163 lines
21 KiB
XML
163 lines
21 KiB
XML
<document id="83DB8DE7A5E3A37E68410DF817DCF18F" ID="10.11646/zootaxa.4205.2.3" ID-DOI="10.11646/zootaxa.4205.2.3" ID-GBIF-Dataset="4d80dadb-048e-402d-97eb-4261b8a7bd20" ID-ISSN="1175-5326" ID-Zenodo-Dep="192909" ID-ZooBank="87489FA9-6C1C-4830-9D5D-51383B4221AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481009146824" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2016" docId="711EE309A607FFA7419AFEA41822BEBE" docLanguage="en" docName="zootaxa.4205.2.3.pdf" docOrigin="Zootaxa 4205 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lesteva steeli Lohse 1982" docType="treatment" docVersion="10" lastPageNumber="147" masterDocId="8D279B71A603FFA3410DFFEF1D74B90C" masterDocTitle="New taxonomic and faunistic data on the Himalayan Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="150" masterPageNumber="143" pageNumber="147" updateTime="1698395911389" updateUser="plazi">
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<mods:title id="8EF837600F3FAD6F751B24DF224549EE">New taxonomic and faunistic data on the Himalayan Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="84FC34B0EBA14842E8475FC0CE7C6847">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="E0F0E1F678DD349F1E29E50AEAFA71E2">Zootaxa</mods:title>
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<mods:date id="A90A8EEB4CC16A99244BC8A9759E9F5E">2016</mods:date>
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<mods:number id="6D8CBCCB18B7EAE2C7469BA889F00BB8">4205</mods:number>
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<treatment id="711EE309A607FFA7419AFEA41822BEBE" ID-DOI="http://doi.org/10.5281/zenodo.6058328" ID-GBIF-Taxon="124955204" ID-Zenodo-Dep="6058328" LSID="urn:lsid:plazi:treatment:711EE309A607FFA7419AFEA41822BEBE" httpUri="http://treatment.plazi.org/id/711EE309A607FFA7419AFEA41822BEBE" lastPageNumber="147" pageId="4" pageNumber="147">
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<subSubSection id="B1AD0194A607FFA7419AFEA41C66B88B" pageId="4" pageNumber="147" type="nomenclature">
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<paragraph id="F908521FA607FFA7419AFEA41F10B86A" blockId="4.[151,612,331,391]" box="[151,612,331,358]" pageId="4" pageNumber="147">
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<heading id="A240E573A607FFA7419AFEA41F10B86A" bold="true" box="[151,612,331,358]" fontSize="11" level="1" pageId="4" pageNumber="147" reason="1">
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<treatmentCitationGroup id="D9A77531A607FFA7419AFEA41F10B86A" box="[151,612,331,358]" pageId="4" pageNumber="147">
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<taxonomicName id="3EB7299CA607FFA7419AFEA41F10B86A" ID-CoL="6PQSF" authority="Lohse, 1982: 203" authorityName="Lohse" authorityPageNumber="203" authorityYear="1982" box="[151,612,331,358]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA7419AFEA41F10B86A" bold="true" box="[151,612,331,358]" pageId="4" pageNumber="147">
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<emphasis id="CBC38E0DA607FFA7419AFEA41D86B869" bold="true" box="[151,242,331,357]" italics="true" pageId="4" pageNumber="147">Lesteva</emphasis>
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(s.str.)
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<emphasis id="CBC38E0DA607FFA74040FEA41CFFB869" bold="true" box="[333,395,331,357]" italics="true" pageId="4" pageNumber="147">steeli</emphasis>
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<treatmentCitation id="7816740EA607FFA7409EFEA31F10B86A" author="Lohse" box="[403,612,332,358]" page="203" pageId="4" pageNumber="147" year="1982">
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<bibRefCitation id="9D262FEEA607FFA7409EFEA31F5FB86A" author="Lohse" box="[403,555,332,358]" pageId="4" pageNumber="147" refString="Lohse, G. A. (1982) The species of the genus Lesteva Latreille (Col., Staphylinidae) known from northern India. The Entomologist's Monthly Magazine, 118, 201 - 205." type="journal article" year="1982">Lohse, 1982</bibRefCitation>
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: 203
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</treatmentCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="F908521FA607FFA7419AFE811C66B88B" blockId="4.[151,612,331,391]" box="[151,274,366,391]" pageId="4" pageNumber="147">
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(
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<figureCitation id="618C4E9AA607FFA74192FE811C7CB88B" box="[159,264,366,391]" captionStart="FIGURES 1 – 2" captionStartId="5.[151,264,1332,1355]" captionTargetBox="[283,1353,203,1288]" captionTargetId="figure@5.[180,1407,195,1311]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Aedeagus of L. (s. str.) steeli: 1 — parameral view, 2 — lateral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/192910/files/figure.png" pageId="4" pageNumber="147">Figs. 1–2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B1AD0194A607FFA7419AFE571CA2B8C0" box="[151,470,438,461]" pageId="4" pageNumber="147" type="reference_group">
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<paragraph id="F908521FA607FFA7419AFE571CA2B8C0" blockId="4.[151,470,438,461]" box="[151,470,438,461]" pageId="4" pageNumber="147">
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<treatmentCitationGroup id="D9A77531A607FFA7419AFE571CA2B8C0" box="[151,470,438,461]" pageId="4" pageNumber="147">
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<taxonomicName id="3EB7299CA607FFA7419AFE571CA2B8C0" ID-CoL="6PQSF" authority="Lohse, 1982: 203" authorityName="Lohse" authorityPageNumber="203" authorityYear="1982" box="[151,470,438,461]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA7419AFE571C6BB8C0" box="[151,287,438,461]" italics="true" pageId="4" pageNumber="147">Lesteva steeli</emphasis>
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<treatmentCitation id="7816740EA607FFA7402BFE591CA2B8C0" author="Lohse" box="[294,470,438,460]" page="203" pageId="4" pageNumber="147" year="1982">
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<bibRefCitation id="9D262FEEA607FFA7402BFE591CD2B8C0" author="Lohse" box="[294,422,438,460]" pageId="4" pageNumber="147" refString="Lohse, G. A. (1982) The species of the genus Lesteva Latreille (Col., Staphylinidae) known from northern India. The Entomologist's Monthly Magazine, 118, 201 - 205." type="journal article" year="1982">Lohse, 1982</bibRefCitation>
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: 203
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1AD0194A607FFA7419AFE161C71BA02" pageId="4" pageNumber="147" type="materials_examined">
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<paragraph id="F908521FA607FFA7419AFE161C71BA02" blockId="4.[151,1437,505,1970]" pageId="4" pageNumber="147">
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<emphasis id="CBC38E0DA607FFA7419AFE161CB2BB1E" bold="true" box="[151,454,505,530]" pageId="4" pageNumber="147">Type material examined:</emphasis>
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Holotype of
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<taxonomicName id="3EB7299CA607FFA7436EFE151EE2BB1D" authority="Lohse, 1982" authorityName="Lohse" authorityYear="1982" box="[611,918,505,530]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA7436EFE151F89BB1D" box="[611,765,505,529]" italics="true" pageId="4" pageNumber="147">Lesteva steeli</emphasis>
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<bibRefCitation id="9D262FEEA607FFA7420BFE161EE2BB1D" author="Lohse" box="[774,918,505,530]" pageId="4" pageNumber="147" refString="Lohse, G. A. (1982) The species of the genus Lesteva Latreille (Col., Staphylinidae) known from northern India. The Entomologist's Monthly Magazine, 118, 201 - 205." type="journal article" year="1982">Lohse, 1982</bibRefCitation>
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</taxonomicName>
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, ♂ [The specimen is damaged: without right antennomeres 7–11, and left hind leg; the specimen was reglued on a new card, old card pinned under the new card; abdominal sternite VIII was glued under the beetle; the preparation of the aedeagus is in a microvial and was pinned under all labels]: ‘Holo-| type’ <round label with red margin, printed>, ‘Kashmir | Gulmarg’ <rectangular label, handwritten in black>, M.Cameron. | Bequest.| B.M. 1955-147.’ <rectangular printed label>, ‘HOLOTYPUS’ <red rectangular label, handwritten in black>, ‘
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<taxonomicName id="3EB7299CA607FFA74275FD411EBABBC9" box="[888,974,686,709]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="genus">
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<emphasis id="CBC38E0DA607FFA74275FD411EBABBC9" box="[888,974,686,709]" italics="true" pageId="4" pageNumber="147">Lesteva</emphasis>
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</taxonomicName>
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<taxonomicName id="3EB7299CA607FFA742E8FD421954BBC9" box="[997,1056,685,709]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
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<emphasis id="CBC38E0DA607FFA742E8FD421954BBC9" box="[997,1056,685,709]" italics="true" pageId="4" pageNumber="147">steeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D0F03376A607FFA74524FD41192ABBC9" box="[1065,1118,686,709]" pageId="4" pageNumber="147" rank="species">n.sp.</taxonomicNameLabel>
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| Holotypus [handwritten] | det.Dr.G.A.Lohse’ <rectangular printed label>, ‘
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<taxonomicName id="3EB7299CA607FFA743C5FD3D1E69BBE5" box="[712,797,722,745]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="genus">
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<emphasis id="CBC38E0DA607FFA743C5FD3D1E69BBE5" box="[712,797,722,745]" italics="true" pageId="4" pageNumber="147">Lesteva</emphasis>
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</taxonomicName>
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(s.str.) |
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<taxonomicName id="3EB7299CA607FFA74283FD3E1911BBE6" authority="Lohse, 1982" authorityName="Lohse" authorityYear="1982" box="[910,1125,721,746]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA74283FD3E1EBDBBE5" box="[910,969,721,745]" italics="true" pageId="4" pageNumber="147">steeli</emphasis>
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<bibRefCitation id="9D262FEEA607FFA742D8FD3E1911BBE6" author="Lohse" box="[981,1125,721,746]" pageId="4" pageNumber="147" refString="Lohse, G. A. (1982) The species of the genus Lesteva Latreille (Col., Staphylinidae) known from northern India. The Entomologist's Monthly Magazine, 118, 201 - 205." type="journal article" year="1982">Lohse, 1982</bibRefCitation>
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</taxonomicName>
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| Shavrin A.V. det. 2016’ (BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="B1AD0194A607FFA741CAFCF618F0BE82" pageId="4" pageNumber="147" type="description">
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<paragraph id="F908521FA607FFA741CAFCF6190ABA5A" blockId="4.[151,1437,505,1970]" pageId="4" pageNumber="147">
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<emphasis id="CBC38E0DA607FFA741CAFCF61C00BA3E" bold="true" box="[199,372,793,818]" pageId="4" pageNumber="147">Redescription.</emphasis>
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Measurements of the holotype: HW: 0.70; HL: 0.50; OL: 0.25; LT: 0.10; AL: 1.62; PL: 0.65; PWMax: 0.82; PWMin: 0.60; ElL: 1.42; EW: 1.37; AW(IV): 1.25; AedL: 0.79; TL: 3.95.
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</paragraph>
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<paragraph id="F908521FA607FFA741CAFC8E1E79BCB5" blockId="4.[151,1437,505,1970]" pageId="4" pageNumber="147">Body brownish black, glossy; legs and antennomeres reddish brown; tarsi, mouthparts and bases of antennomeres 1–3 yellow. Punctation of head deep, distinctly denser on infraorbital ridges; punctation of pronotum regular, slightly deeper than that on head, interspaces between punctures as diameter of 2–3 nearest punctures; scutellum with small irregular punctures; elytra with somewhat larger punctation than that on pronotum; abdomen with small regular punctation. Forebody without microsculpture; abdomen with reticulate microsculpture on abdominal tergite IV and without visible microsculpture on other tergites. Head 1.4 times wider than long, with convex frons and moderately prominent infraorbital ridges, with rather deep depressions between them. Eyes large and convex, two and half times longer than temples. Ocelli poorly visible (same coloration as that of head), distance between ocelli about same length of distance between ocellus and posterior margin of eye. Antennae reaching one-third of elytral length when reclined; measurements of antennomeres (length/width): 1: 0.21 × 0.08; 2: 0.14 × 0.05; 3: 0.15 × 0.06; 4¯6: 0.13 × 0.06; 7¯8: 0.14 × 0.06; 9¯10: 0.14 × 0.07; 11: 0.17 × 0.07. Pronotum cordate, convex, 1.2 times as wide as long, wider in apical third; disc of pronotum without longitudinal impressions; mediobasal portion of pronotum with very weak small transverse depression. Elytra moderately convex, slightly more than twice longer than pronotum and slightly wider than length, insignificantly expanded posteriorly, with rounded hind margins. Metatarsus slightly more than twice longer than metatibia; tarsomere 1 of metatarsus shorter than tarsomeres 2–3, approximately as long as tarsomere 5. Abdomen slightly narrower than elytra; abdominal tergite V with paired tomentose spots.</paragraph>
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<paragraph id="F908521FA607FFA741CAFA2A1CD1BF46" blockId="4.[151,1437,505,1970]" pageId="4" pageNumber="147">
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Male. Abdominal sternite VIII with widely emarginated posterior margin. Aedeagus (
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<figureCitation id="618C4E9AA607FFA745B9FA2A198DBCD2" box="[1204,1273,1477,1502]" captionStart="FIGURES 1 – 2" captionStartId="5.[151,264,1332,1355]" captionTargetBox="[283,1353,203,1288]" captionTargetId="figure@5.[180,1407,195,1311]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Aedeagus of L. (s. str.) steeli: 1 — parameral view, 2 — lateral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/192910/files/figure.png" pageId="4" pageNumber="147">Fig. 1</figureCitation>
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) with narrow median lobe and small rounded apex; parameres with wide apical portions, exceeding apex of aedeagus; internal sac moderately long, with strongly sclerotized areas of thorns in basal and median part of aedeagus. Aedeagus in lateral view as in
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<figureCitation id="618C4E9AA607FFA74051F9DE1CD5BF46" box="[348,417,1585,1610]" captionStart="FIGURES 1 – 2" captionStartId="5.[151,264,1332,1355]" captionTargetBox="[283,1353,203,1288]" captionTargetId="figure@5.[180,1407,195,1311]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Aedeagus of L. (s. str.) steeli: 1 — parameral view, 2 — lateral view. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/192910/files/figure.png" pageId="4" pageNumber="147">Fig. 2</figureCitation>
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.
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</paragraph>
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<paragraph id="F908521FA607FFA741CAF9BA1CE4BF62" blockId="4.[151,1437,505,1970]" box="[199,400,1621,1646]" pageId="4" pageNumber="147">Female unknown.</paragraph>
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<paragraph id="F908521FA607FFA741CAF99618F0BE82" blockId="4.[151,1437,505,1970]" pageId="4" pageNumber="147">
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<emphasis id="CBC38E0DA607FFA741CAF9961CC4BF9E" bold="true" box="[199,432,1657,1682]" pageId="4" pageNumber="147">Comparative notes.</emphasis>
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Based on the body size and coloration, the punctation, microsculpture and pubescence of the body, the proportions of metatibia and metatarsi, as well as the general structure of the aedeagus,
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<taxonomicName id="3EB7299CA607FFA745FAF97118E8BFB9" box="[1271,1436,1693,1718]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA745FAF9711872BFB9" box="[1271,1286,1694,1717]" italics="true" pageId="4" pageNumber="147">L</emphasis>
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. (s.str.)
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<emphasis id="CBC38E0DA607FFA7446CF97218E8BFB9" box="[1377,1436,1693,1717]" italics="true" pageId="4" pageNumber="147">steeli</emphasis>
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</taxonomicName>
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is most similar to the Nepalese
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<emphasis id="CBC38E0DA607FFA740FEF92D1F76BFD5" box="[499,514,1730,1753]" italics="true" pageId="4" pageNumber="147">L</emphasis>
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. (s.str.)
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<emphasis id="CBC38E0DA607FFA74350F92D1FD8BFD5" box="[605,684,1730,1753]" italics="true" pageId="4" pageNumber="147">jaegeri</emphasis>
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<bibRefCitation id="9D262FEEA607FFA743BEF92E1E26BFD6" author="Shavrin" box="[691,850,1729,1754]" pageId="4" pageNumber="147" refString="Shavrin, A. V. (2012) New and poorly known species of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae) from Nepal. Zootaxa, 3547, 78 - 84." type="journal article" year="2012">Shavrin, 2012</bibRefCitation>
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, from which it differs by the narrower antennomeres and apical part of the aedeagus, by the narrower and longer elytra, by the wider parameres and by details of the structure of the internal sac. The shape of the median lobe of the aedeagus of
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<taxonomicName id="3EB7299CA607FFA7451FF8E519C7BE2D" box="[1042,1203,1801,1826]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis id="CBC38E0DA607FFA7451FF8E51955BE2D" box="[1042,1057,1802,1825]" italics="true" pageId="4" pageNumber="147">L</emphasis>
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(s.str.)
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<emphasis id="CBC38E0DA607FFA74575F8E619C7BE2D" box="[1144,1203,1801,1825]" italics="true" pageId="4" pageNumber="147">steeli</emphasis>
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</taxonomicName>
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is similar on that of
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<collectingRegion id="3B739CFDA607FFA7419AF8C21D9EBE4A" box="[151,234,1837,1862]" country="Sri Lanka" name="Central" pageId="4" pageNumber="147">Central</collectingRegion>
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Asian
|
||
<emphasis id="CBC38E0DA607FFA7403AF8C11C32BE49" box="[311,326,1838,1861]" italics="true" pageId="4" pageNumber="147">L</emphasis>
|
||
. (s.str.)
|
||
<emphasis id="CBC38E0DA607FFA740ACF8C21F67BE49" box="[417,531,1837,1861]" italics="true" pageId="4" pageNumber="147">bucharica</emphasis>
|
||
Fauvel, 1900 (see Figs. 30¯
|
||
<quantity id="3E4FFFFAA607FFA74246F8C21EF2BE4A" box="[843,902,1837,1862]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="4" pageNumber="147" unit="in" value="35.0">35 in</quantity>
|
||
<bibRefCitation id="9D262FEEA607FFA74281F8C21945BE4A" author="Shavrin" box="[908,1073,1837,1862]" pageId="4" pageNumber="147" refString="Shavrin, A. V. (2015 b) Review of Lesteva Latreille, 1797 of Central Asia (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 3974 (1), 29 - 48." type="journal article" year="2015" yearSuffix="b">Shavrin 2015b</bibRefCitation>
|
||
) but differs by the shape of apex of the parameres and structure of the internal sac, as well as by the completely different external characters (
|
||
<emphasis id="CBC38E0DA607FFA74488F8BD18E0BE65" box="[1413,1428,1874,1897]" italics="true" pageId="4" pageNumber="147">L</emphasis>
|
||
. (s.str.)
|
||
<emphasis id="CBC38E0DA607FFA741E9F89A1C22BE81" box="[228,342,1909,1933]" italics="true" pageId="4" pageNumber="147">bucharica</emphasis>
|
||
is significantly larger, coloration paler, antennae very long, reaching half-length of elytra, etc.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1AD0194A607FFA741CAF8761822BEBE" box="[199,1366,1945,1970]" pageId="4" pageNumber="147" type="discussion">
|
||
<paragraph id="F908521FA607FFA741CAF8761822BEBE" blockId="4.[151,1437,505,1970]" box="[199,1366,1945,1970]" pageId="4" pageNumber="147">
|
||
<emphasis id="CBC38E0DA607FFA741CAF8761C4FBEBE" bold="true" box="[199,315,1945,1970]" pageId="4" pageNumber="147">Remarks.</emphasis>
|
||
The species, known only on the type material, was described by
|
||
<bibRefCitation id="9D262FEEA607FFA74519F87619DEBEBE" author="Lohse" box="[1044,1194,1945,1970]" pageId="4" pageNumber="147" refString="Lohse, G. A. (1982) The species of the genus Lesteva Latreille (Col., Staphylinidae) known from northern India. The Entomologist's Monthly Magazine, 118, 201 - 205." type="journal article" year="1982">Lohse (1982)</bibRefCitation>
|
||
from Kashmir.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |