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<document id="C6532DA44A30C3E7728A373BBC0904C2" ID-DOI="10.5281/zenodo.5355939" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355939" ID-ZooBank="566024F1-56F3-41D0-B122-45255A075E61" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630216184017" checkinUser="valdenar" docAuthor="Tumpeesuwan, Chanidaporn &amp; Tumpeesuwan, Sakboworn" docDate="2017" docId="03DD8D18CB104159C0C23255FD7AF161" docLanguage="en" docName="65rbz181-188.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Aenigmatoconcha Tumpeesuwan &amp; Tumpeesuwan 2017, new genus" docType="treatment" docUuid="CF52946C-F14C-4CEB-AF7A-D419E1EFAA13" docUuidSource="ZooBank" docVersion="3" lastPageNumber="184" masterDocId="FFE4F560CB11415AC3063465FF8BF334" masterDocTitle="Discovery of an overlooked Helicarionid land snail (Helicarionidae: Durgellinae) from northeastern Thailand, with description of a new genus and new species, and note on radula morphology and genital system" masterLastPageNumber="188" masterPageNumber="181" pageNumber="182" updateTime="1699189763151" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CEF85F362F2477B5B219540067B51BE0">Discovery of an overlooked Helicarionid land snail (Helicarionidae: Durgellinae) from northeastern Thailand, with description of a new genus and new species, and note on radula morphology and genital system</mods:title>
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<mods:date id="A198144A313CFF734BFD685EA99BAA7F">2017</mods:date>
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<mods:number id="14870CA177E611FD18D9C8D1BF4B4A1A">2017-06-07</mods:number>
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<subSubSection id="C36E6F85CB10415BC0C23255FA98F57D" box="[964,1299,1584,1609]" pageId="1" pageNumber="182" type="nomenclature">
<paragraph id="8BCB3C0ECB10415BC0C23255FA98F57D" blockId="1.[964,1299,1584,1609]" box="[964,1299,1584,1609]" pageId="1" pageNumber="182">
<heading id="D0838B62CB10415BC0C23255FA98F57D" bold="true" box="[964,1299,1584,1609]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="182" reason="2">
<emphasis id="B900E01CCB10415BC0C23255FA98F57D" bold="true" box="[964,1299,1584,1609]" pageId="1" pageNumber="182">
<emphasis id="B900E01CCB10415BC0C23255FB1FF57C" bold="true" box="[964,1172,1584,1608]" italics="true" pageId="1" pageNumber="182">
<taxonomicName id="4C74478DCB10415BC0C23255FB06F57C" ID-CoL="7NN6M" authority="Tumpeesuwan &amp; Tumpeesuwan, 2017" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[964,1165,1584,1608]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="182" phylum="Mollusca" rank="genus" status="gen. nov.">Aenigmatoconcha</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2335D67CB10415BC79A3254FA98F57D" box="[1180,1299,1585,1609]" pageId="1" pageNumber="182" rank="genus">new genus</taxonomicNameLabel>
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<subSubSection id="C36E6F85CB10415BC02B3214FA00F5BD" box="[813,1419,1649,1673]" pageId="1" pageNumber="182" type="type_taxon">
<paragraph id="8BCB3C0ECB10415BC02B3214FA00F5BD" blockId="1.[813,1419,1649,1673]" box="[813,1419,1649,1673]" pageId="1" pageNumber="182">
<emphasis id="B900E01CCB10415BC02B3214FC4EF5BD" bold="true" box="[813,965,1649,1673]" pageId="1" pageNumber="182">
<typeStatus id="54CF82ACCB10415BC02B3214FCECF5BD" box="[813,871,1649,1673]" pageId="1" pageNumber="182">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C74478DCB10415BC0C93214FB79F5BD" ID-CoL="64Y97" authority="Tumpeesuwan &amp; Tumpeesuwan, 2017" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[975,1266,1649,1673]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="182" phylum="Mollusca" rank="species" species="clivicola" status="sp. nov.">
<emphasis id="B900E01CCB10415BC0C93214FB79F5BD" box="[975,1266,1649,1673]" italics="true" pageId="1" pageNumber="182">Aenigmatoconcha clivicola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2335D67CB10415BC6073214FA0DF5BD" box="[1281,1414,1649,1673]" pageId="1" pageNumber="182" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
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<subSubSection id="C36E6F85CB10415BC02B32D4FB8EF41D" pageId="1" pageNumber="182" type="diagnosis">
<paragraph id="8BCB3C0ECB10415BC02B32D4FB8EF41D" blockId="1.[813,1452,1713,1833]" pageId="1" pageNumber="182">
<emphasis id="B900E01CCB10415BC02B32D4FC2AF5FD" bold="true" box="[813,929,1713,1737]" pageId="1" pageNumber="182">Diagnosis.</emphasis>
The radula possesses similar unicuspid spatulate teeth arranged in V-shaped rows. The genitalia lack an amatorial organ and possess a large, stout penis and slender elongate epiphallus.
</paragraph>
</subSubSection>
<subSubSection id="C36E6F85CB10415BC02B3334FB44F49D" pageId="1" pageNumber="182" type="etymology">
<paragraph id="8BCB3C0ECB10415BC02B3334FB44F49D" blockId="1.[813,1452,1873,1961]" pageId="1" pageNumber="182">
<emphasis id="B900E01CCB10415BC02B3334FC3CF45D" bold="true" box="[813,951,1873,1897]" pageId="1" pageNumber="182">Etymology.</emphasis>
“
<taxonomicName id="4C74478DCB10415BC0C93334FBB8F45C" box="[975,1075,1873,1896]" class="Insecta" family="Carabidae" genus="Aenigma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B900E01CCB10415BC0C93334FBB8F45C" box="[975,1075,1873,1896]" italics="true" pageId="1" pageNumber="182">Aenigma</emphasis>
</taxonomicName>
” from the Latin word meaning something obscure, a riddle or mystery, and “
<emphasis id="B900E01CCB10415BC6113314FAEFF4BD" box="[1303,1380,1905,1929]" italics="true" pageId="1" pageNumber="182">concha</emphasis>
” from the Latin word meaning snail or shell.
</paragraph>
</subSubSection>
<subSubSection id="C36E6F85CB104159C02B33B4FD7AF161" lastPageId="3" lastPageNumber="184" pageId="1" pageNumber="182" type="discussion">
<paragraph id="8BCB3C0ECB10415BC02B33B4FA20FB1D" blockId="1.[813,1452,2001,2089]" pageId="1" pageNumber="182">
<emphasis id="B900E01CCB10415BC02B33B4FC10F4DD" bold="true" box="[813,923,2001,2025]" pageId="1" pageNumber="182">Remarks.</emphasis>
The radula and genital organs form the unique characters distinguishing
<taxonomicName id="4C74478DCB10415BC7573394FA92FB3D" authority="Tumpeesuwan &amp; Tumpeesuwan, 2017" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[1105,1305,2033,2057]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="182" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B900E01CCB10415BC7573394FA92FB3D" box="[1105,1305,2033,2057]" italics="true" pageId="1" pageNumber="182">Aenigmatoconcha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2335D67CB10415BC62D3394FA2CFB3D" box="[1323,1447,2033,2057]" pageId="1" pageNumber="182" rank="genus">new genus</taxonomicNameLabel>
, from other morphologically close helicarionid genera.
</paragraph>
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<paragraph id="8BCB3C0ECB134158C38E34D8FC2EF3E6" blockId="2.[136,933,189,211]" box="[136,933,189,211]" pageId="2" pageNumber="183">
Table 1. Classification of
<taxonomicName id="4C74478DCB134158C28034D8FDEFF3E7" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" box="[390,612,189,211]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C28034D8FE5EF3E6" box="[390,469,189,210]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
<bibRefCitation id="EFE541FFCB134158C2DB34DBFDEFF3E7" author="Benson WH" box="[477,612,189,211]" pageId="2" pageNumber="183" pagination="26 - 27" refId="ref3866" refString="Benson WH (1860) On the Burmese genus Sophina. The Annals and Magazine of Natural History, Series 3, Volume 5: 26 - 27." type="journal article" year="1860">Benson, 1860</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C74478DCB134158C19A34D8FC2AF3E6" authority="Ancey, 1887" authorityName="Ancey" authorityYear="1887" box="[668,929,189,210]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C19A34D8FC90F3E6" box="[668,795,189,210]" italics="true" pageId="2" pageNumber="183">Chalepotaxis</emphasis>
<bibRefCitation id="EFE541FFCB134158C02734D8FC2AF3E6" author="Ancey CF" box="[801,929,189,210]" pageId="2" pageNumber="183" pagination="38 - 39" refId="ref3649" refString="Ancey CF (1887) Description of new genera or subgenera of Helicidae. The Conchologists' Exchange, 1, 53 - 54, 64, 75 - 76; 2, 22 - 23, 38 - 39." type="journal article" year="1887">Ancey, 1887</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BCB3C0ECB134158C0F63497FA1FF2FF" pageId="2" pageNumber="183">
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<emphasis id="B900E01CCB134158C0F63497FBAFF233" bold="true" box="[1008,1060,242,263]" pageId="2" pageNumber="183">Taxa</emphasis>
</th>
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<emphasis id="B900E01CCB134158C391356CFF60F22A" bold="true" box="[151,235,265,286]" pageId="2" pageNumber="183">Authors</emphasis>
</th>
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<td id="76955730CB13BEA5C17F3545FC8DF202" box="[633,774,288,310]" gridcol="1" gridrow="2" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C19A3545FD6FF202" bold="true" box="[668,740,288,310]" pageId="2" pageNumber="183">Family</emphasis>
</td>
<td id="76955730CB13BEA5C0513545FBAFF202" box="[855,1060,288,310]" gridcol="2" gridrow="2" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C0613545FC5BF202" bold="true" box="[871,976,288,310]" pageId="2" pageNumber="183">Subfamily</emphasis>
</td>
<td id="76955730CB13BEA5C7413545FB21F202" box="[1095,1194,288,310]" gridcol="3" gridrow="2" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C75D3545FB1FF202" bold="true" box="[1115,1172,288,310]" pageId="2" pageNumber="183">Tribe</emphasis>
</td>
<td id="76955730CB13BEA5C6133545FA1FF202" box="[1301,1428,288,310]" gridcol="4" gridrow="2" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C6353545FAFFF202" bold="true" box="[1331,1396,288,310]" pageId="2" pageNumber="183">Genus</emphasis>
</td>
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<th id="76955730CB13BEA5C3913532FE7BF259" box="[151,496,343,365]" gridcol="0" gridrow="3" pageId="2" pageNumber="183">
<bibRefCitation id="EFE541FFCB134158C3913532FE7BF259" author="Blanford WT &amp; Godwin-Austen HH" box="[151,496,343,365]" pageId="2" pageNumber="183" refId="ref3895" refString="Blanford WT &amp; Godwin-Austen HH (1908). The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, 311 pp." type="book" year="1908">Blanford &amp; Godwin-Austen (1908)</bibRefCitation>
</th>
<td id="76955730CB13BEA5C17F3532FC8DF259" box="[633,774,343,365]" gridcol="1" gridrow="3" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C1943532FD65F259" authorityName="Morch" authorityYear="1864" box="[658,750,343,365]" class="Gastropoda" family="Zonitidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="family">Zonitidae</taxonomicName>
</td>
<td id="76955730CB13BEA5C0513532FBAFF259" box="[855,1060,343,365]" gridcol="2" gridrow="3" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C0603532FC59F259" box="[870,978,343,365]" pageId="2" pageNumber="181" rank="subFamily" subFamily="Sophininae">Sophininae</taxonomicName>
</td>
<td id="76955730CB13BEA5C7413532FB21F259" box="[1095,1194,343,365]" gridcol="3" gridrow="3" pageId="2" pageNumber="183">-</td>
<td id="76955730CB13BEA5C6133532FA1FF259" box="[1301,1428,343,365]" gridcol="4" gridrow="3" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C62B3532FAF7F258" authorityName="Benson" authorityYear="1860" box="[1325,1404,343,364]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C62B3532FAF7F258" box="[1325,1404,343,364]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="35443E4CCB13BEA5C3913512FA1FF2B9" box="[151,1428,375,397]" gridrow="4" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913512FE7BF2B9" box="[151,496,375,397]" gridcol="0" gridrow="4" pageId="2" pageNumber="183">
<bibRefCitation id="EFE541FFCB134158C3913512FEB4F2B9" author="Schileyko AA" box="[151,319,375,397]" pageId="2" pageNumber="183" pagination="1309 - 1466" refId="ref4232" refString="Schileyko AA (2003) Treatise on recent terrestrial pulmonate molluscs. Part 10. Ariophantidae, Ostracolethidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, supplement 2, 1309 - 1466." type="journal article" year="2003">Schileyko (2003)</bibRefCitation>
</th>
<td id="76955730CB13BEA5C17F3512FC8DF2B9" box="[633,774,375,397]" gridcol="1" gridrow="4" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C17D3512FC8FF2B9" authorityName="Godwin-Austen" authorityYear="1883" box="[635,772,375,397]" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
</td>
<td id="76955730CB13BEA5C0513512FBAFF2B9" box="[855,1060,375,397]" gridcol="2" gridrow="4" pageId="2" pageNumber="183">Ariophantinae</td>
<td id="76955730CB13BEA5C7413512FB21F2B9" box="[1095,1194,375,397]" gridcol="3" gridrow="4" pageId="2" pageNumber="183">Satiellini</td>
