFFFD4CAE19FCAAFBEC" box="[653,831,1076,1101]" pageId="8" pageNumber="177">I.L.J. Galbraith</collectorName>
|
||
,
|
||
<collectionCode id="ED0EAD8BFFB8FFFFFC8CAE19FC34FBEA" box="[845,929,1076,1099]" pageId="8" pageNumber="177">BMNH</collectionCode>
|
||
1950.3.7.127-136 (
|
||
<figureCitation id="132429CBFFB8FFFFFB45AE19FB5EFBEC" box="[1156,1227,1076,1101]" captionStart="FIGURE 8" captionStartId="7.[151,249,1625,1647]" captionTargetBox="[161,1425,575,1618]" captionTargetId="figure-252@7.[161,1425,575,1618]" captionTargetPageId="7" captionText="FIGURE 8. Otostigmus (Otostigmus) cuneiventris, O. (Parotostigmus) coltellus n. sp., O. (P.) gemmifer and O. (P.) productus, distribution records of specimens from Africa housed in the NHM collection." figureDoi="http://doi.org/10.5281/zenodo.5275605" httpUri="https://zenodo.org/record/5275605/files/figure.png" pageId="8" pageNumber="177">Fig. 8</figureCitation>
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||
);
|
||
<typeStatus id="54A48BECFFB8FFFFFB24AE19FAD0FBED" box="[1253,1349,1076,1100]" pageId="8" pageNumber="177" type="paratype">paratype</typeStatus>
|
||
female from same collection
|
||
</materialsCitation>
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||
.
|
||
<materialsCitation id="3B773F13FFB8FFFFFE5AAE75FB39FB34" collectionCode="T" collectorName="Godman Exploration" country="Guinea-Bissau" county="Tams" location="Gulf" municipality="Percy Sladen Memorial" pageId="8" pageNumber="177" specimenCode="BMNH 1933.8" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54A48BECFFB8FFFFFE5AAE75FE68FBD1" box="[411,509,1112,1136]" pageId="8" pageNumber="177" type="paratype">Paratype</typeStatus>
|
||
(
|
||
<figureCitation id="132429CBFFB8FFFFFDCEAE7AFD03FBD1" box="[527,662,1111,1136]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 11, 14</figureCitation>
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||
):
|
||
<location id="8EC06395FFB8FFFFFD6FAE7AFD74FBD1" LSID="urn:lsid:plazi:treatment:03B68458FFB8FFFDFF56A8FBFEE8FC66:8EC06395FFB8FFFFFD6FAE7AFD74FBD1" box="[686,737,1111,1136]" country="Guinea-Bissau" county="Tams" municipality="Percy Sladen Memorial" name="Gulf" pageId="8" pageNumber="177">Gulf</location>
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||
of
|
||
<collectingCountry id="F30875DEFFB8FFFFFCCCAE7AFCF7FBD1" box="[781,866,1111,1136]" name="Guinea-Bissau" pageId="8" pageNumber="177">Guinea</collectingCountry>
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||
:
|
||
<collectingCountry id="F30875DEFFB8FFFFFCB1AE7AFC76FBD1" box="[880,995,1111,1136]" name="Sao Tome and Principe" pageId="8" pageNumber="177">São Tomé</collectingCountry>
|
||
Isl.,
|
||
<location id="8EC06395FFB8FFFFFBE3AE75FB3BFBD1" LSID="urn:lsid:plazi:treatment:03B68458FFB8FFFDFF56A8FBFEE8FC66:8EC06395FFB8FFFFFBE3AE75FB3BFBD1" box="[1058,1198,1111,1136]" country="Guinea-Bissau" county="Tams" municipality="Percy Sladen Memorial" name="West Africa" pageId="8" pageNumber="177">West Africa</location>
|
||
, 1932,
|
||
<collectingMunicipality id="6BC4AF34FFB8FFFFFAC7AE75FE90FB34" pageId="8" pageNumber="177">Percy Sladen Memorial</collectingMunicipality>
|
||
and
|
||
<collectorName id="26EA5098FFB8FFFFFEFAAE51FDBFFB34" box="[315,554,1148,1173]" pageId="8" pageNumber="177">Godman Exploration</collectorName>
|
||
, leg. W.H.
