165 lines
21 KiB
XML
165 lines
21 KiB
XML
<document id="C50954B4211CC624D12980F6B0FBEA41" ID-DOI="10.5070/P940561331" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913477" ID-ZooBank="1756B24A-813B-423F-896F-91B21FF58A79" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712169731283" checkinUser="felipe" docAuthor="Hickman, Carole S." docDate="2023" docId="C23987DDFFF12936FC18FE84EFC3BE39" docLanguage="en" docName="PaleoBios.40.5.1-51.pdf" docOrigin="PaleoBios 40 (5)" docSource="http://dx.doi.org/10.5070/p940561331" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Crenella washingtonensis Weaver 1916" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="3E00FFA5FFF8293CFF9FFFA4EE04BD6F" masterDocTitle="Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs" masterLastPageNumber="51" masterPageNumber="1" pageNumber="8" updateTime="1712170163329" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="90D95CBDC57C6A74645E4EE9DD704CD6">Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs</mods:title>
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<mods:namePart id="6FBED514FDB7E09CE4D7D056DC2E5591">Hickman, Carole S.</mods:namePart>
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<mods:title id="0D61C56C1495C684E5E4B1EC52D4039B">PaleoBios</mods:title>
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<mods:part id="1D64E5400DF658E1D940AB70264D72AC">
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<mods:date id="8547A958791C612308B193FAAC658D05">2023</mods:date>
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<mods:number id="D89F96F9334AC5A77E0AEA18BAA13895">2023-06-08</mods:number>
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<mods:identifier id="97FE82F8011EAEAA8FF8B34C37A5C032" type="DOI">10.5070/P940561331</mods:identifier>
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<treatment id="C23987DDFFF12936FC18FE84EFC3BE39" ID-DOI="http://doi.org/10.5281/zenodo.10913499" ID-Zenodo-Dep="10913499" LSID="urn:lsid:plazi:treatment:C23987DDFFF12936FC18FE84EFC3BE39" httpUri="http://treatment.plazi.org/id/C23987DDFFF12936FC18FE84EFC3BE39" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<subSubSection id="028A6540FFF12935FC18FE84EBADBC54" box="[903,1449,288,316]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="4A2F36CBFFF12935FC18FE84EBADBC54" blockId="9.[832,1483,288,417]" box="[903,1449,288,316]" pageId="9" pageNumber="8">
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<heading id="116781A7FFF12935FC18FE84EBADBC54" allCaps="true" box="[903,1449,288,316]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="8" reason="2">
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<taxonomicName id="8D904D48FFF12935FC18FE84EB61BC53" authority="WEAVER" authorityName="Weaver" authorityYear="1916" box="[903,1381,288,316]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FC18FE84EAFBBC53" box="[903,1279,288,316]" italics="true" pageId="9" pageNumber="8">CRENELLA WASHINGTONENSIS</emphasis>
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WEAVER
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</taxonomicName>
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, 1916
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="028A6540FFF12935FBD7FEE0EDCDBF5E" pageId="9" pageNumber="8" type="description">
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<paragraph id="4A2F36CBFFF12935FBD7FEE0EAECBC33" blockId="9.[832,1483,288,417]" box="[1096,1256,324,348]" pageId="9" pageNumber="8">
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<figureCitation id="D2AB2A4EFFF12935FBD7FEE0EABABC34" box="[1096,1214,324,348]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (A–C, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="9" pageNumber="8">FIGS. 3D, F</figureCitation>
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;
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<figureCitation id="D2AB2A4EFFF12935FB56FEE0EAECBC33" box="[1225,1256,324,348]" captionStart="Figure 4" captionStartId="9.[112,187,1863,1887]" captionTargetBox="[115,1515,1136,1835]" captionTargetId="figure-396@9.[115,1515,1136,1835]" captionTargetPageId="9" captionText="Figure 4. A. Crenella porterensis detail of coarser ribs and rib increase by splitting (yellow arrow) and by intercalation (black arrows) (enlarged from Fig. 3A). B. Crenella washingtonensis detail of finer ribs and rib increase by intercalation (black arrows) (comparable enlargement from Fig. 3D)." figureDoi="http://doi.org/10.5281/zenodo.10913485" httpUri="https://zenodo.org/record/10913485/files/figure.png" pageId="9" pageNumber="8">4B</figureCitation>
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</paragraph>
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<paragraph id="4A2F36CBFFF12935FCDFFECEEDB9BCCE" blockId="9.[832,1483,288,417]" pageId="9" pageNumber="8">
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<taxonomicName id="8D904D48FFF12935FCDFFECEEB3CBCEB" authority="Weaver (1916)" authorityName="Weaver" authorityYear="1916" box="[832,1336,362,390]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FCDFFECEEA71BCE9" box="[832,1141,362,390]" italics="true" pageId="9" pageNumber="8">Crenella washingtonensis</emphasis>
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Weaver (1916)
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</taxonomicName>
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. p. 37; pl. 3, fig. 40.
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</paragraph>
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<paragraph id="4A2F36CBFFF12935FCDFFE16EA33BC86" blockId="9.[832,1513,434,561]" pageId="9" pageNumber="8">
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<taxonomicName id="8D904D48FFF12935FCDFFE16EBA4BCA3" authority="Weaver, Weaver (1943)" authorityName="Weaver, Weaver" authorityYear="1943" box="[832,1440,434,462]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FCDFFE16EA71BCA1" box="[832,1141,434,462]" italics="true" pageId="9" pageNumber="8">Crenella washingtonensis</emphasis>
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Weaver, Weaver (1943)
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</taxonomicName>
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. p. 116; pl. 25, fig. 4
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</paragraph>
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<paragraph id="4A2F36CBFFF12935FCDFFE5EEDCDBF5E" blockId="9.[832,1513,434,561]" pageId="9" pageNumber="8">
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<taxonomicName id="8D904D48FFF12935FCDFFE5EEB67BF7B" authority="Weaver, Tegland (1933)" authorityName="Weaver, Tegland" authorityYear="1933" box="[832,1379,504,534]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
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<emphasis id="78E4EAD9FFF12935FCDFFE5EEA30BF7B" box="[832,1076,504,534]" italics="true" pageId="9" pageNumber="8">Crenella porterensis</emphasis>
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Weaver, Tegland (1933)
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</taxonomicName>
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. p. 112; pl. 6, fig. 2.
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</paragraph>
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</subSubSection>
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<subSubSection id="028A6540FFF12935FCDFFDE6EB4BBF15" pageId="9" pageNumber="8" type="reference_group">
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<paragraph id="4A2F36CBFFF12935FCDFFDE6EB4BBF15" blockId="9.[832,1482,578,634]" pageId="9" pageNumber="8">
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<taxonomicName id="8D904D48FFF12935FCDFFDE6EBACBF33" authority="Weaver, Durham (1944)" authorityName="Weaver, Durham" authorityYear="1944" box="[832,1448,578,606]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FCDFFDE6EA71BF31" box="[832,1141,578,606]" italics="true" pageId="9" pageNumber="8">Crenella washingtonensis</emphasis>
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Weaver,
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<bibRefCitation id="2E014B3AFFF12935FB7CFDE6EBACBF33" author="Durham, J. W." box="[1251,1448,578,604]" pageId="9" pageNumber="8" pagination="101 - 212" refId="ref29671" refString="Durham, J. W. 1944. Megafaunal zones of the Oligocene of northwestern Washington. University of California Publications, Department of Geological Sciences, Bulletin 27 (5): 101 - 212. pls. 13 - 18, figs. 1 - 7." type="journal article" year="1944">Durham (1944)</bibRefCitation>
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</taxonomicName>
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. p. 140, unfigured hypotype
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<collectionCode id="2C81AE0EFFF12935FB3AFDFBEAF5BF16" box="[1189,1265,607,633]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
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35164.
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</paragraph>
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</subSubSection>
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<subSubSection id="028A6540FFF12936FCFFFD28EFC3BE39" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="discussion">
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<paragraph id="4A2F36CBFFF12936FCFFFD28EC60BCE2" blockId="9.[832,1520,652,1098]" lastBlockId="10.[112,801,195,854]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<emphasis id="78E4EAD9FFF12935FCFFFD28EDF4BFC9" bold="true" box="[864,1008,652,678]" pageId="9" pageNumber="8">Discussion</emphasis>
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—
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<taxonomicName id="8D904D48FFF12935FB92FD29EB39BFC6" authorityName="Weaver, Durham" authorityYear="1944" box="[1037,1341,653,681]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FB92FD29EB39BFC6" box="[1037,1341,653,681]" italics="true" pageId="9" pageNumber="8">Crenella washingtonensis</emphasis>
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</taxonomicName>
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was described from the Lincoln Creek Formation after description of
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<taxonomicName id="8D904D48FFF12935FCDFFD77EADDBF82" box="[832,1241,721,751]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="subSpecies" species="porterensis" subSpecies="porterensis">
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<emphasis id="78E4EAD9FFF12935FCDFFD77EADDBF82" box="[832,1241,721,751]" italics="true" pageId="9" pageNumber="8">Crenella porterensis porterensis</emphasis>
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</taxonomicName>
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and from a different
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<typeStatus id="952B8869FFF12935FCDFFD52ED7CBE7F" box="[832,888,758,784]" pageId="9" pageNumber="8">type</typeStatus>
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locality in the Lincoln Creek. It is distinguished by its smaller shell, ovate outline, and inconspicuous sculpture. It is recognized here from a single, distinctly ovate specimen from a locality in the upper part of the middle member of the Keasey Formation. The shell is a left valve with the exterior surface imbedded in matrix (
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<figureCitation id="D2AB2A4EFFF12935FCD6FC63EDA3BE8E" box="[841,935,967,993]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (A–C, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="9" pageNumber="8">Fig. 3F</figureCitation>
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). The shell is considerably thinner and more fragile than that of
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<taxonomicName id="8D904D48FFF12935FBB5FC4EEACAB96A" authorityName="Weaver, Tegland" authorityYear="1933" box="[1066,1230,1001,1030]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
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<emphasis id="78E4EAD9FFF12935FBB5FC4EEACAB96A" box="[1066,1230,1001,1030]" italics="true" pageId="9" pageNumber="8">C. porterensis</emphasis>
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</taxonomicName>
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. Portions of the opalescent interior layer remain intact, and thin exterior layer is sufficiently translucent that exceedingly fine radial sculpture is visible with a microscope under oblique lighting. The hinge plate is missing and probably was not well developed. The ovate shape may be primarily a correlate of smaller size, although the thin fragile shell and fine ornamentation serve to differentiate it from its more elongate, robust-shelled congener.
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</paragraph>
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<caption id="1EEF6643FFF12935FFEFF8E3EC22BAFB" ID-DOI="http://doi.org/10.5281/zenodo.10913485" ID-Zenodo-Dep="10913485" httpUri="https://zenodo.org/record/10913485/files/figure.png" pageId="9" pageNumber="8" startId="9.[112,187,1863,1887]" targetBox="[115,1515,1136,1835]" targetPageId="9" targetType="figure">
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<paragraph id="4A2F36CBFFF12935FFEFF8E3EC22BAFB" blockId="9.[112,1491,1863,1940]" pageId="9" pageNumber="8">
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<emphasis id="78E4EAD9FFF12935FFEFF8E3EEE8BA30" bold="true" box="[112,236,1863,1887]" pageId="9" pageNumber="8">Figure 4. A</emphasis>
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.
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<taxonomicName id="8D904D48FFF12935FF69F8ECEFCDBA0F" authorityName="Weaver, Tegland" authorityYear="1933" box="[246,457,1864,1888]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="porterensis">
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<emphasis id="78E4EAD9FFF12935FF69F8ECEFCDBA0F" box="[246,457,1864,1888]" italics="true" pageId="9" pageNumber="8">Crenella porterensis</emphasis>
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</taxonomicName>
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detail of coarser ribs and rib increase by splitting (yellow arrow) and by intercalation (black arrows) (enlarged from
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<emphasis id="78E4EAD9FFF12935FEC2F8C5EFA8BA16" bold="true" box="[349,428,1889,1913]" pageId="9" pageNumber="8">Fig. 3A</emphasis>
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).
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<emphasis id="78E4EAD9FFF12935FE5FF8C6EFD4BA15" bold="true" box="[448,464,1890,1914]" pageId="9" pageNumber="8">B</emphasis>
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.
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<taxonomicName id="8D904D48FFF12935FE44F8C7ECE3BA14" authorityName="Weaver, Durham" authorityYear="1944" box="[475,743,1891,1915]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="washingtonensis">
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<emphasis id="78E4EAD9FFF12935FE44F8C7ECE3BA14" box="[475,743,1891,1915]" italics="true" pageId="9" pageNumber="8">Crenella washingtonensis</emphasis>
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||
</taxonomicName>
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detail of finer ribs and rib increase by intercalation (black arrows) (comparable enlargement from
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<emphasis id="78E4EAD9FFF12935FE58F8D8EC1CBAFB" bold="true" box="[455,536,1916,1940]" pageId="9" pageNumber="8">Fig. 3D</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph id="4A2F36CBFFF22936FF0FFE31EEE5BF83" blockId="10.[112,801,195,854]" pageId="10" pageNumber="9">
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The stratigraphic range of this rarer species is unclear. It appears to extend into younger Oligocene strata. Close examination of Tegland’s (1933) hypotype of
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<taxonomicName id="8D904D48FFF22936FC95FE78EEFDBF76" authorityName="Weaver, Tegland" authorityYear="1933" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="10" pageNumber="9" phylum="Mollusca" rank="species" species="porterensis">
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<emphasis id="78E4EAD9FFF22936FC95FE78EEFDBF76" italics="true" pageId="10" pageNumber="9">C. porterensis</emphasis>
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</taxonomicName>
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from the Oligocene
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<typeStatus id="952B8869FFF22936FE6AFE5BEC28BF76" box="[501,556,511,537]" pageId="10" pageNumber="9">type</typeStatus>
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locality of Blakeley Formation at Restoration Point,
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<collectingRegion id="8854F829FFF22936FD9BFD86EC9DBF53" box="[516,665,546,572]" country="United States of America" name="Washington" pageId="10" pageNumber="9">Washington</collectingRegion>
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reveals an ovate shape and finer radial sculpture (
|
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<figureCitation id="D2AB2A4EFFF22936FDD6FDE1ECA0BF30" box="[585,676,581,607]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (A–C, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="10" pageNumber="9">Fig. 3D</figureCitation>
|
||
). It is here re-assigned to
|
||
<taxonomicName id="8D904D48FFF22936FEBFFDCDEC05BFEA" authorityName="Weaver, Durham" authorityYear="1944" box="[288,513,617,645]" class="Bivalvia" family="Mytilidae" genus="Crenella" kingdom="Animalia" order="Mytilida" pageId="10" pageNumber="9" phylum="Mollusca" rank="species" species="washingtonensis">
|
||
<emphasis id="78E4EAD9FFF22936FEBFFDCDEC05BFEA" box="[288,513,617,645]" italics="true" pageId="10" pageNumber="9">C. washingtonensis</emphasis>
|
||
</taxonomicName>
|
||
. In addition to finer and more numerous radial ribs, the Tegland specimen also shows numerous insertions of new ribs by intercalation (
|
||
<figureCitation id="D2AB2A4EFFF22936FFE6FD76EED7BF83" box="[121,211,722,748]" captionStart="Figure 3" captionStartId="8.[112,187,1396,1420]" captionTargetBox="[115,1515,197,1370]" captionTargetId="figure-220@8.[115,1515,195,1374]" captionTargetPageId="8" captionText="Figure 3. Crenella porterensis Weaver (1912) (A–C, E) and C. washingtonensis Weaver (1916) (D, F). A. Articulated, left valve, hypotype UCMP 32410, Lincoln Creek Formation, height 18.2 mm. B. articulated, hypotype USNM 561803, Keasey Formation, height 13.5 mm. C. Exterior mold in matrix, hypotype UCMP 110725, Keasey Formation, height 10.8 mm. D. Right valve, hypotype UCMP 32250, Blakeley Formation, height 12.5 mm. E. Partial shell exterior in matrix, hypotype UCMP 110726, Keasey Formation, height 10.3 mm. F. Partial shell exterior in matrix, hypotype 110727, Keasey Formation, height 5.4 mm." figureDoi="http://doi.org/10.5281/zenodo.10913483" httpUri="https://zenodo.org/record/10913483/files/figure.png" pageId="10" pageNumber="9">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4A2F36CBFFF22936FF0FFD51EFC3BE39" blockId="10.[112,801,195,854]" pageId="10" pageNumber="9">
|
||
<emphasis id="78E4EAD9FFF22936FF0FFD51EFE3BE60" bold="true" box="[144,487,757,783]" pageId="10" pageNumber="9">Figured Keasey hypotype</emphasis>
|
||
—
|
||
<collectionCode id="2C81AE0EFFF22936FD9DFD51EC54BE60" box="[514,592,757,783]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="10" pageNumber="9" type="University or college">UCMP</collectionCode>
|
||
110727 (in matrix), approximate length 7.4 mm, approximate height 5.4 mm, Loc.
|
||
<collectionCode id="2C81AE0EFFF22936FE8CFC98EF5BBE39" box="[275,351,828,854]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="10" pageNumber="9" type="University or college">UCMP</collectionCode>
|
||
IP7984.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |