treatments-xml/data/F0/07/87/F007878A4057FFA799D6FB43C9A6FADC.xml

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<mods:title id="D888D8510ABBCF7EF22090647E36D86B">A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia</mods:title>
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<mods:namePart id="09B070C29FDBBD5AB8031CD7AE4D23D7">Ng, Peter K. L.</mods:namePart>
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<treatment id="F007878A4057FFA799D6FB43C9A6FADC" ID-DOI="http://doi.org/10.5281/zenodo.5964457" ID-GBIF-Taxon="148421672" ID-Zenodo-Dep="5964457" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC" httpUri="http://treatment.plazi.org/id/F007878A4057FFA799D6FB43C9A6FADC" lastPageId="9" lastPageNumber="364" pageId="1" pageNumber="356">
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<paragraph id="7811369C4057FFAF99D6FB43CBDBFAD3" blockId="1.[151,562,1255,1315]" box="[151,452,1255,1282]" pageId="1" pageNumber="356">
<heading id="235981F04057FFAF99D6FB43CBDBFAD3" bold="true" box="[151,452,1255,1282]" fontSize="11" level="1" pageId="1" pageNumber="356" reason="1">
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<taxonomicName id="BFAE4D1F4057FFAF99D6FB43CB64FAD0" ID-CoL="84FXF" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[151,379,1255,1281]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="356" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4057FFAF99D6FB43CB64FAD0" bold="true" box="[151,379,1255,1281]" italics="true" pageId="1" pageNumber="356">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54057FFAF98C0FB4CCBDBFAD3" box="[385,452,1256,1282]" pageId="1" pageNumber="356" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="E0952A194057FFAF99DEFAAECAF1FAF2" box="[159,238,1290,1315]" captionStart="FIGURE 2" captionStartId="4.[151,250,1971,1993]" captionTargetBox="[257,1323,193,1932]" captionTargetId="figure@4.[257,1323,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Karstarma malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax." httpUri="https://zenodo.org/record/1440640/files/figure.png" pageId="1" pageNumber="356">Figs. 2</figureCitation>
, 
<figureCitation id="E0952A194057FFAF99BBFAAFCB20FAF2" box="[250,319,1291,1315]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="1" pageNumber="356">3E–H</figureCitation>
, 
<figureCitation id="E0952A194057FFAF980BFAAFCB73FAF2" box="[330,364,1291,1315]" captionStart="FIGURE 4" captionStartId="6.[151,250,1966,1988]" captionTargetBox="[240,1347,193,1932]" captionTargetId="figure@6.[240,1347,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Right ambulatory legs. A, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633)." httpUri="https://zenodo.org/record/1440644/files/figure.png" pageId="1" pageNumber="356">4B</figureCitation>
, 
<figureCitation id="E0952A194057FFAF9839FAAECBE1FAF3" box="[376,510,1290,1315]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="1" pageNumber="356">5E–H, K–L</figureCitation>
, 
<figureCitation id="E0952A194057FFAF9B48FAAEC835FAF2" box="[521,554,1290,1315]" captionStart="FIGURE 6" captionStartId="8.[151,250,1888,1910]" captionTargetBox="[151,1437,193,1848]" captionTargetId="figure@8.[151,1437,193,1848]" captionTargetPageId="8" captionText="FIGURE 6. Colour in life. A, Karstarma jacobsoni (Ihle, 1912), specimen not collected; B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). Photographs by C. Rahmadi." httpUri="https://zenodo.org/record/1440648/files/figure.png" pageId="1" pageNumber="356">6B</figureCitation>
)
</paragraph>
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<materialsCitation id="C8C63CC14057FFAF99D6FAF6C8C9FA62" ID-GBIF-Occurrence="1913889663" collectingDate="2011-10-05" collectionCode="MZB" collectorName="C. Rahmadi &amp; S. Wiantoro" county="Material" elevation="85" latitude="-8.376917" location="Malang Regency" longLatPrecision="1" longitude="112.43102" municipality="Cru" pageId="1" pageNumber="356" specimenCount="1" stateProvince="East Java Province" typeStatus="holotype">
<emphasis id="4ADAEA8E4057FFAF99D6FAF6CB96FABA" bold="true" box="[151,393,1362,1387]" pageId="1" pageNumber="356">
<collectingCounty id="91704E104057FFAF99D6FAF6CB1CFABA" box="[151,259,1362,1387]" pageId="1" pageNumber="356">Material</collectingCounty>
examined.
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: male (18.3 x 
<quantity id="BF569B794057FFAF9BD9FAF6C8E3FABA" box="[664,764,1362,1387]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="356" unit="mm" value="16.7">16.7 mm</quantity>
) (
<collectionCode id="1EBFAE594057FFAF9A53FAF7C94FFABA" box="[786,848,1363,1387]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<collectingMunicipality id="9875ACE64057FFAF9A16FAF7C99EFABA" box="[855,897,1363,1387]" pageId="1" pageNumber="356">Cru</collectingMunicipality>
4633), Gua Lowo, Dusun Sumber Celeng, Desa Banjarejo, Donomulyo District, 
<location id="7D7160474057FFAF9B4DFAD2C8D0FA5E" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9B4DFAD2C8D0FA5E" box="[524,719,1398,1423]" county="Material" latitude="-8.376917" longLatPrecision="1" longitude="112.43102" municipality="Cru" name="Malang Regency" pageId="1" pageNumber="356" stateProvince="East Java Province">Malang Regency</location>
, 
<collectingRegion id="BA6AF87E4057FFAF9B9FFAD3C9A4FA5E" box="[734,955,1398,1423]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="356">
East 
<collectingRegion id="BA6AF87E4057FFAF9A59FAD3C9A4FA5E" box="[792,955,1398,1423]" inferredBySuffix="true" pageId="1" pageNumber="356">Java Province</collectingRegion>
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, 
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<geoCoordinate id="1D9A505B4057FFAF9D21FAD2CF14FA5E" box="[1120,1291,1398,1423]" direction="east" orientation="longitude" pageId="1" pageNumber="356" precision="1" value="112.43102">112°25’51.7”E</geoCoordinate>
, 
<elevation id="F383D1AF4057FFAF9C5BFAD2CF87FA5E" box="[1306,1432,1398,1423]" metricMagnitude="1" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="356" unit="m" value="85.3">
<quantity id="BF569B794057FFAF9C5BFAD2CF73FA5F" box="[1306,1388,1398,1423]" metricMagnitude="1" metricUnit="m" metricValue="8.53" pageId="1" pageNumber="356" unit="m" value="85.3">85.3 m</quantity>
asl
</elevation>
, coll. 
<collectorName id="D55B534A4057FFAF9993FA3FCB45FA62" box="[210,346,1434,1459]" pageId="1" pageNumber="356">C. Rahmadi</collectorName>
&amp; 
<collectorName id="D55B534A4057FFAF98C5FA3FC80EFA62" box="[388,529,1434,1459]" pageId="1" pageNumber="356">S. Wiantoro</collectorName>
, 
<collectingDate id="1C54E9B44057FFAF9B5EFA3EC8C9FA62" box="[543,726,1434,1459]" pageId="1" pageNumber="356" value="2011-10-05">
<date id="0C10105C4057FFAF9B5EFA3EC8C9FA62" box="[543,726,1434,1459]" pageId="1" pageNumber="356" value="2011-10-05">5 October 2011</date>
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. Paratypes: 1 male (12.5 x 
<quantity id="BF569B794057FFAF9D57FA3FCE9FFA62" box="[1046,1152,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="1" pageNumber="356" unit="mm" value="11.1">11.1 mm</quantity>
), 
<materialsCitation id="C8C63CC14057FFAF9DD7FA3FC96AFA06" ID-GBIF-Occurrence="1913889665" collectingDate="2011-10-05" collectionCode="MZB" collectorName="C. Rahmadi &amp; S. Wiantoro" county="Material" elevation="85" latitude="-8.376917" location="Cru" longLatPrecision="1" longitude="112.43102" municipality="Cru" pageId="1" pageNumber="356" specimenCount="3" specimenCount-female="3" stateProvince="East Java Province" typeStatus="holotype">
<specimenCount id="6EA8FD154057FFAF9DD7FA3FCF1AFA62" box="[1174,1285,1434,1459]" pageId="1" pageNumber="356" type="female">3 females</specimenCount>
(13.4 x 12.0 mm–18.8 x 
<quantity id="BF569B794057FFAF9862FA1ACB91FA06" box="[291,398,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="356" unit="mm" value="16.7">16.7 mm</quantity>
), same data as holotype (
<collectionCode id="1EBFAE594057FFAF9BF5FA1BC8EDFA06" box="[692,754,1471,1495]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<location id="7D7160474057FFAF9BBDFA1BC939FA06" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9BBDFA1BC939FA06" box="[764,806,1471,1495]" name="Cru" pageId="1" pageNumber="356">Cru</location>
4635)
</materialsCitation>
; 
<materialsCitation id="C8C63CC14057FFAF9AC2FA1BC8BAF9CE" ID-GBIF-Occurrence="1913889671" collectionCode="MZB" county="Cru" elevation="105" latitude="-8.37525" location="Desa Kedungsalam" longLatPrecision="1" longitude="112.451416" municipality="Gua Rampal" pageId="1" pageNumber="356" specimenCode="ZRC 2017.1027" specimenCount="1" specimenCount-male="1" stateProvince="East Java Province" typeStatus="holotype">
<specimenCount id="6EA8FD154057FFAF9AC2FA1BC9CEFA06" box="[899,977,1470,1495]" pageId="1" pageNumber="356" type="male">1 male</specimenCount>
(22.8 x 
<quantity id="BF569B794057FFAF9D76FA1ACEBFFA06" box="[1079,1184,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="356" unit="mm" value="20.6">20.6 mm</quantity>
) (
<specimenCode id="28089EE74057FFAF9DF9FA1BCF6BFA06" box="[1208,1396,1471,1495]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="356">ZRC 2017.1027</specimenCode>
, ex 
<collectionCode id="1EBFAE594057FFAF99D6FA47CACBFA2B" box="[151,212,1507,1530]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<collectingCounty id="91704E104057FFAF999FFA47CB17FA2A" box="[222,264,1507,1531]" pageId="1" pageNumber="356">Cru</collectingCounty>
4655), 
<collectingMunicipality id="9875ACE64057FFAF9824FA47CBE7FA2A" box="[357,504,1506,1531]" pageId="1" pageNumber="356">Gua Rampal</collectingMunicipality>
, 
<location id="7D7160474057FFAF9B47FA47C8F9FA2A" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9B47FA47C8F9FA2A" box="[518,742,1506,1531]" county="Cru" latitude="-8.37525" longLatPrecision="1" longitude="112.451416" municipality="Gua Rampal" name="Desa Kedungsalam" pageId="1" pageNumber="356" stateProvince="East Java Province">Desa Kedungsalam</location>
, 
<location id="7D7160474057FFAF9BB5FA46C9C4FA2A" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9BB5FA46C9C4FA2A" box="[756,987,1506,1531]" county="Cru" latitude="-8.37525" longLatPrecision="1" longitude="112.451416" municipality="Gua Rampal" name="Donomulyo District" pageId="1" pageNumber="356" stateProvince="East Java Province">Donomulyo District</location>
, 
<location id="7D7160474057FFAF9AA8FA46CEB2FA2A" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9AA8FA46CEB2FA2A" box="[1001,1197,1506,1531]" county="Cru" latitude="-8.37525" longLatPrecision="1" longitude="112.451416" municipality="Gua Rampal" name="Malang Regency" pageId="1" pageNumber="356" stateProvince="East Java Province">Malang Regency</location>
, 
<collectingRegion id="BA6AF87E4057FFAF9DFAFA47CF87FA2A" box="[1211,1432,1506,1531]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="356">
East 
<collectingRegion id="BA6AF87E4057FFAF9DB4FA47CF87FA2A" box="[1269,1432,1506,1531]" inferredBySuffix="true" pageId="1" pageNumber="356">Java Province</collectingRegion>
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, 
<geoCoordinate id="1D9A505B4057FFAF99D6F9A3CB3DF9CE" box="[151,290,1543,1567]" direction="south" orientation="latitude" pageId="1" pageNumber="356" precision="1" value="-8.37525">8°22’30.9”S</geoCoordinate>
<geoCoordinate id="1D9A505B4057FFAF986AF9A2CBCAF9CE" box="[299,469,1542,1567]" direction="east" orientation="longitude" pageId="1" pageNumber="356" precision="1" value="112.451416">112°27’05.1”E</geoCoordinate>
, 
<elevation id="F383D1AF4057FFAF98A2F9A3C875F9CE" box="[483,618,1542,1567]" metricMagnitude="2" metricUnit="m" metricValue="1.047" pageId="1" pageNumber="356" unit="m" value="104.7">
<quantity id="BF569B794057FFAF98A2F9A3C85FF9CF" box="[483,576,1543,1567]" metricMagnitude="2" metricUnit="m" metricValue="1.047" pageId="1" pageNumber="356" unit="m" value="104.7">104.7 m</quantity>
asl
</elevation>
, coll
</materialsCitation>
. EGKN, 
<date id="0C10105C4057FFAF9A4FF9A3C9BAF9CE" box="[782,933,1542,1567]" pageId="1" pageNumber="356" value="2013-07-19">19 July 2013</date>
; 1 male (14.5 x 
<quantity id="BF569B794057FFAF9D23F9A2CED4F9CE" box="[1122,1227,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="356" unit="mm" value="12.5">12.5 mm</quantity>
), 
<materialsCitation id="C8C63CC14057FFAF9DA1F9A3CBCBF992" ID-GBIF-Occurrence="1913889666" collectionCode="MZB" location="Cru" pageId="1" pageNumber="356" specimenCount="1" specimenCount-female="1" stateProvince="East Java Province" typeStatus="holotype">
<specimenCount id="6EA8FD154057FFAF9DA1F9A3CF5DF9CE" box="[1248,1346,1542,1567]" pageId="1" pageNumber="356" type="female">1 female</specimenCount>
(22.0 x 
<quantity id="BF569B794057FFAF99D6F98FCAE1F992" box="[151,254,1579,1603]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="1" pageNumber="356" unit="mm" value="20.1">20.1 mm</quantity>
) (
<collectionCode id="1EBFAE594057FFAF9857F98FCB4BF992" box="[278,340,1579,1603]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<location id="7D7160474057FFAF981CF98FCB98F992" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF981CF98FCB98F992" box="[349,391,1579,1603]" name="Cru" pageId="1" pageNumber="356" stateProvince="East Java Province">Cru</location>
4634)
</materialsCitation>
, 
<materialsCitation id="C8C63CC14057FFAF98A0F98FC8C0F95A" ID-GBIF-Occurrence="1913889668" collectingDate="2012-05-19" collectionCode="MZB" collectorName="Kusmayasari &amp; E. Tazkiyah" county="Cru" elevation="214" latitude="-8.347416" location="Malang Regency" longLatPrecision="1" longitude="112.448" municipality="Donomulyo District" pageId="1" pageNumber="356" specimenCode="ZRC 2017.1028" specimenCount="1" specimenCount-female="1" stateProvince="East Java Province" typeStatus="holotype">
<specimenCount id="6EA8FD154057FFAF98A0F98FC85BF992" box="[481,580,1578,1603]" pageId="1" pageNumber="356" type="female">1 female</specimenCount>
(21.6 x 
<quantity id="BF569B794057FFAF9BE7F98FC912F992" box="[678,781,1579,1603]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="1" pageNumber="356" unit="mm" value="19.9">19.9 mm</quantity>
) (
<specimenCode id="28089EE74057FFAF9A64F98FC9FFF992" box="[805,992,1579,1603]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="356">ZRC 2017.1028</specimenCode>
, ex 
<collectionCode id="1EBFAE594057FFAF9D4EF98FCE53F993" box="[1039,1100,1579,1602]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<collectingCounty id="91704E104057FFAF9D14F98FCE60F992" box="[1109,1151,1579,1603]" pageId="1" pageNumber="356">Cru</collectingCounty>
4634), Gua Kerek, Desa Kedungsalam, 
<collectingMunicipality id="9875ACE64057FFAF9803F9EAC837F9B6" box="[322,552,1614,1639]" pageId="1" pageNumber="356">Donomulyo District</collectingMunicipality>
, 
<location id="7D7160474057FFAF9B77F9EAC8E7F9B6" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9B77F9EAC8E7F9B6" box="[566,760,1614,1639]" county="Cru" latitude="-8.347416" longLatPrecision="1" longitude="112.448" municipality="Donomulyo District" name="Malang Regency" pageId="1" pageNumber="356" stateProvince="East Java Province">Malang Regency</location>
, 
<collectingRegion id="BA6AF87E4057FFAF9A47F9EBC9FDF9B6" box="[774,994,1614,1639]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="356">
East 
<collectingRegion id="BA6AF87E4057FFAF9A01F9EBC9FDF9B6" box="[832,994,1614,1639]" inferredBySuffix="true" pageId="1" pageNumber="356">Java Province</collectingRegion>
</collectingRegion>
, 
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<geoCoordinate id="1D9A505B4057FFAF9DC4F9EACF2FF9B6" box="[1157,1328,1614,1639]" direction="east" orientation="longitude" pageId="1" pageNumber="356" precision="1" value="112.448">112°26’52.8”E</geoCoordinate>
, 
<elevation id="F383D1AF4057FFAF9C7FF9EBCAA6F95A" metricMagnitude="2" metricUnit="m" metricValue="2.1369999999999996" pageId="1" pageNumber="356" unit="m" value="213.7">
<quantity id="BF569B794057FFAF9C7FF9EBCF83F9B7" box="[1342,1436,1615,1639]" metricMagnitude="2" metricUnit="m" metricValue="2.1369999999999996" pageId="1" pageNumber="356" unit="m" value="213.7">213.7 m</quantity>
asl
</elevation>
, coll. 
<collectorName id="D55B534A4057FFAF99BDF9D6CB91F95A" box="[252,398,1650,1675]" pageId="1" pageNumber="356">Kusmayasari</collectorName>
&amp; 
<collectorName id="D55B534A4057FFAF98F3F9D7C825F95A" box="[434,570,1650,1675]" pageId="1" pageNumber="356">E. Tazkiyah</collectorName>
, 
<collectingDate id="1C54E9B44057FFAF9B04F9D7C8C0F95A" box="[581,735,1651,1675]" pageId="1" pageNumber="356" value="2012-05-19">
<date id="0C10105C4057FFAF9B04F9D7C8C0F95A" box="[581,735,1651,1675]" pageId="1" pageNumber="356" value="2012-05-19">19 May 2012</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="7811369C4057FFAF9986F932CF2EF97E" blockId="1.[151,1437,1362,2035]" box="[199,1329,1686,1711]" pageId="1" pageNumber="356">
<materialsCitation id="C8C63CC14057FFAF9986F932CF2EF97E" ID-GBIF-Occurrence="1913889672" box="[199,1329,1686,1711]" country="Indonesia" location="Caves in Donomulyo District" municipality="southern Malang Regency" pageId="1" pageNumber="356" specimenCount="1" stateProvince="East Java Province" typeStatus="holotype">
<emphasis id="4ADAEA8E4057FFAF9986F932CB7CF97E" bold="true" box="[199,355,1686,1711]" pageId="1" pageNumber="356">
<typeStatus id="A715883E4057FFAF9986F932CB1FF97E" box="[199,256,1686,1711]" pageId="1" pageNumber="356">Type</typeStatus>
locality.
</emphasis>
<location id="7D7160474057FFAF9828F933C8AAF97E" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9828F933C8AAF97E" box="[361,693,1686,1711]" country="Indonesia" municipality="southern Malang Regency" name="Caves in Donomulyo District" pageId="1" pageNumber="356" stateProvince="East Java Province">Caves in Donomulyo District</location>
, 
<collectingMunicipality id="9875ACE64057FFAF9B81F932C9C8F97E" box="[704,983,1686,1711]" pageId="1" pageNumber="356">southern Malang Regency</collectingMunicipality>
, 
<collectingRegion id="BA6AF87E4057FFAF9AA1F933CEAEF97E" box="[992,1201,1686,1711]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="356">
East 
<collectingRegion id="BA6AF87E4057FFAF9D55F933CEAEF97E" box="[1044,1201,1686,1711]" inferredBySuffix="true" pageId="1" pageNumber="356">Java Province</collectingRegion>
</collectingRegion>
, 
<collectingCountry id="00B9760C4057FFAF9DFDF932CF32F97E" box="[1212,1325,1686,1711]" name="Indonesia" pageId="1" pageNumber="356">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="7811369C4057FFAC9986F91EC910FF29" blockId="1.[151,1437,1362,2035]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="357" pageId="1" pageNumber="356">
<materialsCitation id="C8C63CC14057FFAF9986F91ECEFDF926" ID-GBIF-Occurrence="1913889670" collectionCode="ZRC" country="Indonesia" county="Djumblang Cave" location="Gunung Sewu" municipality="Bedojo District" pageId="1" pageNumber="356" specimenCount="2" specimenCount-male="2" stateProvince="central Java" typeStatus="syntype">
<emphasis id="4ADAEA8E4057FFAF9986F91ECBC5F902" bold="true" box="[199,474,1722,1747]" pageId="1" pageNumber="356">Comparative material.</emphasis>
<taxonomicName id="BFAE4D1F4057FFAF98A2F91FC978F902" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[483,871,1722,1747]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="356" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4057FFAF98A2F91FC8CCF902" box="[483,723,1723,1747]" italics="true" pageId="1" pageNumber="356">Karstarma jacobsoni</emphasis>
(
<bibRefCitation id="1C3F4B6D4057FFAF9BA5F91EC941F902" box="[740,862,1722,1747]" pageId="1" pageNumber="356" refId="ref4291" refString="Ihle, J. E. W. (1912) Ueber eine kleine Brachyuren-Sammlung aus unterirdischen Flussen von Java. Notes from the Leyden Museum, 34 (3 &amp; 4), 177 - 182, pl. 9." type="journal article">Ihle, 1912</bibRefCitation>
)
</taxonomicName>
: 
<typeStatus id="A715883E4057FFAF9A35F91FC9C0F902" box="[884,991,1723,1747]" pageId="1" pageNumber="356" type="syntype">Syntypes</typeStatus>
: 
<specimenCount id="6EA8FD154057FFAF9AAAF91FCE5CF902" box="[1003,1091,1722,1747]" pageId="1" pageNumber="356" type="male">2 males</specimenCount>
(18.9 x 
<quantity id="BF569B794057FFAF9DE7F91FCF12F903" box="[1190,1293,1723,1747]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="356" unit="mm" value="17.1">17.1 mm</quantity>
, 18.5 x 17.0 mm) (
<collectionCode id="1EBFAE594057FFAF999BF97BCB0DF926" box="[218,274,1759,1783]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="356">ZRC</collectionCode>
1970.3.1.1), 
<collectingCounty id="91704E104057FFAF98E8F97AC875F926" box="[425,618,1758,1783]" pageId="1" pageNumber="356">Djumblang Cave</collectingCounty>
, 
<collectingMunicipality id="9875ACE64057FFAF9B37F97AC93AF926" box="[630,805,1758,1783]" pageId="1" pageNumber="356">Bedojo District</collectingMunicipality>
, 
<location id="7D7160474057FFAF9A70F97BC9CCF926" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9A70F97BC9CCF926" box="[817,979,1759,1783]" country="Indonesia" county="Djumblang Cave" municipality="Bedojo District" name="Gunung Sewu" pageId="1" pageNumber="356" stateProvince="central Java">Gunung Sewu</location>
, 
<collectingRegion id="BA6AF87E4057FFAF9A9EF97ACE79F927" box="[991,1126,1758,1783]" country="Indonesia" name="Jawa Tengah" pageId="1" pageNumber="356">central Java</collectingRegion>
, 
<collectingCountry id="00B9760C4057FFAF9D33F97ACEFDF926" box="[1138,1250,1758,1783]" name="Indonesia" pageId="1" pageNumber="356">Indonesia</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="C8C63CC14057FFAF9DAFF97BC9E4F8B2" ID-GBIF-Occurrence="1913889664" collectingDate="2006-08-07" collectionCode="MZB" collectorName="C. Rahmadi" country="Indonesia" elevation="134" latitude="-8.028333" location="Cru" longLatPrecision="1" longitude="110.60408" pageId="1" pageNumber="356" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Yogyakarta" typeStatus="syntype">
<specimenCount id="6EA8FD154057FFAF9DAFF97BCF5AF926" box="[1262,1349,1758,1783]" pageId="1" pageNumber="356" type="male">4 males</specimenCount>
(14.5 x 
<quantity id="BF569B794057FFAF99D6F8A7CAE3F8CA" box="[151,252,1795,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="356" unit="mm" value="12.8">12.8 mm</quantity>
–19.4 x 18.0 mm), 
<specimenCount id="6EA8FD154057FFAF9892F8A7C820F8CA" box="[467,575,1794,1819]" pageId="1" pageNumber="356" type="female">2 females</specimenCount>
(16.1 x 14.0 mm, 19.3 x 
<quantity id="BF569B794057FFAF9A1FF8A7C9DBF8CA" box="[862,964,1795,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="1" pageNumber="356" unit="mm" value="17.4">17.4 mm</quantity>
) (
<collectionCode id="1EBFAE594057FFAF9A9AF8A7CE06F8CA" box="[987,1049,1795,1819]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<location id="7D7160474057FFAF9D60F8A7CE54F8CA" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9D60F8A7CE54F8CA" box="[1057,1099,1795,1819]" country="Indonesia" latitude="-8.028333" longLatPrecision="1" longitude="110.60408" name="Cru" pageId="1" pageNumber="356" stateProvince="Yogyakarta">Cru</location>
1590), Gua Ngingrong Mulo, Dusun Serpeng, Desa Pacarejo, Semanu District, Gunung Kidul Regency, Special Region of 
<collectingRegion id="BA6AF87E4057FFAF9DEBF882CF35F8EE" box="[1194,1322,1830,1855]" country="Indonesia" name="Yogyakarta" pageId="1" pageNumber="356">Yogyakarta</collectingRegion>
Province, 
<geoCoordinate id="1D9A505B4057FFAF99D6F8EFCB2EF8B2" box="[151,305,1867,1891]" direction="south" orientation="latitude" pageId="1" pageNumber="356" precision="1" value="-8.028333">08°01’42.0”S</geoCoordinate>
<geoCoordinate id="1D9A505B4057FFAF9879F8EECBFDF8B2" box="[312,482,1866,1891]" direction="east" orientation="longitude" pageId="1" pageNumber="356" precision="1" value="110.60408">110°36’14.7”E</geoCoordinate>
, 
<elevation id="F383D1AF4057FFAF98AEF8EFC869F8B2" box="[495,630,1866,1891]" metricMagnitude="2" metricUnit="m" metricValue="1.343" pageId="1" pageNumber="356" unit="m" value="134.3">
<quantity id="BF569B794057FFAF98AEF8EFC853F8B3" box="[495,588,1867,1891]" metricMagnitude="2" metricUnit="m" metricValue="1.343" pageId="1" pageNumber="356" unit="m" value="134.3">134.3 m</quantity>
asl
</elevation>
, coll. 
<collectorName id="D55B534A4057FFAF9BFBF8EFC95BF8B2" box="[698,836,1866,1891]" pageId="1" pageNumber="356">C. Rahmadi</collectorName>
, 
<collectingDate id="1C54E9B44057FFAF9A11F8EFC9E4F8B2" box="[848,1019,1866,1891]" pageId="1" pageNumber="356" value="2006-08-07">
<date id="0C10105C4057FFAF9A11F8EFC9E4F8B2" box="[848,1019,1866,1891]" pageId="1" pageNumber="356" value="2006-08-07">7 August 2006</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="C8C63CC14057FFAF9D47F8EFC99DF81E" ID-GBIF-Occurrence="1913889669" collectingDate="2006-08-13" collectionCode="MZB" collectorName="A. F. Hidayat" country="Indonesia" elevation="387" latitude="-8.004972" location="Cru" longLatPrecision="1" longitude="110.758224" pageId="1" pageNumber="356" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yogyakarta" typeStatus="syntype">
<specimenCount id="6EA8FD154057FFAF9D47F8EFCE4CF8B2" box="[1030,1107,1866,1891]" pageId="1" pageNumber="356" type="male">1 male</specimenCount>
(22.8 x 
<quantity id="BF569B794057FFAF9DF3F8EFCF04F8B2" box="[1202,1307,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="1" pageNumber="356" unit="mm" value="21.2">21.2 mm</quantity>
), 
<specimenCount id="6EA8FD154057FFAF9C6EF8EFCF83F8B2" box="[1327,1436,1866,1891]" pageId="1" pageNumber="356" type="female">2 females</specimenCount>
(11.7 x 
<quantity id="BF569B794057FFAF99B8F8CBCB79F857" box="[249,358,1903,1927]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="1" pageNumber="356" unit="mm" value="10.4">10.4 mm</quantity>
, 15.6 x 
<quantity id="BF569B794057FFAF9893F8CBC821F856" box="[466,574,1903,1927]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="356" unit="mm" value="13.1">13.1 mm</quantity>
) (
<collectionCode id="1EBFAE594057FFAF9B1AF8CBC886F856" box="[603,665,1903,1927]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<location id="7D7160474057FFAF9BE6F8CBC8CEF856" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9BE6F8CBC8CEF856" box="[679,721,1903,1927]" country="Indonesia" latitude="-8.004972" longLatPrecision="1" longitude="110.758224" name="Cru" pageId="1" pageNumber="356" stateProvince="Yogyakarta">Cru</location>
1597), Gua Jomblang Bedoyo, Dusun Jomblang Lor, Desa Karangasam, Ponjong District, Gunung Kidul Regency, Special Region of 
<collectingRegion id="BA6AF87E4057FFAF9D43F836CE9DF87A" box="[1026,1154,1938,1963]" country="Indonesia" name="Yogyakarta" pageId="1" pageNumber="356">Yogyakarta</collectingRegion>
Province, 
<geoCoordinate id="1D9A505B4057FFAF9C43F837CF83F87A" box="[1282,1436,1939,1963]" direction="south" orientation="latitude" pageId="1" pageNumber="356" precision="1" value="-8.004972">08°00’17.9”S</geoCoordinate>
<geoCoordinate id="1D9A505B4057FFAF99D6F812CB5DF81E" box="[151,322,1974,1999]" direction="east" orientation="longitude" pageId="1" pageNumber="356" precision="1" value="110.758224">110°45’29.6”E</geoCoordinate>
, 
<elevation id="F383D1AF4057FFAF980EF812CBC8F81E" box="[335,471,1974,1999]" metricMagnitude="2" metricUnit="m" metricValue="3.867" pageId="1" pageNumber="356" unit="m" value="386.7">
<quantity id="BF569B794057FFAF980EF812CBB2F81F" box="[335,429,1974,1999]" metricMagnitude="2" metricUnit="m" metricValue="3.867" pageId="1" pageNumber="356" unit="m" value="386.7">386.7 m</quantity>
asl
</elevation>
, coll. 
<collectorName id="D55B534A4057FFAF9B5FF813C8A3F81E" box="[542,700,1974,1999]" pageId="1" pageNumber="356">A. F. Hidayat</collectorName>
, 
<collectingDate id="1C54E9B44057FFAF9B89F813C99DF81E" box="[712,898,1974,1999]" pageId="1" pageNumber="356" value="2006-08-13">
<date id="0C10105C4057FFAF9B89F813C99DF81E" box="[712,898,1974,1999]" pageId="1" pageNumber="356" value="2006-08-13">13 August 2006</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="C8C63CC14057FFAC9AD1F813C913FF29" ID-GBIF-Occurrence="1913889667" collectingDate="2007-07-20" collectionCode="MZB" collectorName="D. Wowor &amp; Wahyudin" country="Indonesia" elevation="474" lastPageId="2" lastPageNumber="357" latitude="-7.965389" location="Cru" longLatPrecision="1" longitude="110.75467" pageId="1" pageNumber="356" specimenCode="ZRC 2009.0807" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Yogyakarta" typeStatus="syntype">
<specimenCount id="6EA8FD154057FFAF9AD1F813C9F7F81E" box="[912,1000,1974,1999]" pageId="1" pageNumber="356" type="male">3 males</specimenCount>
(21.6 x 
<quantity id="BF569B794057FFAF9D0AF813CEAEF81E" box="[1099,1201,1975,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="356" unit="mm" value="19.7">19.7 mm</quantity>
–24.3 x 
<quantity id="BF569B794057FFAF9C4EF813CF66F81E" box="[1295,1401,1975,1999]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="1" pageNumber="356" unit="mm" value="22.3">22.3 mm</quantity>
), 
<specimenCount id="6EA8FD154057FFAF9CCFF813CAFCF822" pageId="1" pageNumber="356" type="female">1 female</specimenCount>
(22.4 x 
<quantity id="BF569B794057FFAF9805F87FCBB1F822" box="[324,430,2011,2035]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="1" pageNumber="356" unit="mm" value="19.9">19.9 mm</quantity>
), 1 ovigerous female (24.0 x 
<quantity id="BF569B794057FFAF9BBDF87EC97CF823" box="[764,867,2010,2035]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="1" pageNumber="356" unit="mm" value="21.6">21.6 mm</quantity>
, eggs 
<quantity id="BF569B794057FFAF9AEDF87FCE1CF823" box="[940,1027,2011,2035]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="356" unit="mm" value="1.8">1.8 mm</quantity>
diameter) (
<collectionCode id="1EBFAE594057FFAF9DC7F87FCEDBF822" box="[1158,1220,2011,2035]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="356">MZB</collectionCode>
<location id="7D7160474057FFAF9D8CF87FCEE8F822" LSID="urn:lsid:plazi:treatment:F007878A4057FFA799D6FB43C9A6FADC:7D7160474057FFAF9D8CF87FCEE8F822" box="[1229,1271,2011,2035]" country="Indonesia" latitude="-7.965389" longLatPrecision="1" longitude="110.75467" name="Cru" pageId="1" pageNumber="356" stateProvince="Yogyakarta">Cru</location>
4656), 
<specimenCount id="6EA8FD154057FFAF9C0EF87FCF83F822" box="[1359,1436,2010,2035]" pageId="1" pageNumber="356" type="male">1 male</specimenCount>
(20.7 x 
<quantity id="BF569B794054FFAC99B7FF3CCB7DFF61" box="[246,354,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="2" pageNumber="357" unit="mm" value="19.4">19.4 mm</quantity>
), 
<specimenCount id="6EA8FD154054FFAC983BFF3CCBC0FF61" box="[378,479,151,176]" pageId="2" pageNumber="357" type="female">1 female</specimenCount>
(21.1 x 
<quantity id="BF569B794054FFAC9B0BFF3CC8ACFF61" box="[586,691,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="2" pageNumber="357" unit="mm" value="18.8">18.8 mm</quantity>
) (
<specimenCode id="28089EE74054FFAC9B8FFF3CC992FF61" box="[718,909,152,176]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="357">ZRC 2009.0807</specimenCode>
), Gua Gilap, Dusun Klumpit, Desa Kenteng, Ponjong District, Gunung Kidul Regency, Special Region of 
<collectingRegion id="BA6AF87E4054FFAC9A11FF18C9CFFF04" box="[848,976,188,213]" country="Indonesia" name="Yogyakarta" pageId="2" pageNumber="357">Yogyakarta</collectingRegion>
Province, 
<geoCoordinate id="1D9A505B4054FFAC9D0AFF18CEFAFF04" box="[1099,1253,188,213]" direction="south" orientation="latitude" pageId="2" pageNumber="357" precision="1" value="-7.965389">07°57’55.4”S</geoCoordinate>
<geoCoordinate id="1D9A505B4054FFAC9DACFF18CF87FF04" box="[1261,1432,188,213]" direction="east" orientation="longitude" pageId="2" pageNumber="357" precision="1" value="110.75467">110°45’16.8”E</geoCoordinate>
, 
<elevation id="F383D1AF4054FFAC99D6FF44CB03FF29" box="[151,284,223,248]" metricMagnitude="2" metricUnit="m" metricValue="4.738" pageId="2" pageNumber="357" unit="m" value="473.8">
<quantity id="BF569B794054FFAC99D6FF44CAECFF26" box="[151,243,224,248]" metricMagnitude="2" metricUnit="m" metricValue="4.738" pageId="2" pageNumber="357" unit="m" value="473.8">473.8 m</quantity>
asl
</elevation>
, coll. 
<collectorName id="D55B534A4054FFAC981EFF44CBCCFF26" box="[351,467,224,247]" pageId="2" pageNumber="357">D. Wowor</collectorName>
&amp; 
<collectorName id="D55B534A4054FFAC98B6FF7BC873FF29" box="[503,620,223,248]" pageId="2" pageNumber="357">Wahyudin</collectorName>
, 
<collectingDate id="1C54E9B44054FFAC9B36FF44C913FF29" box="[631,780,223,248]" pageId="2" pageNumber="357" value="2007-07-20">
<date id="0C10105C4054FFAC9B36FF44C913FF29" box="[631,780,223,248]" pageId="2" pageNumber="357" value="2007-07-20">20 July 2007</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="30B465174054FFAC9986FEA0C8CEFB71" pageId="2" pageNumber="357" type="diagnosis">
<paragraph id="7811369C4054FFAC9986FEA0C8CEFB71" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="357">
<emphasis id="4ADAEA8E4054FFAC9986FEA0CB5DFECC" bold="true" box="[199,322,260,285]" pageId="2" pageNumber="357">Diagnosis.</emphasis>
Carapace trapezoidal in shape, widest between bases of second and third ambulatory legs (third and fourth pereopods); dorsal surface distinctly rugose especially on branchial region, without setae; external orbital tooth directly anteriorly, separated from first anterolateral tooth by small but distinct V-shaped cleft, anterolateral margin straight; frontal margin bilobed, with distinct median notch; epigastric crista well defined, 4- lobed; postorbital crista present as 2 small bulges; gastric region well defined, groove present along median epi- and protogastric regions, diverging on meso- and metagastric regions and surrounding gastro-cardiac groove, cervical groove indistinct; posterolateral region with long striae; lateral margin with numerous short stiff setae. Eyes developed with well-pigmented cornea, cornea narrower than peduncle in dorsal view, peduncle smooth and swollen. Thoracic sternites 2 and 3 fused, suture indistinct; suture between 3 and 4 discernible. Chelipeds equal, relatively more robust in males, all margins granulated, dactylus with 12 tubercles at about 0.7 of proximo-lateral margin, cutting edges with row of numerous triangular teeth. Ambulatory legs relatively long, slender, third pair (fourth pereopod) longest, fourth pair shortest, dorsal margins of meri serrated, with subdistal spine. First ambulatory leg (second pereopod) with whole lateral dactylus surface (except corneous tip) and half distolateral surface of propodus bearing dense mat of short black setae covering most of surface, merus length 4 times as long as wide; second ambulatory leg (third pereopod) with dense mat of short black setae on about 0.8 of proximolateral surface of dactylus and 0.25 of distolateral surface of propodus, merus length 3.2 times as long as wide; third ambulatory leg with scattered clumps of short black setae on about 0.8 proximolateral surface of dactylus and 0.2 distolateral surface of propodus, merus length 3.53 times as long as wide; fourth ambulatory leg without mat of short black setae on dactylus and propodus, merus length 3.4 times as long as wide. All pleonal somites and telson free; male pleon triangular, wide; somites 1 and 2 short; somite 3 with convex lateral margins; somites 4 and 5 trapezoidal in shape with slightly convex lateral margins; somite 6 with convex margins; apex of telson rounded, 1.3 times as long as wide. Female pleon rounded, very wide; somite 3 trapezoidal in shape with straight lateral margins; somites 4–6 each with convex lateral margins; telson distinctly sunken into distal margin of somite 6. G1 relatively long, slender, with submedian hump along inner margin, distal part bent outwards approximately 45° to longitudinal axis, outer margin and subdistal areas covered with dense long setae; distal chitinous tip elongate, relatively slender, spade-like with rounded margin.
</paragraph>
</subSubSection>
<subSubSection id="30B465174054FFAC9986FB08C96AFADC" pageId="2" pageNumber="357" type="etymology">
<paragraph id="7811369C4054FFAC9986FB08CBE9FB39" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="357">
<emphasis id="4ADAEA8E4054FFAC9986FB08CB52FB14" bold="true" box="[199,333,1196,1221]" pageId="2" pageNumber="357">Etymology.</emphasis>
The species name is derived from the 
<typeStatus id="A715883E4054FFAC9A43FB09C92DFB14" box="[770,818,1197,1221]" pageId="2" pageNumber="357">type</typeStatus>
locality, Malang, in eastern 
<collectingRegion id="BA6AF87E4054FFAC9D30FB09CEBAFB15" box="[1137,1189,1197,1220]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="357">Java</collectingRegion>
, 
<collectingCountry id="00B9760C4054FFAC9DF0FB08CF3DFB14" box="[1201,1314,1196,1221]" name="Indonesia" pageId="2" pageNumber="357">Indonesia</collectingCountry>
. The name is used as a noun in apposition.
</paragraph>
<paragraph id="7811369C4054FFAC9986FB50C96AFADC" blockId="2.[151,1437,151,2013]" box="[199,885,1268,1293]" pageId="2" pageNumber="357">
<emphasis id="4ADAEA8E4054FFAC9986FB50CB02FADC" bold="true" box="[199,285,1268,1293]" pageId="2" pageNumber="357">Colour.</emphasis>
Carapace light brown with pincers whitish (
<figureCitation id="E0952A194054FFAC9A4EFB50C977FADC" box="[783,872,1268,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,1888,1910]" captionTargetBox="[151,1437,193,1848]" captionTargetId="figure@8.[151,1437,193,1848]" captionTargetPageId="8" captionText="FIGURE 6. Colour in life. A, Karstarma jacobsoni (Ihle, 1912), specimen not collected; B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). Photographs by C. Rahmadi." httpUri="https://zenodo.org/record/1440648/files/figure.png" pageId="2" pageNumber="357">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="30B465174054FFA79986FAB3C98FFDC9" lastPageId="9" lastPageNumber="364" pageId="2" pageNumber="357" type="discussion">
<paragraph id="7811369C4054FFAC9986FAB3CF73F89C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="357">
<emphasis id="4ADAEA8E4054FFAC9986FAB3CB24FAE1" bold="true" box="[199,315,1303,1328]" pageId="2" pageNumber="357">Remarks.</emphasis>
<taxonomicName id="BFAE4D1F4054FFAC9808FABDC83BFAE1" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[329,548,1304,1328]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4054FFAC9808FABDC83BFAE1" box="[329,548,1304,1328]" italics="true" pageId="2" pageNumber="357">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54054FFAC9B73FAB3C868FAE1" box="[562,631,1303,1328]" pageId="2" pageNumber="357" rank="species">
<emphasis id="4ADAEA8E4054FFAC9B73FAB3C868FAE1" bold="true" box="[562,631,1303,1328]" pageId="2" pageNumber="357">n. sp.</emphasis>
</taxonomicNameLabel>
closely resembles 
<taxonomicName id="BFAE4D1F4054FFAC9A21FABDCE92FAE1" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[864,1165,1303,1328]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9A21FABDC9EAFAE1" box="[864,1013,1304,1328]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
(
<bibRefCitation id="1C3F4B6D4054FFAC9D4BFAB3CE9BFAE1" box="[1034,1156,1303,1328]" pageId="2" pageNumber="357" refId="ref4291" refString="Ihle, J. E. W. (1912) Ueber eine kleine Brachyuren-Sammlung aus unterirdischen Flussen von Java. Notes from the Leyden Museum, 34 (3 &amp; 4), 177 - 182, pl. 9." type="journal article">Ihle, 1912</bibRefCitation>
)
</taxonomicName>
in having a distinctly trapezoidal carapace, a sharp external orbital tooth that is separated from first anterolateral tooth by a distinct Vshaped cleft, a straight anterolateral margin, reduced corneas, proportions of the third ambulatory legs (
<figureCitation id="E0952A194054FFAC9C60FAFBCF71FAA9" box="[1313,1390,1375,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1978,2000]" captionTargetBox="[257,1318,195,1934]" captionTargetId="figure@3.[257,1318,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807). A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax." httpUri="https://zenodo.org/record/1440638/files/figure.png" pageId="2" pageNumber="357">Figs. 1</figureCitation>
, 
<figureCitation id="E0952A194054FFAC9C3CFAC4CF92FAA9" box="[1405,1421,1376,1400]" captionStart="FIGURE 2" captionStartId="4.[151,250,1971,1993]" captionTargetBox="[257,1323,193,1932]" captionTargetId="figure@4.[257,1323,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Karstarma malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax." httpUri="https://zenodo.org/record/1440640/files/figure.png" pageId="2" pageNumber="357">2</figureCitation>
); and similarly shaped and positioned vulvae (
<figureCitation id="E0952A194054FFAC9BCDFA20C918FA4D" box="[652,775,1412,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3C, G</figureCitation>
). 
<taxonomicName id="BFAE4D1F4054FFAC9A5AFA21C9EFFA4D" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[795,1008,1412,1436]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4054FFAC9A5AFA21C9EFFA4D" box="[795,1008,1412,1436]" italics="true" pageId="2" pageNumber="357">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54054FFAC9AB6FA20CE28FA4C" box="[1015,1079,1412,1437]" pageId="2" pageNumber="357" rank="species">
<emphasis id="4ADAEA8E4054FFAC9AB6FA20CE28FA4C" bold="true" box="[1015,1079,1412,1437]" pageId="2" pageNumber="357">n. sp.</emphasis>
</taxonomicNameLabel>
, however, can be differentiated from 
<taxonomicName id="BFAE4D1F4054FFAC9995FA0DCB7DFA11" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[212,354,1448,1472]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9995FA0DCB7DFA11" box="[212,354,1448,1472]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
by several characters: the relatively larger cornea (
<figureCitation id="E0952A194054FFAC9AE2FA03CE07FA11" box="[931,1048,1447,1472]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">Fig. 5K, L</figureCitation>
versus 
<figureCitation id="E0952A194054FFAC9D2FFA03CECAFA11" box="[1134,1237,1447,1472]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">
Fig. 
<date id="0C10105C4054FFAC9DE0FA03CEA4FA11" box="[1185,1211,1447,1472]" pageId="2" pageNumber="357">5I</date>
, J
</figureCitation>
); the less swollen ocular peduncle, of which the median part lacks a ridge (
<figureCitation id="E0952A194054FFAC9A7AFA68C98CFA34" box="[827,915,1484,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">Fig. 5L</figureCitation>
) (vs. distinctly more swollen with a distinct median ridge in 
<taxonomicName id="BFAE4D1F4054FFAC9810FA55CBC0F9D9" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[337,479,1520,1544]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9810FA55CBC0F9D9" box="[337,479,1520,1544]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
, 
<figureCitation id="E0952A194054FFAC98AFFA4BC821F9D9" box="[494,574,1519,1544]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">Fig. 5J</figureCitation>
); the teeth on the cutting edges of the chela fingers are relatively smaller and less prominent in the new species than in 
<taxonomicName id="BFAE4D1F4054FFAC9B24F9B1C8EDF9FD" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[613,754,1556,1580]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9B24F9B1C8EDF9FD" box="[613,754,1556,1580]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
(
<figureCitation id="E0952A194054FFAC9A40F9B0C948F9FD" box="[769,855,1556,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3H</figureCitation>
vs. 
<figureCitation id="E0952A194054FFAC9AC4F9B0C9C3F9FD" box="[901,988,1556,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3D</figureCitation>
); the proportionately shorter male third ambulatory leg with the ratio of the total leg length to CW 2.1–2.6 (
<figureCitation id="E0952A194054FFAC9AF6F993CE0EF981" box="[951,1041,1591,1616]" captionStart="FIGURE 4" captionStartId="6.[151,250,1966,1988]" captionTargetBox="[240,1347,193,1932]" captionTargetId="figure@6.[240,1347,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Right ambulatory legs. A, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633)." httpUri="https://zenodo.org/record/1440644/files/figure.png" pageId="2" pageNumber="357">Fig. 4B</figureCitation>
) (vs. proportionately longer in 
<taxonomicName id="BFAE4D1F4054FFAC9CC3F99DCB1BF9A5" baseAuthorityName="Ihle" baseAuthorityYear="1912" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9CC3F99DCB1BF9A5" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
the ratio being 2.5–2.7, 
<figureCitation id="E0952A194054FFAC9B68F9F8C899F9A5" box="[553,646,1628,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,1966,1988]" captionTargetBox="[240,1347,193,1932]" captionTargetId="figure@6.[240,1347,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Right ambulatory legs. A, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633)." httpUri="https://zenodo.org/record/1440644/files/figure.png" pageId="2" pageNumber="357">Fig. 4A</figureCitation>
); the male pleon has the lateral margins of somites 4 and 5 slightly convex in the new species (
<figureCitation id="E0952A194054FFAC988EF9DBC83BF949" box="[463,548,1663,1688]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3E</figureCitation>
) (vs. straight in 
<taxonomicName id="BFAE4D1F4054FFAC9B9FF925C972F949" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[734,877,1664,1688]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9B9FF925C972F949" box="[734,877,1664,1688]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
, 
<figureCitation id="E0952A194054FFAC9A3DF9DBC9C9F949" box="[892,982,1663,1688]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3A</figureCitation>
); the female pleon has a relatively more rounded lateral margin with the lateral margins of somite 5 distinctly convex in the new species (
<figureCitation id="E0952A194054FFAC9D9FF900CF2DF96D" box="[1246,1330,1700,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3F</figureCitation>
) (vs. less rounded and the lateral margins of somite 5 less convex in 
<taxonomicName id="BFAE4D1F4054FFAC9A71F96DC9A0F931" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[816,959,1736,1760]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9A71F96DC9A0F931" box="[816,959,1736,1760]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
, 
<figureCitation id="E0952A194054FFAC9A8FF963CE3AF931" box="[974,1061,1735,1760]" captionStart="FIGURE 3" captionStartId="5.[151,250,1886,1908]" captionTargetBox="[150,1435,311,1848]" captionTargetId="figure@5.[150,1435,311,1848]" captionTargetPageId="5" captionText="FIGURE 3. A, D, Karstarma jacobsoni (Ihle, 1912), male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, K. jacobsoni (Ihle, 1912), female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, K. malang n. sp., paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela." httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="2" pageNumber="357">Fig. 3B</figureCitation>
); and the relatively more slender G1, with the width of the distal chitinous tip less than half of that of the main G1 shaft in the new species (
<figureCitation id="E0952A194054FFAC9C2FF948CAC5F8F9" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">Fig. 5E–H</figureCitation>
) (vs. G1 is more robust with a wider distal chitinous tip which is more than half of the main shaft in 
<taxonomicName id="BFAE4D1F4054FFAC9CC3F8B5CB1AF89D" baseAuthorityName="Ihle" baseAuthorityYear="1912" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9CC3F8B5CB1AF89D" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
, 
<figureCitation id="E0952A194054FFAC9852F890CB8EF89C" box="[275,401,1844,1869]" captionStart="FIGURE 5" captionStartId="7.[151,250,1933,1955]" captionTargetBox="[171,1412,219,1892]" captionTargetId="figure@7.[151,1436,193,1913]" captionTargetPageId="7" captionText="FIGURE 5. A–D, I, J, Karstarma jacobsoni (Ihle, 1912), male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm." httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="2" pageNumber="357">Fig. 5A–D</figureCitation>
). The good series of specimens of both species confirm the validity of these characters.
</paragraph>
<paragraph id="7811369C4054FFAB9986F8F3CF20FF29" blockId="2.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="360" pageId="2" pageNumber="357">
The living colours of 
<taxonomicName id="BFAE4D1F4054FFAC98FFF8FDC88CF8A1" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[446,659,1880,1904]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4054FFAC98FFF8FDC88CF8A1" box="[446,659,1880,1904]" italics="true" pageId="2" pageNumber="357">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54054FFAC9BD8F8F3C8C8F8A1" box="[665,727,1879,1904]" pageId="2" pageNumber="357" rank="species">
<emphasis id="4ADAEA8E4054FFAC9BD8F8F3C8C8F8A1" bold="true" box="[665,727,1879,1904]" pageId="2" pageNumber="357">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="BFAE4D1F4054FFAC9A4CF8FDC985F8A1" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[781,922,1880,1904]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9A4CF8FDC985F8A1" box="[781,922,1880,1904]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
are also different; the new species has opaque light brown colour (
<figureCitation id="E0952A194054FFAC983CF8D8CBCAF844" box="[381,469,1916,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1888,1910]" captionTargetBox="[151,1437,193,1848]" captionTargetId="figure@8.[151,1437,193,1848]" captionTargetPageId="8" captionText="FIGURE 6. Colour in life. A, Karstarma jacobsoni (Ihle, 1912), specimen not collected; B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). Photographs by C. Rahmadi." httpUri="https://zenodo.org/record/1440648/files/figure.png" pageId="2" pageNumber="357">Fig. 6B</figureCitation>
) while 
<taxonomicName id="BFAE4D1F4054FFAC9B6FF8D9C8A2F845" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[558,701,1916,1940]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4054FFAC9B6FF8D9C8A2F845" box="[558,701,1916,1940]" italics="true" pageId="2" pageNumber="357">K. jacobsoni</emphasis>
</taxonomicName>
is a more translucent white (
<figureCitation id="E0952A194054FFAC9D4FF8D8CE75F844" box="[1038,1130,1916,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1888,1910]" captionTargetBox="[151,1437,193,1848]" captionTargetId="figure@8.[151,1437,193,1848]" captionTargetPageId="8" captionText="FIGURE 6. Colour in life. A, Karstarma jacobsoni (Ihle, 1912), specimen not collected; B, K. malang n. sp., holotype male (18.3 x 16.7 mm) (MZB Cru 4633). Photographs by C. Rahmadi." httpUri="https://zenodo.org/record/1440648/files/figure.png" pageId="2" pageNumber="357">Fig. 6A</figureCitation>
). Another cave species, 
<taxonomicName id="BFAE4D1F4054FFAC9CC3F8D9CB53F869" baseAuthorityName="Naruse &amp; Ng" baseAuthorityYear="2007" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="microphthalmus">
<emphasis id="4ADAEA8E4054FFAC9CC3F8D9CB53F869" italics="true" pageId="2" pageNumber="357">K. microphthalmus</emphasis>
</taxonomicName>
, from southern 
<collectingRegion id="BA6AF87E4054FFAC9B4BF804C86DF868" box="[522,626,1952,1977]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="357">Sulawesi</collectingRegion>
, is also yellowish-cream in color (
<bibRefCitation id="1C3F4B6D4054FFAC9D48F804CEEFF869" box="[1033,1264,1952,1977]" pageId="2" pageNumber="357" refId="ref4396" refString="Naruse, T. &amp; Ng, P. K. L. (2007) On a new species of cavernicolous crab of the genus Sesarmoides Serene &amp; Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae) from Sulawesi, Indonesia. Raffles Bulletin of Zoology, 55 (1), 127 - 130." type="journal article">Naruse &amp; Ng 2007</bibRefCitation>
). All the other known 
<taxonomicName id="BFAE4D1F4054FFAC99ABF861CB7CF80D" authorityName="Davie &amp; Ng" authorityYear="2007" box="[234,355,1989,2012]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E4054FFAC99ABF861CB7CF80D" box="[234,355,1989,2012]" italics="true" pageId="2" pageNumber="357">Karstarma</emphasis>
</taxonomicName>
species are distinctly coloured: 
<taxonomicName id="BFAE4D1F4054FFAC9B88F861C936F80D" authorityName="Wowor &amp; Ng" authorityYear="2009" box="[713,809,1988,2012]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="ardea">
<emphasis id="4ADAEA8E4054FFAC9B88F861C936F80D" box="[713,809,1988,2012]" italics="true" pageId="2" pageNumber="357">K. ardea</emphasis>
</taxonomicName>
, 
<taxonomicName id="BFAE4D1F4054FFAC9A76F861C9E7F80D" authorityName="Husana, Naruse &amp; Kase" authorityYear="2010" box="[823,1016,1988,2012]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="4ADAEA8E4054FFAC9A76F861C9E7F80D" box="[823,1016,1988,2012]" italics="true" pageId="2" pageNumber="357">K. philippinarum</emphasis>
</taxonomicName>
and 
<taxonomicName id="BFAE4D1F4054FFAC9D75F861CEAFF80D" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" box="[1076,1200,1988,2012]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="357" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="4ADAEA8E4054FFAC9D75F861CEAFF80D" box="[1076,1200,1988,2012]" italics="true" pageId="2" pageNumber="357">K. ultrapes</emphasis>
</taxonomicName>
have bright orange carapaces (
<bibRefCitation id="1C3F4B6D4053FFAB9852FF3CCBA4FF61" box="[275,443,152,176]" pageId="5" pageNumber="360" refId="ref4683" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1994) Sesarmoides ultrapes new species, a remarkable sesarmine crab from caves in the Solomon Islands (Decapoda: Brachyura: Grapsidae). Crustacean Research, 23, 12 - 22. https: // doi. org / 10.18353 / crustacea. 23.0 _ 12" type="journal article">
Ng 
<emphasis id="4ADAEA8E4053FFAB987EFF3DCB68FF61" box="[319,375,152,176]" italics="true" pageId="5" pageNumber="360">et al.</emphasis>
1994
</bibRefCitation>
; 
<bibRefCitation id="1C3F4B6D4053FFAB9886FF3CC8BBFF61" box="[455,676,152,176]" pageId="5" pageNumber="360" refId="ref5145" refString="Wowor, D. &amp; Ng, P. K. L. (2009) Two new species of sesarmid crabs (Crustacea: Decapoda: Brachyura) associated with limestone formations in West Papua, Indonesia. Zootaxa, 2025, 21 - 31." type="journal article">Wowor &amp; Ng 2009</bibRefCitation>
; 
<bibRefCitation id="1C3F4B6D4053FFAB9BF1FF3CC996FF61" box="[688,905,152,176]" pageId="5" pageNumber="360" refId="ref4233" refString="Husana, D. E. M., Naruse, T. &amp; Kase, T. (2010) A new species of the genus Karstarma (Crustacea: Decapoda: Brachyura: Sesarmidae) from anchialine caves in the Philippines. Raffles Bulletin of Zoology, 58 (1), 51 - 55." type="journal article">
Husana 
<emphasis id="4ADAEA8E4053FFAB9A4DFF3DC95AFF61" box="[780,837,152,176]" italics="true" pageId="5" pageNumber="360">et al.</emphasis>
2010
</bibRefCitation>
), while the dorsal carapace surfaces of most of the other 
<taxonomicName id="BFAE4D1F4053FFAB99BEFF19CB67FF05" authorityName="Davie &amp; Ng" authorityYear="2007" box="[255,376,189,212]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E4053FFAB99BEFF19CB67FF05" box="[255,376,189,212]" italics="true" pageId="5" pageNumber="360">Karstarma</emphasis>
</taxonomicName>
species are light orange to orangish-brown or brown to dark purplish-brown (
<bibRefCitation id="1C3F4B6D4053FFAB9D9BFF18CF5FFF05" box="[1242,1344,188,212]" pageId="5" pageNumber="360" refId="ref4572" refString="Ng, P. K. L. (2002) New species of cavernicolous crabs of the genus Sesarmoides from the western Pacific, with a key to the genus (Crustacea: Decapoda: Brachyura: Sesarmidae). Raffles Bulletin of Zoology, 50 (2), 419 - 435." type="journal article">Ng 2002</bibRefCitation>
; 
<bibRefCitation id="1C3F4B6D4053FFAB9C0BFF19CB07FF29" pageId="5" pageNumber="360" refId="ref5145" refString="Wowor, D. &amp; Ng, P. K. L. (2009) Two new species of sesarmid crabs (Crustacea: Decapoda: Brachyura) associated with limestone formations in West Papua, Indonesia. Zootaxa, 2025, 21 - 31." type="journal article">Wowor &amp; Ng 2009</bibRefCitation>
; C. Rahmadi, pers. comm.). The living colours of 
<taxonomicName id="BFAE4D1F4053FFAB9A0AFF45CE14FF29" baseAuthorityName="Ng" baseAuthorityYear="1988" box="[843,1035,224,248]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="novabritannia">
<emphasis id="4ADAEA8E4053FFAB9A0AFF45CE14FF29" box="[843,1035,224,248]" italics="true" pageId="5" pageNumber="360">K. novabritannia</emphasis>
</taxonomicName>
and 
<taxonomicName id="BFAE4D1F4053FFAB9D00FF45CE8FFF29" baseAuthorityName="Ng" baseAuthorityYear="2002" box="[1089,1168,224,248]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="sulu">
<emphasis id="4ADAEA8E4053FFAB9D00FF45CE8FFF29" box="[1089,1168,224,248]" italics="true" pageId="5" pageNumber="360">K. sulu</emphasis>
</taxonomicName>
are not known.
</paragraph>
<caption id="2CD166144055FFAD99D6F81EC851F83E" httpUri="https://zenodo.org/record/1440638/files/figure.png" pageId="3" pageNumber="358" startId="3.[151,250,1978,2000]" targetBox="[257,1318,195,1934]" targetPageId="3">
<paragraph id="7811369C4055FFAD99D6F81EC851F83E" blockId="3.[151,1435,1978,2031]" pageId="3" pageNumber="358">
<emphasis id="4ADAEA8E4055FFAD99D6F81ECB12F801" bold="true" box="[151,269,1978,2000]" pageId="3" pageNumber="358">FIGURE 1</emphasis>
. 
<taxonomicName id="BFAE4D1F4055FFAD9858F81FC86FF801" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[281,624,1978,2000]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="358" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4055FFAD9858F81FCBF0F801" box="[281,495,1978,2000]" italics="true" pageId="3" pageNumber="358">Karstarma jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, male (20.7 x 19.4 mm) (ZRC 2009.0807). A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax.
</paragraph>
</caption>
<caption id="2CD166144052FFAA99D6F817C851F839" httpUri="https://zenodo.org/record/1440640/files/figure.png" pageId="4" pageNumber="359" startId="4.[151,250,1971,1993]" targetBox="[257,1323,193,1932]" targetPageId="4">
<paragraph id="7811369C4052FFAA99D6F817C851F839" blockId="4.[151,1435,1971,2024]" pageId="4" pageNumber="359">
<emphasis id="4ADAEA8E4052FFAA99D6F817CB13F818" bold="true" box="[151,268,1971,1993]" pageId="4" pageNumber="359">FIGURE 2</emphasis>
. 
<taxonomicName id="BFAE4D1F4052FFAA9858F810CBC6F818" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[281,473,1971,1993]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="359" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4052FFAA9858F810CBC6F818" box="[281,473,1971,1993]" italics="true" pageId="4" pageNumber="359">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54052FFAA989EF817C807F818" box="[479,536,1971,1993]" pageId="4" pageNumber="359" rank="species">
<emphasis id="4ADAEA8E4052FFAA989EF817C807F818" bold="true" box="[479,536,1971,1993]" pageId="4" pageNumber="359">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax.
</paragraph>
</caption>
<caption id="2CD166144053FFAB99D6F8FACF06F801" httpUri="https://zenodo.org/record/1440642/files/figure.png" pageId="5" pageNumber="360" startId="5.[151,250,1886,1908]" targetBox="[150,1435,311,1848]" targetPageId="5">
<paragraph id="7811369C4053FFAB99D6F8FACF06F801" blockId="5.[151,1437,1886,2000]" pageId="5" pageNumber="360">
<emphasis id="4ADAEA8E4053FFAB99D6F8FACB13F8A5" bold="true" box="[151,268,1886,1908]" pageId="5" pageNumber="360">FIGURE 3</emphasis>
. A, D, 
<taxonomicName id="BFAE4D1F4053FFAB9815F8FBC8B6F8A5" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[340,681,1886,1908]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4053FFAB9815F8FBC834F8A5" box="[340,555,1886,1908]" italics="true" pageId="5" pageNumber="360">Karstarma jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, male (20.7 x 19.4 mm) (ZRC 2009.0807); B, C, 
<taxonomicName id="BFAE4D1F4053FFAB9DD8F8FBCF88F8A5" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[1177,1431,1886,1908]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4053FFAB9DD8F8FBCF07F8A5" box="[1177,1304,1886,1908]" italics="true" pageId="5" pageNumber="360">K. jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, female (21.1 x 18.8 mm) (ZRC 2009.0807) E, H, 
<taxonomicName id="BFAE4D1F4053FFAB9BD1F8DAC8E5F843" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[656,762,1916,1939]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4053FFAB9BD1F8DAC8E5F843" box="[656,762,1916,1939]" italics="true" pageId="5" pageNumber="360">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54053FFAB9A41F8D8C925F843" box="[768,826,1916,1938]" pageId="5" pageNumber="360" rank="species">
<emphasis id="4ADAEA8E4053FFAB9A41F8D8C925F843" bold="true" box="[768,826,1916,1938]" pageId="5" pageNumber="360">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (18.3 x 16.7 mm) (MZB Cru 4633); F, G, 
<taxonomicName id="BFAE4D1F4053FFAB9CC5F8DACAFCF860" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="360" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4053FFAB9CC5F8DACAFCF860" italics="true" pageId="5" pageNumber="360">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54053FFAB99ADF83FCB37F860" box="[236,296,1947,1969]" pageId="5" pageNumber="360" rank="species">
<emphasis id="4ADAEA8E4053FFAB99ADF83FCB37F860" bold="true" box="[236,296,1947,1969]" pageId="5" pageNumber="360">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype female (18.8 x 16.7 mm) (MZB Cru 4635). A, E, male anterior thoracic sternum and pleon; B, F, female thoracic sternum and pleon; C, G, female thoracic sternum and vulvae; D, H, outer view of left male chela.
</paragraph>
</caption>
<caption id="2CD166144050FFA899D6F80AC92FF833" httpUri="https://zenodo.org/record/1440644/files/figure.png" pageId="6" pageNumber="361" startId="6.[151,250,1966,1988]" targetBox="[240,1347,193,1932]" targetPageId="6">
<paragraph id="7811369C4050FFA899D6F80AC92FF833" blockId="6.[151,1437,1966,2018]" pageId="6" pageNumber="361">
<emphasis id="4ADAEA8E4050FFA899D6F80ACB10F815" bold="true" box="[151,271,1966,1988]" pageId="6" pageNumber="361">FIGURE 4</emphasis>
. Right ambulatory legs. A, 
<taxonomicName id="BFAE4D1F4050FFA89B77F80BC98BF815" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[566,916,1966,1988]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4050FFA89B77F80BC90FF815" box="[566,784,1966,1988]" italics="true" pageId="6" pageNumber="361">Karstarma jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, male (20.7 x 19.4 mm) (ZRC 2009.0807); B, 
<taxonomicName id="BFAE4D1F4050FFA89CC5F80BCAFCF833" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4050FFA89CC5F80BCAFCF833" italics="true" pageId="6" pageNumber="361">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54050FFA899A8F868CB3DF833" box="[233,290,1996,2018]" pageId="6" pageNumber="361" rank="species">
<emphasis id="4ADAEA8E4050FFA899A8F868CB3DF833" bold="true" box="[233,290,1996,2018]" pageId="6" pageNumber="361">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (18.3 x 16.7 mm) (MZB Cru 4633).
</paragraph>
</caption>
<caption id="2CD166144051FFA999D6F829CF83F830" httpUri="https://zenodo.org/record/1440646/files/figure.png" pageId="7" pageNumber="362" startId="7.[151,250,1933,1955]" targetBox="[171,1412,219,1892]" targetPageId="7">
<paragraph id="7811369C4051FFA999D6F829CF83F830" blockId="7.[151,1436,1933,2017]" pageId="7" pageNumber="362">
<emphasis id="4ADAEA8E4051FFA999D6F829CB11F872" bold="true" box="[151,270,1933,1955]" pageId="7" pageNumber="362">FIGURE 5</emphasis>
. A–D, I, J, 
<taxonomicName id="BFAE4D1F4051FFA998C6F82BC8FFF872" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[391,736,1933,1956]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E4051FFA998C6F82BC840F872" box="[391,607,1933,1956]" italics="true" pageId="7" pageNumber="362">Karstarma jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, male (18.6 x 16.7 mm) (MZB Cru 1590); E–H, K, L, 
<taxonomicName id="BFAE4D1F4051FFA99C54F82BCF9FF872" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[1301,1408,1933,1956]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E4051FFA99C54F82BCF9FF872" box="[1301,1408,1933,1956]" italics="true" pageId="7" pageNumber="362">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F54051FFA99CC6F829CAAAF813" pageId="7" pageNumber="362" rank="species">
<emphasis id="4ADAEA8E4051FFA99CC6F829CAAAF813" bold="true" pageId="7" pageNumber="362">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (18.3 x 16.7 mm) (MZB Cru 4633). A, B, E, F, ventral and dorsal views of G1; D, H, tips of G1, ventral view; I, K, dorsal view of eyes; J, L, frontal view of eyes. Scale bars: A–C, E–G = 2.0 mm; D, H = 1.0 mm; I–J, K–L = 5.0 mm.
</paragraph>
</caption>
<caption id="2CD16614405EFFA699D6F8C4C933F844" httpUri="https://zenodo.org/record/1440648/files/figure.png" pageId="8" pageNumber="363" startId="8.[151,250,1888,1910]" targetBox="[151,1437,193,1848]" targetPageId="8">
<paragraph id="7811369C405EFFA699D6F8C4C933F844" blockId="8.[151,1436,1888,1941]" pageId="8" pageNumber="363">
<emphasis id="4ADAEA8E405EFFA699D6F8C4CB11F8A7" bold="true" box="[151,270,1888,1910]" pageId="8" pageNumber="363">FIGURE 6</emphasis>
. Colour in life. A, 
<taxonomicName id="BFAE4D1F405EFFA69895F8C5C932F8A7" authority="(Ihle, 1912)" authorityName="Ihle" authorityYear="1912" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[468,813,1888,1910]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E405EFFA69895F8C5C8B4F8A7" box="[468,683,1888,1910]" italics="true" pageId="8" pageNumber="363">Karstarma jacobsoni</emphasis>
(Ihle, 1912)
</taxonomicName>
, specimen not collected; B, 
<taxonomicName id="BFAE4D1F405EFFA69D10F8C5CEA4F8A7" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[1105,1211,1888,1910]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405EFFA69D10F8C5CEA4F8A7" box="[1105,1211,1888,1910]" italics="true" pageId="8" pageNumber="363">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405EFFA69D82F8C4CEE2F8A7" box="[1219,1277,1888,1910]" pageId="8" pageNumber="363" rank="species">
<emphasis id="4ADAEA8E405EFFA69D82F8C4CEE2F8A7" bold="true" box="[1219,1277,1888,1910]" pageId="8" pageNumber="363">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (18.3 x 16.7 mm) (MZB Cru 4633). Photographs by C. Rahmadi.
</paragraph>
</caption>
<paragraph id="7811369C405FFFA79986FF3DC98FFDC9" blockId="9.[151,1437,151,1293]" pageId="9" pageNumber="364">
<taxonomicName id="BFAE4D1F405FFFA79986FF3DCBA7FF61" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[199,440,152,176]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E405FFFA79986FF3DCBA7FF61" box="[199,440,152,176]" italics="true" pageId="9" pageNumber="364">Karstarma jacobsoni</emphasis>
</taxonomicName>
is known to have a large egg size (
<quantity id="BF569B79405FFFA79A1EFF3CC9A5FF7E" box="[863,954,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="364" unit="mm" value="1.8">1.8 mm</quantity>
diameter, present study, see Comparative Material) and almost certainly has an abbreviated larval development with the eggs probably hatching out into juvenile crabs. While no ovigerous specimens of 
<taxonomicName id="BFAE4D1F405FFFA79B8BFF45C95FFF29" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[714,832,224,248]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79B8BFF45C95FFF29" box="[714,832,224,248]" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA79A08FF7BC995FF29" box="[841,906,223,248]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA79A08FF7BC995FF29" bold="true" box="[841,906,223,248]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
were collected, but the relatively large size of their vulvae suggests their eggs are probably of a similar size to that of 
<taxonomicName id="BFAE4D1F405FFFA79A80FEA1CE4FFECD" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[961,1104,260,284]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E405FFFA79A80FEA1CE4FFECD" box="[961,1104,260,284]" italics="true" pageId="9" pageNumber="364">K. jacobsoni</emphasis>
</taxonomicName>
. This suggests that these two species are probably land-locked which do not need to migrate to the sea to release their larvae. Considering that the two 
<taxonomicName id="BFAE4D1F405FFFA799B4FEE9CB71FEB5" authorityName="Davie &amp; Ng" authorityYear="2007" box="[245,366,333,356]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E405FFFA799B4FEE9CB71FEB5" box="[245,366,333,356]" italics="true" pageId="9" pageNumber="364">Karstarma</emphasis>
</taxonomicName>
species from 
<collectingRegion id="BA6AF87E405FFFA79B53FEE9C85BFEB5" box="[530,580,333,356]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="364">Java</collectingRegion>
belong to different underground watershed systems and karst ranges, their populations are effectively isolated, geographically and reproductively from each other. It is important to note that all the other coloured 
<taxonomicName id="BFAE4D1F405FFFA798D2FE31C813FE7D" authorityName="Davie &amp; Ng" authorityYear="2007" box="[403,524,405,428]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E405FFFA798D2FE31C813FE7D" box="[403,524,405,428]" italics="true" pageId="9" pageNumber="364">Karstarma</emphasis>
</taxonomicName>
species occur near or adjacent to the sea shore, or in anchialine systems which, though inland, are still connected to sea. Most are restricted to small islands. Many of these species inhabiting near sea shores (e.g., 
<taxonomicName id="BFAE4D1F405FFFA79810FE79CBD0FE25" baseAuthorityName="Balss" baseAuthorityYear="1934" box="[337,463,476,500]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="jacksoni">
<emphasis id="4ADAEA8E405FFFA79810FE79CBD0FE25" box="[337,463,476,500]" italics="true" pageId="9" pageNumber="364">K. jacksoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="BFAE4D1F405FFFA7989FFE79C876FE25" baseAuthorityName="Ng" baseAuthorityYear="2002" box="[478,617,476,500]" class="Malacostraca" family="Sesarmidae" genus="Sesarmoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="boholano">
<emphasis id="4ADAEA8E405FFFA7989FFE79C876FE25" box="[478,617,476,500]" italics="true" pageId="9" pageNumber="364">K. boholano</emphasis>
</taxonomicName>
, 
<taxonomicName id="BFAE4D1F405FFFA79B36FE79C917FE25" baseAuthorityName="Ng" baseAuthorityYear="2002" box="[631,776,477,500]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="guamense">
<emphasis id="4ADAEA8E405FFFA79B36FE79C917FE25" box="[631,776,477,500]" italics="true" pageId="9" pageNumber="364">K. guamense</emphasis>
</taxonomicName>
, 
<taxonomicName id="BFAE4D1F405FFFA79A56FE79C971FE25" baseAuthorityName="Ng &amp; Whitten" baseAuthorityYear="1995" box="[791,878,476,500]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="emdi">
<emphasis id="4ADAEA8E405FFFA79A56FE79C971FE25" box="[791,878,476,500]" italics="true" pageId="9" pageNumber="364">K. emdi</emphasis>
</taxonomicName>
and 
<taxonomicName id="BFAE4D1F405FFFA79AE4FE79CE08FE25" box="[933,1047,476,500]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="balicus">
<emphasis id="4ADAEA8E405FFFA79AE4FE79CE08FE25" box="[933,1047,476,500]" italics="true" pageId="9" pageNumber="364">K. balicus</emphasis>
</taxonomicName>
) have small eggs and release their zoeae into the open sea (
<bibRefCitation id="1C3F4B6D405FFFA798E8FE5BC99CFDC9" box="[425,899,511,536]" pageId="9" pageNumber="364" refId="ref4883" refString="Orchard, M. (2012) Crabs of Christmas Island. Christmas Island Natural History Association, 287 pp." type="book">Orchard 2012; PKL Ng, unpublished data</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="30B46517405FFFA79986FD80C955FB71" pageId="9" pageNumber="364" type="description">
<paragraph id="7811369C405FFFA79986FD80CECCFC75" blockId="9.[151,1437,151,1293]" pageId="9" pageNumber="364">
<emphasis id="4ADAEA8E405FFFA79986FD80CB93FDEC" bold="true" box="[199,396,548,573]" pageId="9" pageNumber="364">Notes on habitat</emphasis>
. Both 
<taxonomicName id="BFAE4D1F405FFFA7989BFD81C8D6FDED" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[474,713,548,572]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E405FFFA7989BFD81C8D6FDED" box="[474,713,548,572]" italics="true" pageId="9" pageNumber="364">Karstarma jacobsoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="BFAE4D1F405FFFA79A40FD81C968FDED" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[769,887,548,572]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79A40FD81C968FDED" box="[769,887,548,572]" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA79A3EFD80C9A0FDEC" box="[895,959,548,573]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA79A3EFD80C9A0FDEC" bold="true" box="[895,959,548,573]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
inhabit the banks of underground streams in the totally dark zone of cave systems (C. Rahmadi, pers. comm.) where the water has zero salinity and the habitats are far from the sea. The 
<typeStatus id="A715883E405FFFA79B55FDC9C85BFD54" box="[532,580,621,645]" pageId="9" pageNumber="364">type</typeStatus>
locality of 
<taxonomicName id="BFAE4D1F405FFFA79B86FDC9C923FD55" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[711,828,620,644]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79B86FDC9C923FD55" box="[711,828,620,644]" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA79A05FDC8C99CFD54" box="[836,899,620,645]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA79A05FDC8C99CFD54" bold="true" box="[836,899,620,645]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
is about three to six km from southern coast of 
<collectingRegion id="BA6AF87E405FFFA799D6FD34CAD6FD76" box="[151,201,656,679]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="364">Java</collectingRegion>
while that of 
<taxonomicName id="BFAE4D1F405FFFA7982BFD35CBE6FD79" baseAuthorityName="Ihle" baseAuthorityYear="1912" box="[362,505,656,680]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="jacobsoni">
<emphasis id="4ADAEA8E405FFFA7982BFD35CBE6FD79" box="[362,505,656,680]" italics="true" pageId="9" pageNumber="364">K. jacobsoni</emphasis>
</taxonomicName>
is further far inland, about 
<quantity id="BF569B79405FFFA79A75FD34C9B5FD79" box="[820,938,655,680]" metricMagnitude="4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="364" unit="km" value="25.0" valueMax="30.0" valueMin="20.0">20–30 km</quantity>
; with the two sites some 
<quantity id="BF569B79405FFFA79D8AFD34CF3DFD79" box="[1227,1314,655,680]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="364" unit="km" value="200.0">200 km</quantity>
from each other (
<figureCitation id="E0952A19405FFFA799A0FD10CB38FD1D" box="[225,295,692,717]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1852]" captionTargetBox="[170,1426,1393,1803]" captionTargetId="figure@9.[150,1436,1348,1809]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Karstarma jacobsoni (Ihle, 1912) and K. malang n. sp., in Java Island, Indonesia. Map prepared by C. Rahmadi." httpUri="https://zenodo.org/record/1440650/files/figure.png" pageId="9" pageNumber="364">Fig. 7</figureCitation>
). 
<taxonomicName id="BFAE4D1F405FFFA7987CFD11C80CFD1D" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[317,531,692,716]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA7987CFD11C80CFD1D" box="[317,531,692,716]" italics="true" pageId="9" pageNumber="364">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA79B5DFD10C843FD1C" box="[540,604,692,717]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA79B5DFD10C843FD1C" bold="true" box="[540,604,692,717]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
was found in three vertical caves where underground freshwater streams were present. Based on the information provided by Cahyo Rahmadi, 
<taxonomicName id="BFAE4D1F405FFFA79AF5FD7DCE36FD21" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[948,1065,728,752]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79AF5FD7DCE36FD21" box="[948,1065,728,752]" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
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<emphasis id="4ADAEA8E405FFFA79D73FD73CE6DFD21" bold="true" box="[1074,1138,727,752]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
is an obligate cavernicole living in total dark zone, i.e., it is a stygobite. It was found along the banks of the underground streams on the muddy cave floor (
<bibRefCitation id="1C3F4B6D405FFFA79837FCBBC82DFCE9" box="[374,562,799,824]" pageId="9" pageNumber="364" refId="ref4954" refString="Rahmadi, C. (2011 b) Memburu kepiting gua (Karstarma sp.) di Malang Selatan. Indonesian Caves Life, 25 Oktober, 2011. Available from: https: // biotagua. org / 2011 / 10 / 25 / kepiting-malang / # more- 2336 (accessed 29 Agustus 2017)" type="journal article">Rahmadi 2011b</bibRefCitation>
). Gua Lowo (lowo or lawa means bat in Javanese), the 
<typeStatus id="A715883E405FFFA79D86FC84CEE8FCE9" box="[1223,1271,800,824]" pageId="9" pageNumber="364">type</typeStatus>
locality of 
<taxonomicName id="BFAE4D1F405FFFA79CC3FC85CAF4FC8D" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79CC3FC85CAF4FC8D" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA799B5FCE0CB2BFC8C" box="[244,308,836,861]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA799B5FCE0CB2BFC8C" bold="true" box="[244,308,836,861]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
is a roosting ground for bats during the day in the middle of forest. Arachnids of the genus 
<taxonomicName id="BFAE4D1F405FFFA79C1AFCE0CF83FC8A" authorityName="Simon" authorityYear="1892" box="[1371,1436,836,859]" class="Arachnida" family="Charinidae" genus="Sarax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E405FFFA79C1AFCE0CF83FC8A" box="[1371,1436,836,859]" italics="true" pageId="9" pageNumber="364">Sarax</emphasis>
</taxonomicName>
(
<taxonomicName id="BFAE4D1F405FFFA799DFFCC3CB3BFC51" authorityName="Thorell" authorityYear="1883" box="[158,292,871,896]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="364" phylum="Arthropoda" rank="order">Amblypygi</taxonomicName>
: 
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) and 
<taxonomicName id="BFAE4D1F405FFFA798B7FCCCC864FC51" box="[502,635,872,896]" class="Arachnida" family="Sparassidae" genus="Heteropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="4ADAEA8E405FFFA798B7FCCCC864FC51" box="[502,635,872,896]" italics="true" pageId="9" pageNumber="364">Heteropoda</emphasis>
</taxonomicName>
(
<taxonomicName id="BFAE4D1F405FFFA79BCFFCCCC8EEFC51" box="[654,753,872,896]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="364" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
: 
<taxonomicName id="BFAE4D1F405FFFA79BBEFCC3C99AFC51" authorityName="Bertkau" authorityYear="1872" box="[767,901,871,896]" class="Arachnida" family="Sparassidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="364" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
) were also present, while 
<taxonomicName id="BFAE4D1F405FFFA79DFEFCC3CF2EFC51" authorityName="" authorityYear="1802" box="[1215,1329,871,896]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
(Insecta: 
<taxonomicName id="BFAE4D1F405FFFA799D6FC28CB0BFC74" authorityName="" authorityYear="1758" box="[151,276,908,933]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) and Isopoda (
<taxonomicName id="BFAE4D1F405FFFA798F8FC28C834FC75" box="[441,555,908,932]" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crustacea" pageId="9" pageNumber="364" phylum="Arthropoda" rank="order">Crustacea</taxonomicName>
) were found in the guano on the cave floor (
<bibRefCitation id="1C3F4B6D405FFFA79D5DFC28CED8FC75" box="[1052,1223,908,933]" pageId="9" pageNumber="364" refId="ref5058" refString="Rahmadi, C. (2012) Sekilas invertebrata gua di Malang Selatan. Indonesian Caves Life, 17 February 2012. Available from: https: // biotagua. org / 2012 / 02 / 17 / invertebrat-malang (accessed 29 Agustus 2017)" type="journal article">Rahmadi 2012</bibRefCitation>
).
</paragraph>
<paragraph id="7811369C405FFFA79986FC0BC955FB71" blockId="9.[151,1437,151,1293]" pageId="9" pageNumber="364">
There are three other caves in the southern Malang karst range that have been explored the fauna: Gua Siem, Gua Lo and Gua Bangi (
<bibRefCitation id="1C3F4B6D405FFFA798F7FC70C872FC3D" box="[438,621,980,1005]" pageId="9" pageNumber="364" refId="ref4904" refString="Rahmadi, C. (2011 a) Malang-Menyusuri gua minim oksigen di Malang Selatan (Part 1). Indonesian Caves Life, 11 Oktober, 2011. https: // biotagua. org / 2011 / 10 / 11 / menyusuri-malang / (accessed 29 Agustus 2017)" type="journal article">Rahmadi 2011a</bibRefCitation>
, 
<bibRefCitation id="1C3F4B6D405FFFA79B3BFC70C896FC3D" box="[634,649,980,1004]" pageId="9" pageNumber="364" refId="ref5010" refString="Rahmadi, C. (2011 c) Malang-merangkak dan berendam (Part 3). Indonesian Caves Life, 10 November 2011. Available from: https: // biotagua. org / 2011 / 11 / 10 / malang-merangkak / (accessed 29 Agustus 2017)" type="journal article">c</bibRefCitation>
). These caves are mostly inhabited by terrestrial fauna such as bats, spiders, insects and isopods. Gua Lo and Gua Bangi are two connected caves and at one point between the two caves, there is subterranean groundwater with white catfish and other fish observed but no crab was seen (
<bibRefCitation id="1C3F4B6D405FFFA79C74FBB8CAC1FB89" pageId="9" pageNumber="364" refId="ref5010" refString="Rahmadi, C. (2011 c) Malang-merangkak dan berendam (Part 3). Indonesian Caves Life, 10 November 2011. Available from: https: // biotagua. org / 2011 / 11 / 10 / malang-merangkak / (accessed 29 Agustus 2017)" type="journal article">Rahmadi 2011c</bibRefCitation>
). So far, 
<taxonomicName id="BFAE4D1F405FFFA79802FBE5CBA8FB89" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[323,439,1088,1112]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79802FBE5CBA8FB89" box="[323,439,1088,1112]" italics="true" pageId="9" pageNumber="364">K. malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA798FFFB9BCBE1FB89" box="[446,510,1087,1112]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA798FFFB9BCBE1FB89" bold="true" box="[446,510,1087,1112]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
has only been found in Gua Kerek, Gua Rampal and Gua Lowo. It is possible that the subterranean streams in these caves are connected and form an underground watershed system but not linked to the subterranean system of the Gua Lo-Bangi system (
<figureCitation id="E0952A19405FFFA79BB9FB23C923FB71" box="[760,828,1159,1184]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1852]" captionTargetBox="[170,1426,1393,1803]" captionTargetId="figure@9.[150,1436,1348,1809]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Karstarma jacobsoni (Ihle, 1912) and K. malang n. sp., in Java Island, Indonesia. Map prepared by C. Rahmadi." httpUri="https://zenodo.org/record/1440650/files/figure.png" pageId="9" pageNumber="364">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="30B46517405FFFA79986FB08C9A6FADC" pageId="9" pageNumber="364" type="distribution">
<paragraph id="7811369C405FFFA79986FB08C9A6FADC" blockId="9.[151,1437,151,1293]" pageId="9" pageNumber="364">
<emphasis id="4ADAEA8E405FFFA79986FB08CB46FB14" bold="true" box="[199,345,1196,1221]" pageId="9" pageNumber="364">Distribution</emphasis>
. 
<taxonomicName id="BFAE4D1F405FFFA79826FB09C824FB15" authority="Wowor &amp; Ng, 2018" authorityName="Wowor &amp; Ng" authorityYear="2018" box="[359,571,1196,1220]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="malang" status="sp. nov.">
<emphasis id="4ADAEA8E405FFFA79826FB09C824FB15" box="[359,571,1196,1220]" italics="true" pageId="9" pageNumber="364">Karstarma malang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51E957F5405FFFA79B00FB08C89FFB14" box="[577,640,1196,1221]" pageId="9" pageNumber="364" rank="species">
<emphasis id="4ADAEA8E405FFFA79B00FB08C89FFB14" bold="true" box="[577,640,1196,1221]" pageId="9" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
is currently known only from the caves in Malang Selatan (or southern Malang) karst range of Donomulyo District, Malang Regency. The caves range about between two to ten kilometers from the Indian Ocean coast of southern 
<collectingRegion id="BA6AF87E405FFFA79B9FFB51C9AAFADC" box="[734,949,1268,1293]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="364">East Java Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>