|
||
<collectionCode id="ED0EAD8BFFB8FFFFFD61AE51FD27FB32" box="[672,690,1148,1171]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="177">T</collectionCode>
|
||
.
|
||
<collectingCounty id="62C14DC2FFB8FFFFFD7DAE51FD6EFB32" box="[700,763,1148,1171]" pageId="8" pageNumber="177">Tams</collectingCounty>
|
||
,
|
||
<specimenCount id="9D19FEC7FFB8FFFFFCC6AE51FCAAFB32" box="[775,831,1148,1173]" pageId="8" pageNumber="177" type="generic">1 ex.</specimenCount>
|
||
,
|
||
<specimenCode id="DBB99D35FFB8FFFFFC8AAE51FC60FB34" box="[843,1013,1148,1173]" collectionCode="BMNH" pageId="8" pageNumber="177">BMNH 1933.8</specimenCode>
|
||
.30.239. (
|
||
<figureCitation id="132429CBFFB8FFFFFB9CAE51FB37FB34" box="[1117,1186,1148,1173]" captionStart="FIGURE 8" captionStartId="7.[151,249,1625,1647]" captionTargetBox="[161,1425,575,1618]" captionTargetId="figure-252@7.[161,1425,575,1618]" captionTargetPageId="7" captionText="FIGURE 8. Otostigmus (Otostigmus) cuneiventris, O. (Parotostigmus) coltellus n. sp., O. (P.) gemmifer and O. (P.) productus, distribution records of specimens from Africa housed in the NHM collection." figureDoi="http://doi.org/10.5281/zenodo.5275605" httpUri="https://zenodo.org/record/5275605/files/figure.png" pageId="8" pageNumber="177">Fig. 8</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30566C5FFB8FFFFFF06AEB2FD1CF854" pageId="8" pageNumber="177" type="description">
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AEB2FD31FB19" blockId="8.[151,1437,932,2037]" box="[199,676,1183,1208]" pageId="8" pageNumber="177">
|
||
<emphasis id="B96BE95CFFB8FFFFFF06AEB2FECCFB19" bold="true" box="[199,345,1183,1208]" inLineHeading="true" pageId="8" pageNumber="177" reason="1">Description.</emphasis>
|
||
Length to
|
||
<quantity id="4CE798ABFFB8FFFFFE13AEB2FDB4FB16" box="[466,545,1183,1208]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="177" unit="mm" value="78.0">78 mm</quantity>
|
||
(
|
||
<typeStatus id="54A48BECFFB8FFFFFDF1AEB2FD0DFB19" box="[560,664,1183,1208]" pageId="8" pageNumber="177" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AEE9FB12FB7C" blockId="8.[151,1437,932,2037]" box="[199,1159,1220,1245]" pageId="8" pageNumber="177">
|
||
Cephalic plate and tergites pale brown to chestnut brown (
|
||
<figureCitation id="132429CBFFB8FFFFFC92AEE9FC0DFB7C" box="[851,920,1220,1245]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Fig. 9</figureCitation>
|
||
); legs orange-brown.
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AECAFBE3FA84" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
|
||
<materialsCitation id="3B773F13FFB8FFFFFF06AECAFBE7FA84" collectionCode="T, ZMB" pageId="8" pageNumber="177" specimenCode="T5, ZMB 927" specimenCount="1">
|
||
Antenna extending back to
|
||
<specimenCode id="DBB99D35FFB8FFFFFE3AAECAFD89FAA1" box="[507,540,1255,1280]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="177">T5</specimenCode>
|
||
, with 17 articles in
|
||
<typeStatus id="54A48BECFFB8FFFFFD39AEC5FCA6FAA1" box="[760,819,1256,1280]" pageId="8" pageNumber="177">types</typeStatus>
|
||
(16 articles in
|
||
<specimenCode id="DBB99D35FFB8FFFFFC1AAEC5FBD9FAA1" box="[987,1100,1255,1280]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="177" type="Museum">ZMB 927</specimenCode>
|
||
), the basal four glabrous (
|
||
<figureCitation id="132429CBFFB8FFFFFAAEAECAFF23FA84" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Fig. 10</figureCitation>
|
||
). Cephalic plate punctate, variably with a pair of longitudinal depressions posteriorly
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AF02FBF5FA14" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
|
||
Coxosternal tooth plates several times wider than long; anterior margin of tooth plate straight, adentate, gently inclined laterally for most of its width, expanding at its lateral corner to from a blunt bulge (
|
||
<figureCitation id="132429CBFFB8FFFFFB7BAF79FAABFACC" box="[1210,1342,1364,1389]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 11, 12</figureCitation>
|
||
). Single hyaline seta near midline of toothplate, inserting at boundary of strongly pigmented margin. Trochanteroprefemoral process a sclerotized blade, lacking distinct teeth (
|
||
<figureCitation id="132429CBFFB8FFFFFC13AFB1FBC7FA14" box="[978,1106,1436,1461]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 11, 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AF92FA82FA79" blockId="8.[151,1437,932,2037]" box="[199,1303,1471,1496]" pageId="8" pageNumber="177">Second maxillary telopodite with a pretarsal accessory spur; dorsal spine on distal part of article 2.</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AFC9FC5FF9E4" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
|
||
<materialsCitation id="3B773F13FFB8FFFFFF06AFC9FC53F9E4" collectionCode="T" location="Sternites" municipality="Margination" pageId="8" pageNumber="177" specimenCode="T3" specimenCount="1" stateProvince="Tete">
|
||
Tergites smooth, lacking keels or rugosity. Paramedian sutures continuous from
|
||
<collectingRegion id="49DBFBACFFB8FFFFFBA4AFC9FB12FA5A" box="[1125,1159,1508,1531]" country="Mozambique" name="Tete" pageId="8" pageNumber="177">TT</collectingRegion>
|
||
3 or 4–20; mostly with incomplete sutures extending about half length of
|
||
<specimenCode id="DBB99D35FFB8FFFFFD0BAC2AFD79F981" box="[714,748,1543,1568]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="177">T3</specimenCode>
|
||
.
|
||
<collectingMunicipality id="6BC4AF34FFB8FFFFFD36AC2AFC14F981" box="[759,897,1543,1568]" pageId="8" pageNumber="177">Margination</collectingMunicipality>
|
||
complete on
|
||
<collectingRegion id="49DBFBACFFB8FFFFFBD9AC2AFBA8F981" box="[1048,1085,1543,1568]" country="Mozambique" name="Tete" pageId="8" pageNumber="177">TT</collectingRegion>
|
||
19–21, incomplete from
|
||
<collectingRegion id="49DBFBACFFB8FFFFFA91AC2AFAE6F981" box="[1360,1395,1543,1568]" country="Mozambique" name="Tete" pageId="8" pageNumber="177">TT</collectingRegion>
|
||
14– 15.
|
||
<location id="8EC06395FFB8FFFFFF00AC01FEB6F9E4" LSID="urn:lsid:plazi:treatment:03B68458FFB8FFFDFF56A8FBFEE8FC66:8EC06395FFB8FFFFFF00AC01FEB6F9E4" box="[193,291,1580,1605]" municipality="Margination" name="Sternites" pageId="8" pageNumber="177" stateProvince="Tete">Sternites</location>
|
||
with continuous paramedian sutures to S19, lacking on S20
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AC7DFCE5F92C" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">Legs 1 and 2 with two tarsal spurs; legs 3 to 16, 17, 18 or 19 with one tarsal spur, lacking on leg 20; all legs lacking tibial spurs apart from leg 1 (right leg only) in ZMB 927.</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06ACBAFC8BF911" blockId="8.[151,1437,932,2037]" box="[199,798,1687,1712]" pageId="8" pageNumber="177">Spiracles elliptical, with strongly humped atrial floor.</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AC91FA1CF959" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">Tergite of ultimate leg-bearing segment with broadly V-shaped posterior margin, lacking median depression. Sternite of ultimate leg-bearing segment narrowing posteriorly, with nearly straight lateral and posterior margins.</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AD29FDBAF8C4" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">
|
||
Coxpleural process short, with two small apical spines in
|
||
<typeStatus id="54A48BECFFB8FFFFFC8EAD29FC1FF8BD" box="[847,906,1796,1820]" pageId="8" pageNumber="177">types</typeStatus>
|
||
(one in ZMB 927) but lacking dorsal or lateral spines. Pore field extends to dorsal and posterior margins of coxopleuron, with only a short, narrow pore-free band at coxopleural process (
|
||
<figureCitation id="132429CBFFB8FFFFFE61AD61FDB7F8C4" box="[416,546,1868,1893]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="8" pageNumber="177">Figs 13, 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFB8FFFFFF06AD42FD1CF854" blockId="8.[151,1437,932,2037]" pageId="8" pageNumber="177">Ultimate leg relatively slender, relative lengths of podomeres in the ratio prefemur 1, femur 1, tibia 0.88, tarsus 1 0.73, tarsus 2 0.39, pretarsus 0.22; prefemur of even width (in dorsal view) along its length, width distally about one-third its length. Prefemur lacking spines apart from one mid-ventral spine near midlength in ZMB 927. Ultimate leg pretarsus with accessory claws.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6065C6FFB9FFFEFF56AD79FB26F844" ID-DOI="http://doi.org/10.5281/zenodo.5275607" ID-Zenodo-Dep="5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="9" pageNumber="178" startId="9.[151,264,1876,1899]" targetBox="[151,1436,193,1856]" targetPageId="9">
|
||
<paragraph id="8BA0354EFFB9FFFEFF56AD79FB26F844" blockId="9.[151,1436,1876,2022]" pageId="9" pageNumber="178">
|
||
<emphasis id="B96BE95CFFB9FFFEFF56AD79FED2F8CB" bold="true" box="[151,327,1876,1899]" pageId="9" pageNumber="178">FIGURES 9–14.</emphasis>
|
||
<taxonomicName id="4C1F4ECDFFB9FFFEFE8FAD78FD5FF8CA" authority="Simaiakis & Edgecombe, 2013" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[334,714,1877,1899]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="coltellus" status="sp. nov." subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFB9FFFEFE8FAD78FE54F8CA" box="[334,449,1877,1899]" italics="true" pageId="9" pageNumber="178">Otostigmus</emphasis>
|
||
(
|
||
<emphasis id="B96BE95CFFB9FFFEFE13AD7BFDF0F8CA" box="[466,613,1878,1899]" italics="true" pageId="9" pageNumber="178">Parotostigmus</emphasis>
|
||
)
|
||
<emphasis id="B96BE95CFFB9FFFEFDB5AD78FD5FF8CA" box="[628,714,1877,1899]" italics="true" pageId="9" pageNumber="178">coltellus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2585427FFB9FFFEFD10AD7BFC9DF8CA" box="[721,776,1878,1899]" pageId="9" pageNumber="178" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30566C5FFBAFFFDFF06AABAFB74FF74" pageId="10" pageNumber="179" type="etymology">
|
||
<paragraph id="8BA0354EFFBAFFFDFF06AABAFBFDFF11" blockId="10.[151,1436,151,968]" box="[199,1128,151,176]" pageId="10" pageNumber="179">
|
||
<emphasis id="B96BE95CFFBAFFFDFF06AABAFEDBFF11" bold="true" box="[199,334,151,176]" inLineHeading="true" pageId="10" pageNumber="179" reason="1">Etymology.</emphasis>
|
||
<emphasis id="B96BE95CFFBAFFFDFE95AABAFE3BFF11" box="[340,430,151,176]" italics="true" pageId="10" pageNumber="179">Coltello</emphasis>
|
||
, Italian for knife, with reference to the blade-like tooth plates.
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFBAFFFDFF06AA91FB74FF74" blockId="10.[151,1436,151,968]" box="[199,1249,188,213]" pageId="10" pageNumber="179">
|
||
<emphasis id="B96BE95CFFBAFFFDFF06AA91FE56FF74" bold="true" box="[199,451,188,213]" inLineHeading="true" pageId="10" pageNumber="179" reason="1">General distribution.</emphasis>
|
||
West-Central Tropical Africa: Gulf of
|
||
<collectingCountry id="F30875DEFFBAFFFDFCB6AA91FC5DFF74" box="[887,968,188,213]" name="Guinea-Bissau" pageId="10" pageNumber="179">Guinea</collectingCountry>
|
||
(
|
||
<collectingCountry id="F30875DEFFBAFFFDFC16AA91FBD9FF74" box="[983,1100,188,213]" name="Sao Tome and Principe" pageId="10" pageNumber="179">São Tomé</collectingCountry>
|
||
, Rolas) (ST).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30566C5FFBAFFFDFF06AAF2FEE8FC66" pageId="10" pageNumber="179" type="discussion">
|
||
<paragraph id="8BA0354EFFBAFFFDFF06AAF2FC74FEE1" blockId="10.[151,1436,151,968]" pageId="10" pageNumber="179">
|
||
<emphasis id="B96BE95CFFBAFFFDFF06AAF2FED9FF59" bold="true" box="[199,332,223,248]" inLineHeading="true" pageId="10" pageNumber="179" reason="1">Discussion.</emphasis>
|
||
The type material is from the island of
|
||
<collectingCountry id="F30875DEFFBAFFFDFCC4AAF2FCECFF59" box="[773,889,223,248]" name="Sao Tome and Principe" pageId="10" pageNumber="179">Säo Tomé</collectingCountry>
|
||
; a conspecific specimen (ZMB 927; Museum für Naturkunde, Berlin) from the island of Rolas is a
|
||
<typeStatus id="54A48BECFFBAFFFDFD0DAB29FCB6FEBD" box="[716,803,260,284]" pageId="10" pageNumber="179" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFC8BAB29FA09FEBC" authority="Karsch, 1888" authorityName="Karsch" authorityYear="1888" box="[842,1436,260,285]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFC8BAB29FC5FFEBA" box="[842,970,260,283]" italics="true" pageId="10" pageNumber="179">Otostigmus</emphasis>
|
||
(
|
||
<emphasis id="B96BE95CFFBAFFFDFC1AAB28FBE8FEBD" box="[987,1149,261,284]" italics="true" pageId="10" pageNumber="179">Parotostigmus</emphasis>
|
||
)
|
||
<emphasis id="B96BE95CFFBAFFFDFB4FAB29FB68FEBC" box="[1166,1277,260,285]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
<bibRefCitation id="EF8E48BFFFBAFFFDFAC4AB29FA09FEBC" author="Karsch, F." box="[1285,1436,260,285]" pageId="10" pageNumber="179" pagination="29 - 31" refId="ref17263" refString="Karsch, F. (1888) Zwei neue Myriopoden von Ecuador. Berliner Entomologische Zeitschrift, 32, 29 - 31." type="journal article" year="1888">Karsch, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
but is excluded from that species with the designation of a
|
||
<typeStatus id="54A48BECFFBAFFFDFCE8AB0AFC05FEE1" box="[809,912,295,320]" pageId="10" pageNumber="179" type="lectotype">lectotype</typeStatus>
|
||
below.
|
||
</paragraph>
|
||
<paragraph id="8BA0354EFFBAFFFDFF06AB61FEE8FC66" blockId="10.[151,1436,151,968]" pageId="10" pageNumber="179">
|
||
Compared to all other African
|
||
<emphasis id="B96BE95CFFBAFFFDFDEAAB61FDAAFEC2" box="[555,575,332,355]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFD98AB60FD6EFEC5" authorityName="Pocock" authorityYear="1895" box="[601,763,333,356]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFD98AB60FD6EFEC5" box="[601,763,333,356]" italics="true" pageId="10" pageNumber="179">Parotostigmus</emphasis>
|
||
</taxonomicName>
|
||
), this species is readily distinguished by its adentate tooth plates. The most similar species,
|
||
<emphasis id="B96BE95CFFBAFFFDFDCAAB5DFD8AFE26" box="[523,543,368,391]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFDF6AB5CFDD0FE29" box="[567,581,369,392]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.).
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFDA5AB42FCB9FE29" authority="Karsch" authorityName="Karsch" authorityYear="1888" box="[612,812,367,392]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="productus">
|
||
<emphasis id="B96BE95CFFBAFFFDFDA5AB42FD46FE29" box="[612,723,367,392]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
Karsch
|
||
</taxonomicName>
|
||
, also occurs on
|
||
<collectingCountry id="F30875DEFFBAFFFDFC1EAB42FBC7FE29" box="[991,1106,367,392]" name="Sao Tome and Principe" pageId="10" pageNumber="179">São Tomé</collectingCountry>
|
||
, and the two taxa share some character states that, in combination, separate them from other African
|
||
<emphasis id="B96BE95CFFBAFFFDFC25ABB9FC6DFE0A" box="[996,1016,404,427]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFBD5ABB8FB23FE0D" authorityName="Pocock" authorityYear="1895" box="[1044,1206,405,428]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFBD5ABB8FB23FE0D" box="[1044,1206,405,428]" italics="true" pageId="10" pageNumber="179">Parotostigmus</emphasis>
|
||
</taxonomicName>
|
||
). These include 17 antennal articles, four of which are glabrous (versus
|
||
<quantity id="4CE798ABFFBAFFFDFD38AB9AFCCDFE71" box="[761,856,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="5.715" metricValueMax="6.35" metricValueMin="5.08" pageId="10" pageNumber="179" unit="in" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 in</quantity>
|
||
other species), complete paramedian sutures from TT3 or 4, complete paramedian sutures on the sternites, and a pair of coxopleural apical spines. The uniquely adentate tooth plates of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFE61A82DFDDAFDB9" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[416,591,511,536]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFE61A82DFE2EFDB6" box="[416,443,512,535]" italics="true" pageId="10" pageNumber="179">O.</emphasis>
|
||
(
|
||
<emphasis id="B96BE95CFFBAFFFDFE0AA82CFE4CFDB9" box="[459,473,513,536]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFE31ABD2FDDAFDB9" box="[496,591,511,536]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
are the same in the four known specimens of the species and this difference from
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFF14A809FE03FD9C" authorityName="Karsch" authorityYear="1888" box="[213,406,548,573]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="176" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFF14A809FF7CFD9A" box="[213,233,548,571]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFEC0A808FE9AFD9D" box="[257,271,549,572]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFEE6A809FE03FD9C" box="[295,406,548,573]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
</taxonomicName>
|
||
cannot be attributed to wear, particularly because the sutures defining the base of the tooth plate are distinct from those of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFE37A865FD23FDC1" authorityName="Karsch" authorityYear="1888" box="[502,694,583,608]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="176" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFE37A865FD9FFDFE" box="[502,522,584,607]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFDE3A864FDA5FDC1" box="[546,560,585,608]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFD86A86AFD23FDC1" box="[583,694,583,608]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
</taxonomicName>
|
||
. In the latter species, the relatively long tooth plates are set off by oblique sutures (for further details see figures in
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFD77A841FCE0FD24" authorityName="Karsch" authorityYear="1888" box="[694,885,620,645]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="176" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFD77A841FD5FFD22" box="[694,714,620,643]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFD20A840FD7AFD25" box="[737,751,621,644]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFCC7A841FCE0FD24" box="[774,885,620,645]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
</taxonomicName>
|
||
), whereas the suture at the base of the tooth plate in
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFF75A8BDFEF1FD09" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[180,356,655,680]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFF75A8BDFF5DFD06" box="[180,200,656,679]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFF21A8BCFF7BFD09" box="[224,238,657,680]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFEC4A8A2FEF1FD09" box="[261,356,655,680]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
is approximately transverse (see
|
||
<figureCitation id="132429CBFFBAFFFDFD1DA8A2FCCEFD09" box="[732,859,655,680]" captionStart="FIGURES 9–14" captionStartId="9.[151,264,1876,1899]" captionTargetBox="[151,1436,193,1856]" captionTargetId="figure-115@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 9–14. Otostigmus (Parotostigmus) coltellus n. sp. Holotype BMNH 1950.3.7.127-136 (São Tomé), except figs 11, 14, paratype female from same collection: Figure 9, posterior part of cephalic plate and TT1–4, dorsal view, scale bar 2 mm; Figure 10, left antenna, dorsal view, scale bar 1 mm; Figure 11, forcipular segment, ventral view, scale bar 1 mm; Figure 12, tooth plates, scale bar 0.5 mm; Figure 13, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm; Figure 14, coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5275607" httpUri="https://zenodo.org/record/5275607/files/figure.png" pageId="10" pageNumber="179">Figs 11, 12</figureCitation>
|
||
). As well, the position of the single hyaline seta on each tooth plate is a reliable marker of the boundary between the sclerotized margin of the tooth plate and its basal plate. The two species are also readily distinguished by the shape of the coxopleural process, being shorter and less slender in
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFECAA8D1FE2EFCB4" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[267,443,764,789]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFECAA8D1FE8AFCB2" box="[267,287,764,787]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFEF6A8D0FED9FCB5" box="[311,332,765,788]" italics="true" pageId="10" pageNumber="179">P.</emphasis>
|
||
)
|
||
<emphasis id="B96BE95CFFBAFFFDFE9DA8D1FE2EFCB4" box="[348,443,764,789]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
. These two species co-occur in the collection from Zampalma that includes the
|
||
<typeStatus id="54A48BECFFBAFFFDFAFBA8D1FA09FCB4" box="[1338,1436,764,789]" pageId="10" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFF74A90DFEF0FC99" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[181,357,799,824]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFF74A90DFF5CFC96" box="[181,201,800,823]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFF20A90CFF7AFC99" box="[225,239,801,824]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFEC7A932FEF0FC99" box="[262,357,799,824]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
. Although a marked size difference separates the two (large) specimens of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFB6EA90DFACAFC99" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[1199,1375,799,824]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFB6EA90DFB56FC96" box="[1199,1219,800,823]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFB1BA90CFB7DFC99" box="[1242,1256,801,824]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFAC1A932FACAFC99" box="[1280,1375,799,824]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
from the nine smaller specimens of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFE37A969FD2FFCFC" authorityName="Karsch" authorityYear="1888" box="[502,698,836,861]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="176" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFE37A969FD9FFCFA" box="[502,522,836,859]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFDE2A968FDA4FCFD" box="[547,561,837,860]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFD8AA969FD2FFCFC" box="[587,698,836,861]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
</taxonomicName>
|
||
, morphological differences cannot be attributed to changes with growth because the smaller
|
||
<typeStatus id="54A48BECFFBAFFFDFE29A945FDDCFC21" box="[488,585,872,896]" pageId="10" pageNumber="179" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFDBBA945FCA3FC21" authorityName="Simaiakis & Edgecombe" authorityYear="2013" box="[634,822,871,896]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="coltellus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFDBBA945FD1BFCDE" box="[634,654,872,895]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFD6DA944FD2FFC21" box="[684,698,873,896]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFD16A94AFCA3FC21" box="[727,822,871,896]" italics="true" pageId="10" pageNumber="179">coltellus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1933.8.30.239) is scarcely larger than the Zalpalma specimens of
|
||
<taxonomicName id="4C1F4ECDFFBAFFFDFE66A9A1FDFEFC04" authorityName="Karsch" authorityYear="1888" box="[423,619,908,933]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="176" phylum="Arthropoda" rank="species" species="productus" subGenus="Parotostigmus">
|
||
<emphasis id="B96BE95CFFBAFFFDFE66A9A1FE2EFC02" box="[423,443,908,931]" italics="true" pageId="10" pageNumber="179">O</emphasis>
|
||
. (
|
||
<emphasis id="B96BE95CFFBAFFFDFE15A9A0FE77FC05" box="[468,482,909,932]" italics="true" pageId="10" pageNumber="179">P</emphasis>
|
||
.)
|
||
<emphasis id="B96BE95CFFBAFFFDFE3DA9A1FDFEFC04" box="[508,619,908,933]" italics="true" pageId="10" pageNumber="179">productus</emphasis>
|
||
</taxonomicName>
|
||
, yet both species exhibit their typical characters of the tooth plates and coxopleural process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |