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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner &amp; Keupp, Helmut" docDate="2022" docId="A4CD9C9FCA3B5EA994A6FF2AA47F9886" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Archaeotolithus bornholmiensis" docType="treatment" docVersion="2" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageId="0" pageNumber="51" updateTime="1649795760618" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schwarzhans, Werner</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark &amp; Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">wwschwarz@aol.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keupp, Helmut</mods:namePart>
<mods:affiliation>Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zitteliana</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zitteliana.96.81737</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:identifier>
<mods:identifier type="Pensoft-Pub">2747-8106-96-51</mods:identifier>
<mods:identifier type="ZooBank">CD8C70BA025345408EFCD430A12F6B35</mods:identifier>
<mods:identifier type="Pensoft-UUID">FCEF2A912BC258BBB022681E86486B06</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A4CD9C9FCA3B5EA994A6FF2AA47F9886" httpUri="http://treatment.plazi.org/id/A4CD9C9FCA3B5EA994A6FF2AA47F9886" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="multiple">
<paragraph pageId="0" pageNumber="51">Animalia</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
Taxon classification
<taxonomicName LSID="A4CD9C9F-CA3B-5EA9-94A6-FF2AA47F9886" authority="(Malling &amp; Gronwall, 1909)" baseAuthorityName="Malling &amp; Gronwall" baseAuthorityYear="1909" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus bornholmiensis" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="bornholmiensis">
Archaeotolithus bornholmiensis (Malling &amp;
<normalizedToken originalValue="Grønwall">Gronwall</normalizedToken>
, 1909)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">
<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling &amp; Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">Plate 4, fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="reference_group">
<paragraph pageId="0" pageNumber="51">
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus bornholmiensis" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="bornholmiensis">Otolithus bornholmiensis</taxonomicName>
1909
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus bornholmiensis" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="bornholmiensis">Otolithus bornholmiensis</taxonomicName>
- Malling &amp;
<normalizedToken originalValue="Grønwall">Gronwall</normalizedToken>
: pl. 11, figs 14-16.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName genus="Arcaeotolithus" lsidName="Arcaeotolithus bornholmiensis" pageId="0" pageNumber="51" rank="species" species="bornholmiensis">Arcaeotolithus bornholmiensis</taxonomicName>
1912
<taxonomicName genus="Arcaeotolithus" lsidName="Arcaeotolithus bornholmiensis" pageId="0" pageNumber="51" rank="species" species="bornholmiensis">Arcaeotolithus bornholmiensis</taxonomicName>
(Malling &amp;
<normalizedToken originalValue="Grønwall">Gronwall</normalizedToken>
, 1909) - Stolley: pl. 7, figs 4-5.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="genus">Otolithus</taxonomicName>
?2014 archaeotoliths group 1 - Hesse: fig. 28A-B.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="materials_examined">
<paragraph pageId="0" pageNumber="51">Material.</paragraph>
<paragraph pageId="0" pageNumber="51">
<specimenCount type="generic">1 specimen</specimenCount>
from the late Pliensbachian,
<taxonomicName genus="Hawkerense" lsidName="Hawkerense" pageId="0" pageNumber="51" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Hawkerense</emphasis>
</taxonomicName>
subzone, of Buttenheim (SNSB-BSPG
<date value="2022-04-07">2022 IV 7</date>
, Plate
<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. Otoliths of Archaeotolithus. fig. 1: Achaeotolithus bornholmiensis (Malling &amp; Gronwall, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon. figs 2 - 4: Archaeotolithus doppelsteini sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon; 2: holotype SNSB-BSPG 2022 IV 8; 3 - 4: paratypes SNSB-BSPG 2022 IV 9. figs 5 - 7: Archaeotolithus trigonalis Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12." figureDoi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51">4</figureCitation>
, fig. 1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="differential diagnosis">
<paragraph pageId="0" pageNumber="51">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName baseAuthorityName="Malling &amp; Gronwall" baseAuthorityYear="1909" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus bornholmiensis" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="bornholmiensis">
<emphasis italics="true" pageId="0" pageNumber="51">Archaeotolithus bornholmiensis</emphasis>
</taxonomicName>
is the only species in the genus with almost no indication of a sulcus and further differs from its congeners in the more strongly convex inner face. It is thus also the one most resembling the characteristics of a lapillus.
</paragraph>
<caption doi="10.3897/zitteliana.96.81737.plate4" httpUri="https://binary.pensoft.net/fig/671634" pageId="0" pageNumber="51" start="Plate 4" startId="F6">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Plate 4.</emphasis>
Otoliths of
<taxonomicName authorityName="Stolley" authorityYear="1912" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus" order="Ophidiiformes" pageId="0" pageNumber="51" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Archaeotolithus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="51">fig. 1</emphasis>
:
<taxonomicName genus="Achaeotolithus" lsidName="Achaeotolithus bornholmiensis" pageId="0" pageNumber="51" rank="species" species="bornholmiensis">
<emphasis italics="true" pageId="0" pageNumber="51">Achaeotolithus bornholmiensis</emphasis>
</taxonomicName>
(Malling &amp;
<normalizedToken originalValue="Grønwall">Gronwall</normalizedToken>
, 1909), from the late Pliensbachian of Buttenheim, holotype SNSB-BSPG 2022 IV 7, EPH horizon.
<emphasis bold="true" pageId="0" pageNumber="51">figs 2-4</emphasis>
:
<taxonomicName authorityName="Schwarzhans &amp; Keupp" authorityYear="2022" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus doppelsteini" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="doppelsteini">
<emphasis italics="true" pageId="0" pageNumber="51">Archaeotolithus doppelsteini</emphasis>
</taxonomicName>
sp. nov., from the late Pliensbachian of Buttenheim, EPH horizon;
<emphasis bold="true" pageId="0" pageNumber="51">2</emphasis>
: holotype SNSB-BSPG 2022 IV 8;
<emphasis bold="true" pageId="0" pageNumber="51">3-4</emphasis>
: paratypes SNSB-BSPG 2022 IV 9.
<emphasis bold="true" pageId="0" pageNumber="51">figs 5-7</emphasis>
:
<taxonomicName authorityName="Stolley" authorityYear="1912" genus="Archaeotolithus" kingdom="Animalia" lsidName="Archaeotolithus trigonalis" order="Ophidiiformes" pageId="0" pageNumber="51" rank="species" species="trigonalis">
<emphasis italics="true" pageId="0" pageNumber="51">Archaeotolithus trigonalis</emphasis>
</taxonomicName>
Stolley, 1912, from the late Pliensbachian of Buttenheim, EPH horizon, figured specimens SNSB-BSPG 2022 IV 12.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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