<td id="76955730CB13BEA5C6133512FA1FF2B9" box="[1301,1428,375,397]" gridcol="4" gridrow="4" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C62B3512FAF7F2B8" authorityName="Benson" authorityYear="1860" box="[1325,1404,375,396]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C62B3512FAF7F2B8" box="[1325,1404,375,396]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
</taxonomicName>
</td>
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<bibRefCitation id="EFE541FFCB134158C39135F3FE19F298" author="Bouchet P &amp; Rocroi JP" box="[151,402,406,428]" pageId="2" pageNumber="183" pagination="1 - 397" refId="ref3689" refString="Bouchet P &amp; Rocroi JP (2005) Classification and Nomenclature of Gastropod Families. Malacologia, 47 (1 - 2): 1 - 397." type="journal article" year="2005">Bouchet &amp; Rocroi (2005)</bibRefCitation>
</th>
<td id="76955730CB13BEA5C17F35F3FC8DF298" box="[633,774,406,428]" gridcol="1" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C17F35F3FC8DF298" authorityName="Bourguignat" authorityYear="1877" box="[633,774,406,428]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
</td>
<td id="76955730CB13BEA5C05135F3FBAFF298" box="[855,1060,406,428]" gridcol="2" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C06535F3FC5FF298" authorityName="Godwin-Austen" authorityYear="1888" box="[867,980,406,428]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="182" phylum="Mollusca" rank="subFamily" subFamily="Durgellinae">Durgellinae</taxonomicName>
</td>
<td id="76955730CB13BEA5C74135F3FB21F298" box="[1095,1194,406,428]" gridcol="3" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C74135F3FB21F298" authorityName="Godwin-Austen" authorityYear="1888" box="[1095,1194,406,428]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="182" phylum="Mollusca" rank="tribe" tribe="Durgellini">Durgellini</taxonomicName>
</td>
<td id="76955730CB13BEA5C61335F3FA1FF298" box="[1301,1428,406,428]" gridcol="4" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C62B35F3FAF7F29F" authorityName="Benson" authorityYear="1860" box="[1325,1404,406,427]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C62B35F3FAF7F29F" box="[1325,1404,406,427]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="35443E4CCB13BEA5C39135D3FA1FF2F8" box="[151,1428,438,460]" gridrow="6" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39135D3FE7BF2F8" box="[151,496,438,460]" gridcol="0" gridrow="6" pageId="2" pageNumber="183">
<bibRefCitation id="EFE541FFCB134158C39135D3FE19F2F8" author="Pall-Gergely B &amp; Feher Z &amp; Othani JU &amp; Asami T" box="[151,402,438,460]" pageId="2" pageNumber="183" refId="ref4159" refString="Pall-Gergely B, Feher Z, Othani JU &amp; Asami T (2016) An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylomatophora: Helicarionidae). Molluscan Research, DOI: 10.1080 / 13235818.2016.1234996." type="book" year="2016">Páll-Gergely et al. (2016)</bibRefCitation>
</th>
<td id="76955730CB13BEA5C17F35D3FC8DF2F8" box="[633,774,438,460]" gridcol="1" gridrow="6" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C17F35D3FC8DF2F8" authorityName="Bourguignat" authorityYear="1877" box="[633,774,438,460]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
</td>
<td id="76955730CB13BEA5C05135D3FBAFF2F8" box="[855,1060,438,460]" gridcol="2" gridrow="6" pageId="2" pageNumber="183">-</td>
<td id="76955730CB13BEA5C74135D3FB21F2F8" box="[1095,1194,438,460]" gridcol="3" gridrow="6" pageId="2" pageNumber="183">-</td>
<td id="76955730CB13BEA5C61335D3FA1FF2F8" box="[1301,1428,438,460]" gridcol="4" gridrow="6" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C61335D3FA1FF2FF" authorityName="Ancey" authorityYear="1887" box="[1301,1428,438,459]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">Chalepotaxis</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<caption id="DF0B6C86CB134158C38E367EFC63F105" box="[136,1000,539,561]" pageId="2" pageNumber="183" startId="2.[136,191,539,561]">
<paragraph id="8BCB3C0ECB134158C38E367EFC63F105" blockId="2.[136,1000,539,561]" box="[136,1000,539,561]" pageId="2" pageNumber="183">
Table 2. Reported distribution of
<taxonomicName id="4C74478DCB134158C2D2367EFDAAF104" authorityName="Benson" authorityYear="1860" box="[468,545,539,560]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C2D2367EFDAAF104" box="[468,545,539,560]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
</taxonomicName>
,
<emphasis id="B900E01CCB134158C128367EFD54F104" box="[558,735,539,560]" italics="true" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C128367EFD24F104" authorityName="Ancey" authorityYear="1887" box="[558,687,539,560]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">Chalepotaxis</taxonomicName>
, and
</emphasis>
<taxonomicName id="4C74478DCB134158C1E1367EFC63F105" box="[743,1000,539,561]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B900E01CCB134158C1E1367EFC1FF104" box="[743,916,539,560]" italics="true" pageId="2" pageNumber="183">Aenigmatoconcha</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BCB3C0ECB134158C38E3631FA49F15F" blockId="2.[136,1474,596,647]" box="[136,1474,596,619]" pageId="2" pageNumber="183">
Reference: 1 =
<bibRefCitation id="EFE541FFCB134158C2263630FE36F15E" author="Benson WH" box="[288,445,596,618]" pageId="2" pageNumber="183" pagination="387 - 392" refId="ref3754" refString="Benson WH (1859 a) New Helicidae collected by W. Theobald, Esq., jun., in Burmah and the Khasia Hills, and characters of a new Burmese Streptaxis described by W. H. Benson. The Annals and Magazine of Natural History, Series 3, Volume 3: 387 - 392." type="journal article" year="1859">Benson (1859a)</bibRefCitation>
; 2 =
<bibRefCitation id="EFE541FFCB134158C2F43630FD1BF15E" author="Benson WH" box="[498,656,596,618]" pageId="2" pageNumber="183" pagination="471 - 474" refId="ref3814" refString="Benson WH (1859 b) Characters of a new Burmese Streptaxis and of two forms belonging to a peculiar section of Helix collected by Captain Richard H. Sankey, Madras Engineers. The Annals and Magazine of Natural History, Series 3, Volume 3: 471 - 474." type="journal article" year="1859">Benson (1859b)</bibRefCitation>
; 3 =
<bibRefCitation id="EFE541FFCB134158C1C23630FCDCF15E" author="Benson WH" box="[708,855,596,618]" pageId="2" pageNumber="183" pagination="26 - 27" refId="ref3866" refString="Benson WH (1860) On the Burmese genus Sophina. The Annals and Magazine of Natural History, Series 3, Volume 5: 26 - 27." type="journal article" year="1860">Benson (1860)</bibRefCitation>
; 4 =
<bibRefCitation id="EFE541FFCB134158C08D3631FBA5F15F" author="Stoliczka F" box="[907,1070,596,619]" pageId="2" pageNumber="183" pagination="143 - 177" refId="ref4305" refString="Stoliczka F (1871) Note on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Province), with descriptions of new species. Journal of the Asiatic Society of Bengal, 40: 143 - 177, pls. 6 - 8; 217 - 259, pls. 15 - 19." type="journal article" year="1871">Stoliczka (1871)</bibRefCitation>
; 5 =
<bibRefCitation id="EFE541FFCB134158C7653631FA35F15F" author="Blanford WT &amp; Godwin-Austen HH" box="[1123,1470,596,619]" pageId="2" pageNumber="183" refId="ref3895" refString="Blanford WT &amp; Godwin-Austen HH (1908). The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, 311 pp." type="book" year="1908">Blanford &amp; Godwin-Austen (1908)</bibRefCitation>
;
</paragraph>
<paragraph id="8BCB3C0ECB134158C38F3614FD37F1B3" blockId="2.[136,1474,596,647]" box="[137,700,625,647]" pageId="2" pageNumber="183">
6 =
<bibRefCitation id="EFE541FFCB134158C3B43614FED6F1B3" author="Schileyko AA" box="[178,349,625,647]" pageId="2" pageNumber="183" pagination="1 - 68" refId="ref4276" refString="Schileyko AA (2011) Check-list of land pulmonate mollucs of Vietnam (Gastropoda: Stylommatophora). Ruthenica, 21 (1): 1 - 68." type="journal article" year="2011">Schileyko (2011)</bibRefCitation>
; and 7 = Páll-Gergely et al., (2016)
</paragraph>
<paragraph id="8BCB3C0ECB134158C39136CEFA2AF60A" pageId="2" pageNumber="183">
<table id="F974CEAECB13BEA5C39136CEFA2DF60A" box="[151,1446,683,1342]" gridcols="4" gridrows="12" pageId="2" pageNumber="183">
<tr id="35443E4CCB13BEA5C39136CEFA2DF1F5" box="[151,1446,683,705]" gridrow="0" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39136CEFF31F1F5" box="[151,186,683,705]" gridcol="0" gridrow="0" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C39136CEFF31F1F4" bold="true" box="[151,186,683,704]" pageId="2" pageNumber="183">No.</emphasis>
</th>
<th id="76955730CB13BEA5C3DC36CEFD29F1F5" box="[218,674,683,705]" gridcol="1" gridrow="0" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C2A336CEFE52F1F4" bold="true" box="[421,473,683,704]" pageId="2" pageNumber="183">Taxa</emphasis>
</th>
<th id="76955730CB13BEA5C1C336CEFA87F1F5" box="[709,1292,683,705]" gridcol="2" gridrow="0" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C0B036CEFB92F1F5" bold="true" box="[950,1049,683,705]" pageId="2" pageNumber="183">Localities</emphasis>
</th>
<th id="76955730CB13BEA5C63036CEFA2DF1F5" box="[1334,1446,683,705]" gridcol="3" gridrow="0" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C63036CEFA2DF1F5" bold="true" box="[1334,1446,683,705]" pageId="2" pageNumber="183">References</emphasis>
</th>
</tr>
<tr id="35443E4CCB13BEA5C3913684FA2DF020" box="[151,1446,737,788]" gridrow="1" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913684FF31F020" box="[151,186,737,788]" gridcol="0" gridrow="1" pageId="2" pageNumber="183">1</th>
<td id="76955730CB13BEA5C3DC3684FD29F020" box="[218,674,737,788]" gridcol="1" gridrow="1" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3687FE66F1C3" authority="(Benson, 1859 a)" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[218,493,737,759]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="forabilis">
<emphasis id="B900E01CCB134158C3DC3687FECFF1C2" box="[218,324,737,759]" italics="true" pageId="2" pageNumber="183">S. forabilis</emphasis>
(
<bibRefCitation id="EFE541FFCB134158C2553687FE6DF1C3" author="Benson WH" box="[339,486,737,759]" pageId="2" pageNumber="183" pagination="387 - 392" refId="ref3754" refString="Benson WH (1859 a) New Helicidae collected by W. Theobald, Esq., jun., in Burmah and the Khasia Hills, and characters of a new Burmese Streptaxis described by W. H. Benson. The Annals and Magazine of Natural History, Series 3, Volume 3: 387 - 392." type="journal article" year="1859">Benson, 1859a</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33684FA87F020" box="[709,1292,737,788]" gridcol="2" gridrow="1" pageId="2" pageNumber="183">Phie-Than Valley, Tenassarim Caves in limestone hills at Damotha, near Moulmein, Myanmar</td>
<td id="76955730CB13BEA5C6303684FA2DF020" box="[1334,1446,737,788]" gridcol="3" gridrow="1" pageId="2" pageNumber="183">1, 3, 4, 5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913748FA2DF077" box="[151,1446,813,835]" gridrow="2" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913748FF31F077" box="[151,186,813,835]" gridcol="0" gridrow="2" pageId="2" pageNumber="183">2</th>
<td id="76955730CB13BEA5C3DC3748FD29F077" box="[218,674,813,835]" gridcol="1" gridrow="2" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3748FECFF076" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[218,324,813,834]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="forabilis">
<emphasis id="B900E01CCB134158C3DC3748FECFF076" box="[218,324,813,834]" italics="true" pageId="2" pageNumber="183">S. forabilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C74478DCB134158C254374BFD29F076" authority="Godwin-Austen, 1908" authorityName="Godwin-Austen" authorityYear="1908" box="[338,674,813,835]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="variety" species="forabilis" variety="bensoni">var bensoni Godwin-Austen, 1908</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33748FA87F077" box="[709,1292,813,835]" gridcol="2" gridrow="2" pageId="2" pageNumber="183">Damotha, near Moulmein, Myanmar</td>
<td id="76955730CB13BEA5C6303748FA2DF077" box="[1334,1446,813,835]" gridcol="3" gridrow="2" pageId="2" pageNumber="183">5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913739FA2DF046" box="[151,1446,860,882]" gridrow="3" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913739FF31F046" box="[151,186,860,882]" gridcol="0" gridrow="3" pageId="2" pageNumber="183">3</th>
<td id="76955730CB13BEA5C3DC3739FD29F046" box="[218,674,860,882]" gridcol="1" gridrow="3" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3739FE5DF046" authority="(Benson, 1859 b)" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[218,470,860,882]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="calias">
<emphasis id="B900E01CCB134158C3DC3739FEA7F045" box="[218,300,860,881]" italics="true" pageId="2" pageNumber="183">S. calias</emphasis>
(
<bibRefCitation id="EFE541FFCB134158C23D3739FE44F046" author="Benson WH" box="[315,463,860,882]" pageId="2" pageNumber="183" pagination="471 - 474" refId="ref3814" refString="Benson WH (1859 b) Characters of a new Burmese Streptaxis and of two forms belonging to a peculiar section of Helix collected by Captain Richard H. Sankey, Madras Engineers. The Annals and Magazine of Natural History, Series 3, Volume 3: 471 - 474." type="journal article" year="1859">Benson, 1859b</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33739FA87F046" box="[709,1292,860,882]" gridcol="2" gridrow="3" pageId="2" pageNumber="183">Near Moulmein, Myanmar.</td>
<td id="76955730CB13BEA5C6303739FA2DF046" box="[1334,1446,860,882]" gridcol="3" gridrow="3" pageId="2" pageNumber="183">2, 3, 4, 5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913719FA2DF09B" box="[151,1446,892,943]" gridrow="4" pageId="2" pageNumber="183" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76955730CB13BEA5C1C33719FA87F09B" box="[709,1292,892,943]" gridcol="2" gridrow="4" pageId="2" pageNumber="183">Limestone hills to east and south of Moulmein at the Farm Caves and Tavoy, Myanmar</td>
</tr>
<tr id="35443E4CCB13BEA5C39137A2FA2DF0E9" box="[151,1446,967,989]" gridrow="5" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39137A2FF31F0E9" box="[151,186,967,989]" gridcol="0" gridrow="5" pageId="2" pageNumber="183">4</th>
<td id="76955730CB13BEA5C3DC37A2FD29F0E9" box="[218,674,967,989]" gridcol="1" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC37ADFD9AF0E9" authority="(Benson, 1859 b)" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[218,529,967,989]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="schistostelis">
<emphasis id="B900E01CCB134158C3DC37ADFEECF0E8" box="[218,359,967,989]" italics="true" pageId="2" pageNumber="183">S. schistostelis</emphasis>
(
<bibRefCitation id="EFE541FFCB134158C27037ADFD81F0E9" author="Benson WH" box="[374,522,967,989]" pageId="2" pageNumber="183" pagination="471 - 474" refId="ref3814" refString="Benson WH (1859 b) Characters of a new Burmese Streptaxis and of two forms belonging to a peculiar section of Helix collected by Captain Richard H. Sankey, Madras Engineers. The Annals and Magazine of Natural History, Series 3, Volume 3: 471 - 474." type="journal article" year="1859">Benson, 1859b</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C337A2FA87F0E9" box="[709,1292,967,989]" gridcol="2" gridrow="5" pageId="2" pageNumber="183">Near Moulmein, Myanmar</td>
<td id="76955730CB13BEA5C63037A2FA2DF0E9" box="[1334,1446,967,989]" gridcol="3" gridrow="5" pageId="2" pageNumber="183">2, 4, 5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913793FA2DF71D" box="[151,1446,1014,1065]" gridrow="6" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913793FF31F71D" box="[151,186,1014,1065]" gridcol="0" gridrow="6" pageId="2" pageNumber="183">5</th>
<td id="76955730CB13BEA5C3DC3793FD29F71D" box="[218,674,1014,1065]" gridcol="1" gridrow="6" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3792FE71F738" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[218,506,1014,1036]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="discoidalis">
<emphasis id="B900E01CCB134158C3DC3792FED0F73F" box="[218,347,1014,1036]" italics="true" pageId="2" pageNumber="183">S. discoidalis</emphasis>
<bibRefCitation id="EFE541FFCB134158C2653793FE71F738" author="Stoliczka F" box="[355,506,1014,1036]" pageId="2" pageNumber="183" pagination="143 - 177" refId="ref4305" refString="Stoliczka F (1871) Note on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Province), with descriptions of new species. Journal of the Asiatic Society of Bengal, 40: 143 - 177, pls. 6 - 8; 217 - 259, pls. 15 - 19." type="journal article" year="1871">Stoliczka, 1871</bibRefCitation>
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33793FA87F71D" box="[709,1292,1014,1065]" gridcol="2" gridrow="6" pageId="2" pageNumber="183">Limestone hills of the Farm Caves on Attaran River and South of Moulmein, Myanmar</td>
<td id="76955730CB13BEA5C6303793FA2DF71D" box="[1334,1446,1014,1065]" gridcol="3" gridrow="6" pageId="2" pageNumber="183">4, 5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913027FA2DF76C" box="[151,1446,1090,1112]" gridrow="7" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913027FF31F76C" box="[151,186,1090,1112]" gridcol="0" gridrow="7" pageId="2" pageNumber="183">6</th>
<td id="76955730CB13BEA5C3DC3027FD29F76C" box="[218,674,1090,1112]" gridcol="1" gridrow="7" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3027FD8BF763" authority="Stoliczka, 1871" authorityName="Stoliczka" authorityYear="1871" box="[218,512,1090,1112]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="conjungens">
<emphasis id="B900E01CCB134158C3DC3027FEEBF76C" box="[218,352,1090,1112]" italics="true" pageId="2" pageNumber="183">S. conjungens</emphasis>
<bibRefCitation id="EFE541FFCB134158C26E3027FD8BF763" author="Stoliczka F" box="[360,512,1090,1112]" pageId="2" pageNumber="183" pagination="143 - 177" refId="ref4305" refString="Stoliczka F (1871) Note on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Province), with descriptions of new species. Journal of the Asiatic Society of Bengal, 40: 143 - 177, pls. 6 - 8; 217 - 259, pls. 15 - 19." type="journal article" year="1871">Stoliczka, 1871</bibRefCitation>
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33027FA87F76C" box="[709,1292,1090,1112]" gridcol="2" gridrow="7" pageId="2" pageNumber="183">South of Moulmein, Myanmar</td>
<td id="76955730CB13BEA5C6303027FA2DF76C" box="[1334,1446,1090,1112]" gridcol="3" gridrow="7" pageId="2" pageNumber="183">4, 5</td>
</tr>
<tr id="35443E4CCB13BEA5C3913014FA2DF7B3" box="[151,1446,1137,1159]" gridrow="8" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913014FF31F7B3" box="[151,186,1137,1159]" gridcol="0" gridrow="8" pageId="2" pageNumber="183">7</th>
<td id="76955730CB13BEA5C3DC3014FD29F7B3" box="[218,674,1137,1159]" gridcol="1" gridrow="8" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC3014FE67F7B3" authority="(Gredler, 1881)" baseAuthorityName="Gredler" baseAuthorityYear="1881" box="[218,492,1137,1159]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="infantilis">
<emphasis id="B900E01CCB134158C3DC3014FEC7F7B2" box="[218,332,1137,1158]" italics="true" pageId="2" pageNumber="183">C. infantilis</emphasis>
(Gredler, 1881)
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C33014FA87F7B3" box="[709,1292,1137,1159]" gridcol="2" gridrow="8" pageId="2" pageNumber="183">Hupei, Guang Xi, and Hunan, Southern China</td>
<td id="76955730CB13BEA5C6303014FA2DF7B3" box="[1334,1446,1137,1159]" gridcol="3" gridrow="8" pageId="2" pageNumber="183">6, 7</td>
</tr>
<tr id="35443E4CCB13BEA5C39130F4FA2DF7D4" box="[151,1446,1169,1248]" gridrow="9" pageId="2" pageNumber="183" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76955730CB13BEA5C1C330F4FA87F7D4" box="[709,1292,1169,1248]" gridcol="2" gridrow="9" pageId="2" pageNumber="183">Ha Giang, Ha Lang, Quang Uyen, Lang Son, Hái Phong, Dĕ Ma Phuc, Ke Bao and Krieu islands, Northern Vietnam Luang Prabang, Laos</td>
</tr>
<tr id="35443E4CCB13BEA5C391309CFA2DF63B" box="[151,1446,1273,1295]" gridrow="10" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C391309CFF31F63B" box="[151,186,1273,1295]" gridcol="0" gridrow="10" pageId="2" pageNumber="183">8</th>
<td id="76955730CB13BEA5C3DC309CFD29F63B" box="[218,674,1273,1295]" gridcol="1" gridrow="10" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC309FFDD4F63B" authority="(Schmacker &amp; Boetter, 1891)" baseAuthorityName="Schmacker &amp; Boetter" baseAuthorityYear="1891" box="[218,607,1273,1295]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="spadix">
<emphasis id="B900E01CCB134158C3DC309FFEBDF63A" box="[218,310,1273,1295]" italics="true" pageId="2" pageNumber="183">C. spadix</emphasis>
(Schmacker &amp; Boetter, 1891)
</taxonomicName>
</td>
<td id="76955730CB13BEA5C1C3309CFA87F63B" box="[709,1292,1273,1295]" gridcol="2" gridrow="10" pageId="2" pageNumber="183">Taiwan</td>
<td id="76955730CB13BEA5C630309CFA2DF63B" box="[1334,1446,1273,1295]" gridcol="3" gridrow="10" pageId="2" pageNumber="183">7</td>
</tr>
<tr id="35443E4CCB13BEA5C391314DFA2DF60A" box="[151,1446,1320,1342]" gridrow="11" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C391314DFF31F60A" box="[151,186,1320,1342]" gridcol="0" gridrow="11" pageId="2" pageNumber="183">9</th>
<td id="76955730CB13BEA5C3DC314DFD29F60A" box="[218,674,1320,1342]" gridcol="1" gridrow="11" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3DC314DFEC2F609" authority="Tumpeesuwan &amp; Tumpeesuwan, 2017" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[218,329,1320,1341]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="clivicola" status="sp. nov.">
<emphasis id="B900E01CCB134158C3DC314DFEC2F609" box="[218,329,1320,1341]" italics="true" pageId="2" pageNumber="183">A. clivicola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2335D67CB134158C250314CFE40F60A" box="[342,459,1320,1342]" pageId="2" pageNumber="183" rank="species">new species</taxonomicNameLabel>
</td>
<td id="76955730CB13BEA5C1C3314DFA87F60A" box="[709,1292,1320,1342]" gridcol="2" gridrow="11" pageId="2" pageNumber="183">Phu Pha Lom, Loei, Thailand</td>
<td id="76955730CB13BEA5C630314DFA2DF60A" box="[1334,1446,1320,1342]" gridcol="3" gridrow="11" pageId="2" pageNumber="183">This study</td>
</tr>
</table>
</paragraph>
<caption id="DF0B6C86CB134158C38E31E8FC63F697" box="[136,1000,1421,1443]" pageId="2" pageNumber="183" startId="2.[136,191,1421,1443]">
<paragraph id="8BCB3C0ECB134158C38E31E8FC63F697" blockId="2.[136,1000,1421,1443]" box="[136,1000,1421,1443]" pageId="2" pageNumber="183">
Table 3. Comparison of shells of
<taxonomicName id="4C74478DCB134158C2D331E8FDA9F696" authorityName="Benson" authorityYear="1860" box="[469,546,1421,1442]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C2D331E8FDA9F696" box="[469,546,1421,1442]" italics="true" pageId="2" pageNumber="183">Sophina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C74478DCB134158C13631E8FD24F696" authorityName="Ancey" authorityYear="1887" box="[560,687,1421,1442]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB134158C13631E8FD24F696" box="[560,687,1421,1442]" italics="true" pageId="2" pageNumber="183">Chalepotaxis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C74478DCB134158C1EE31E8FC63F697" box="[744,1000,1421,1443]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B900E01CCB134158C1EE31E8FC1EF696" box="[744,917,1421,1442]" italics="true" pageId="2" pageNumber="183">Aenigmatoconcha</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BCB3C0ECB134158C64831B1FA13F40F" pageId="2" pageNumber="183">
<table id="F974CEAECB13BEA5C39131B1FA2AF40F" box="[151,1441,1492,1851]" gridcols="7" gridrows="10" pageId="2" pageNumber="183">
<tr id="35443E4CCB13BEA5C39131B1FA2AF6DD" box="[151,1441,1492,1513]" gridrow="0" pageId="2" pageNumber="183" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76955730CB13BEA5C64C31B1FA2AF6DD" box="[1354,1441,1492,1513]" gridcol="6" gridrow="0" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C64831B1FA16F6DD" bold="true" box="[1358,1437,1492,1513]" pageId="2" pageNumber="183">Ventral</emphasis>
</th>
</tr>
<tr id="35443E4CCB13BEA5C3913194FA2AF517" box="[151,1441,1521,1571]" gridrow="1" pageId="2" pageNumber="183" rowspan-0="1">
<td id="76955730CB13BEA5C2A23194FDF4F517" box="[420,639,1521,1571]" gridcol="1" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C2D73194FDD9F533" bold="true" box="[465,594,1521,1543]" pageId="2" pageNumber="183">Shell: Shape</emphasis>
</td>
<td id="76955730CB13BEA5C1983194FCA0F517" box="[670,811,1521,1571]" gridcol="2" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C1AE3194FCA9F533" bold="true" box="[680,802,1521,1543]" pageId="2" pageNumber="183">Shell height</emphasis>
</td>
<td id="76955730CB13BEA5C04A3194FC7AF517" box="[844,1009,1521,1571]" gridcol="3" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C0623194FC51F533" bold="true" box="[868,986,1521,1543]" pageId="2" pageNumber="183">Shell width</emphasis>
</td>
<td id="76955730CB13BEA5C71E3194FBEFF517" box="[1048,1124,1521,1571]" gridcol="4" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C71E3194FBEFF532" bold="true" box="[1048,1124,1521,1542]" pageId="2" pageNumber="183">Whorls</emphasis>
</td>
<td id="76955730CB13BEA5C7963194FA98F517" box="[1168,1299,1521,1571]" gridcol="5" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C79B3194FA8EF533" bold="true" box="[1181,1285,1521,1543]" pageId="2" pageNumber="183">Umbilicus</emphasis>
</td>
<td id="76955730CB13BEA5C64C3194FA2AF517" box="[1354,1441,1521,1571]" gridcol="6" gridrow="1" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C65E3194FA2AF517" bold="true" pageId="2" pageNumber="183">spiral striation</emphasis>
</td>
</tr>
<tr id="35443E4CCB13BEA5C3913221FA2AF56E" box="[151,1441,1604,1626]" gridrow="2" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913221FE0DF56E" box="[151,390,1604,1626]" gridcol="0" gridrow="2" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3913220FF61F56D" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[151,234,1604,1626]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="calias">
<emphasis id="B900E01CCB134158C3913220FF61F56D" box="[151,234,1604,1626]" italics="true" pageId="2" pageNumber="183">S. calias</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A23221FDF4F56E" box="[420,639,1604,1626]" gridcol="1" gridrow="2" pageId="2" pageNumber="183">Dispersedly orbiculate</td>
<td id="76955730CB13BEA5C1983221FCA0F56E" box="[670,811,1604,1626]" gridcol="2" gridrow="2" pageId="2" pageNumber="183">4–6 mm</td>
<td id="76955730CB13BEA5C04A3221FC7AF56E" box="[844,1009,1604,1626]" gridcol="3" gridrow="2" pageId="2" pageNumber="183">9–15 mm</td>
<td id="76955730CB13BEA5C71E3221FBEFF56E" box="[1048,1124,1604,1626]" gridcol="4" gridrow="2" pageId="2" pageNumber="183">5</td>
<td id="76955730CB13BEA5C7963221FA98F56E" box="[1168,1299,1604,1626]" gridcol="5" gridrow="2" pageId="2" pageNumber="183">Narrow</td>
<td id="76955730CB13BEA5C64C3221FA2AF56E" box="[1354,1441,1604,1626]" gridcol="6" gridrow="2" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C3913206FA2AF54E" box="[151,1441,1635,1658]" gridrow="3" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913206FE0DF54E" box="[151,390,1635,1658]" gridcol="0" gridrow="3" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3913201FEAEF54D" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[151,293,1636,1657]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="schistostelis">
<emphasis id="B900E01CCB134158C3913201FEAEF54D" box="[151,293,1636,1657]" italics="true" pageId="2" pageNumber="183">S. schistostelis</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A23206FDF4F54E" box="[420,639,1635,1658]" gridcol="1" gridrow="3" pageId="2" pageNumber="183">Globosely depressed</td>
<td id="76955730CB13BEA5C1983206FCA0F54E" box="[670,811,1635,1658]" gridcol="2" gridrow="3" pageId="2" pageNumber="183">8.5 mm</td>
<td id="76955730CB13BEA5C04A3206FC7AF54E" box="[844,1009,1635,1658]" gridcol="3" gridrow="3" pageId="2" pageNumber="183">16 mm</td>
<td id="76955730CB13BEA5C71E3206FBEFF54E" box="[1048,1124,1635,1658]" gridcol="4" gridrow="3" pageId="2" pageNumber="183">4½–5½</td>
<td id="76955730CB13BEA5C7963206FA98F54E" box="[1168,1299,1635,1658]" gridcol="5" gridrow="3" pageId="2" pageNumber="183">Narrow</td>
<td id="76955730CB13BEA5C64C3206FA2AF54E" box="[1354,1441,1635,1658]" gridcol="6" gridrow="3" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C39132E7FA2AF5AE" box="[151,1441,1666,1690]" gridrow="4" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39132E7FE0DF5AE" box="[151,390,1666,1690]" gridcol="0" gridrow="4" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C39132E1FE92F5AD" authorityName="Stoliczka" authorityYear="1871" box="[151,281,1668,1689]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="discoidalis">
<emphasis id="B900E01CCB134158C39132E1FE92F5AD" box="[151,281,1668,1689]" italics="true" pageId="2" pageNumber="183">S. discoidalis</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A232E7FDF4F5AE" box="[420,639,1666,1690]" gridcol="1" gridrow="4" pageId="2" pageNumber="183">Depressed subdiscoid</td>
<td id="76955730CB13BEA5C19832E7FCA0F5AE" box="[670,811,1666,1690]" gridcol="2" gridrow="4" pageId="2" pageNumber="183">4.5 mm</td>
<td id="76955730CB13BEA5C04A32E7FC7AF5AE" box="[844,1009,1666,1690]" gridcol="3" gridrow="4" pageId="2" pageNumber="183">10.5 mm</td>
<td id="76955730CB13BEA5C71E32E7FBEFF5AE" box="[1048,1124,1666,1690]" gridcol="4" gridrow="4" pageId="2" pageNumber="183">5–5½</td>
<td id="76955730CB13BEA5C79632E7FA98F5AE" box="[1168,1299,1666,1690]" gridcol="5" gridrow="4" pageId="2" pageNumber="183">Wide</td>
<td id="76955730CB13BEA5C64C32E7FA2AF5AE" box="[1354,1441,1666,1690]" gridcol="6" gridrow="4" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C39132C6FA2AF58D" box="[151,1441,1699,1721]" gridrow="5" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39132C6FE0DF58D" box="[151,390,1699,1721]" gridcol="0" gridrow="5" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C39132C1FE95F58D" authorityName="Stoliczka" authorityYear="1871" box="[151,286,1700,1721]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="conjungens">
<emphasis id="B900E01CCB134158C39132C1FE95F58D" box="[151,286,1700,1721]" italics="true" pageId="2" pageNumber="183">S. conjungens</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A232C6FDF4F58D" box="[420,639,1699,1721]" gridcol="1" gridrow="5" pageId="2" pageNumber="183">Globosely orbiculate</td>
<td id="76955730CB13BEA5C19832C6FCA0F58D" box="[670,811,1699,1721]" gridcol="2" gridrow="5" pageId="2" pageNumber="183">5 mm</td>
<td id="76955730CB13BEA5C04A32C6FC7AF58D" box="[844,1009,1699,1721]" gridcol="3" gridrow="5" pageId="2" pageNumber="183">12 mm</td>
<td id="76955730CB13BEA5C71E32C6FBEFF58D" box="[1048,1124,1699,1721]" gridcol="4" gridrow="5" pageId="2" pageNumber="183">5</td>
<td id="76955730CB13BEA5C79632C6FA98F58D" box="[1168,1299,1699,1721]" gridcol="5" gridrow="5" pageId="2" pageNumber="183">Slightly wide</td>
<td id="76955730CB13BEA5C64C32C6FA2AF58D" box="[1354,1441,1699,1721]" gridcol="6" gridrow="5" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C39132A7FA2AF5ED" box="[151,1441,1730,1753]" gridrow="6" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C39132A7FE0DF5ED" box="[151,390,1730,1753]" gridcol="0" gridrow="6" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C39132A1FE89F5EC" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[151,258,1731,1753]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="forabilis">
<emphasis id="B900E01CCB134158C39132A1FE89F5EC" box="[151,258,1731,1753]" italics="true" pageId="2" pageNumber="183">S. forabilis</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A232A7FDF4F5ED" box="[420,639,1730,1753]" gridcol="1" gridrow="6" pageId="2" pageNumber="183">Semiglobosely conoid</td>
<td id="76955730CB13BEA5C19832A7FCA0F5ED" box="[670,811,1730,1753]" gridcol="2" gridrow="6" pageId="2" pageNumber="183">4 mm</td>
<td id="76955730CB13BEA5C04A32A7FC7AF5ED" box="[844,1009,1730,1753]" gridcol="3" gridrow="6" pageId="2" pageNumber="183">8–9 mm</td>
<td id="76955730CB13BEA5C71E32A7FBEFF5ED" box="[1048,1124,1730,1753]" gridcol="4" gridrow="6" pageId="2" pageNumber="183">5½–6</td>
<td id="76955730CB13BEA5C79632A7FA98F5ED" box="[1168,1299,1730,1753]" gridcol="5" gridrow="6" pageId="2" pageNumber="183">Wide</td>
<td id="76955730CB13BEA5C64C32A7FA2AF5ED" box="[1354,1441,1730,1753]" gridcol="6" gridrow="6" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C3913286FA2AF5CF" box="[151,1441,1763,1787]" gridrow="7" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913286FE0DF5CF" box="[151,390,1763,1787]" gridcol="0" gridrow="7" pageId="2" pageNumber="183">
<emphasis id="B900E01CCB134158C3913286FE9BF5CC" box="[151,272,1763,1784]" italics="true" pageId="2" pageNumber="183">S. f. bensoni</emphasis>
</th>
<td id="76955730CB13BEA5C2A23286FDF4F5CF" box="[420,639,1763,1787]" gridcol="1" gridrow="7" pageId="2" pageNumber="183">Globosely conoid</td>
<td id="76955730CB13BEA5C1983286FCA0F5CF" box="[670,811,1763,1787]" gridcol="2" gridrow="7" pageId="2" pageNumber="183">4 mm</td>
<td id="76955730CB13BEA5C04A3286FC7AF5CF" box="[844,1009,1763,1787]" gridcol="3" gridrow="7" pageId="2" pageNumber="183">7.5 mm</td>
<td id="76955730CB13BEA5C71E3286FBEFF5CF" box="[1048,1124,1763,1787]" gridcol="4" gridrow="7" pageId="2" pageNumber="183">
5¼–5
<emphasis id="B900E01CCB134158C7483280FBEFF5CF" bold="true" box="[1102,1124,1765,1787]" pageId="2" pageNumber="183">⅕</emphasis>
</td>
<td id="76955730CB13BEA5C7963286FA98F5CF" box="[1168,1299,1763,1787]" gridcol="5" gridrow="7" pageId="2" pageNumber="183">Slightly wide</td>
<td id="76955730CB13BEA5C64C3286FA2AF5CF" box="[1354,1441,1763,1787]" gridcol="6" gridrow="7" pageId="2" pageNumber="183">Absent</td>
</tr>
<tr id="35443E4CCB13BEA5C3913360FA2AF42F" box="[151,1441,1797,1819]" gridrow="8" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913360FE0DF42F" box="[151,390,1797,1819]" gridcol="0" gridrow="8" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3913360FE81F42E" baseAuthorityName="Gredler" baseAuthorityYear="1881" box="[151,266,1797,1818]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="infantilis">
<emphasis id="B900E01CCB134158C3913360FE81F42E" box="[151,266,1797,1818]" italics="true" pageId="2" pageNumber="183">C. infantilis</emphasis>
</taxonomicName>
</th>
<td id="76955730CB13BEA5C2A23360FDF4F42F" box="[420,639,1797,1819]" gridcol="1" gridrow="8" pageId="2" pageNumber="183">Depressed conical</td>
<td id="76955730CB13BEA5C1983360FCA0F42F" box="[670,811,1797,1819]" gridcol="2" gridrow="8" pageId="2" pageNumber="183">4–5 mm</td>
<td id="76955730CB13BEA5C04A3360FC7AF42F" box="[844,1009,1797,1819]" gridcol="3" gridrow="8" pageId="2" pageNumber="183">6–7 mm</td>
<td id="76955730CB13BEA5C71E3360FBEFF42F" box="[1048,1124,1797,1819]" gridcol="4" gridrow="8" pageId="2" pageNumber="183">6</td>
<td id="76955730CB13BEA5C7963360FA98F42F" box="[1168,1299,1797,1819]" gridcol="5" gridrow="8" pageId="2" pageNumber="183">Narrow</td>
<td id="76955730CB13BEA5C64C3360FA2AF42F" box="[1354,1441,1797,1819]" gridcol="6" gridrow="8" pageId="2" pageNumber="183">—</td>
</tr>
<tr id="35443E4CCB13BEA5C3913346FA2AF40F" box="[151,1441,1827,1851]" gridrow="9" pageId="2" pageNumber="183">
<th id="76955730CB13BEA5C3913346FE0DF40F" box="[151,390,1827,1851]" gridcol="0" gridrow="9" pageId="2" pageNumber="183">
<taxonomicName id="4C74478DCB134158C3913340FE8EF40E" authority="Tumpeesuwan &amp; Tumpeesuwan, 2017" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[151,261,1829,1850]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="183" phylum="Mollusca" rank="species" species="clivicola" status="sp. nov.">
<emphasis id="B900E01CCB134158C3913340FE8EF40E" box="[151,261,1829,1850]" italics="true" pageId="2" pageNumber="183">A. clivicola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2335D67CB134158C2143340FE0DF40F" box="[274,390,1829,1851]" pageId="2" pageNumber="183" rank="species">new species</taxonomicNameLabel>
</th>
<td id="76955730CB13BEA5C2A23346FDF4F40F" box="[420,639,1827,1851]" gridcol="1" gridrow="9" pageId="2" pageNumber="183">Flattened to depressed</td>
<td id="76955730CB13BEA5C1983346FCA0F40F" box="[670,811,1827,1851]" gridcol="2" gridrow="9" pageId="2" pageNumber="183">7.69–9.89 mm</td>
<td id="76955730CB13BEA5C04A3346FC7AF40F" box="[844,1009,1827,1851]" gridcol="3" gridrow="9" pageId="2" pageNumber="183">15.40–18.62 mm</td>
<td id="76955730CB13BEA5C71E3346FBEFF40F" box="[1048,1124,1827,1851]" gridcol="4" gridrow="9" pageId="2" pageNumber="183">5¼–5½</td>
<td id="76955730CB13BEA5C7963346FA98F40F" box="[1168,1299,1827,1851]" gridcol="5" gridrow="9" pageId="2" pageNumber="183">Slightly wide</td>
<td id="76955730CB13BEA5C64C3346FA2AF40F" box="[1354,1441,1827,1851]" gridcol="6" gridrow="9" pageId="2" pageNumber="183">Absent</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCB3C0ECB124159C37734D8FD7AF161" blockId="3.[113,753,189,597]" pageId="3" pageNumber="184">
<taxonomicName id="4C74478DCB124159C37734D8FEBDF3E1" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[113,310,189,213]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB124159C37734D8FEBDF3E1" box="[113,310,189,213]" italics="true" pageId="3" pageNumber="184">Aenigmatoconcha</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C74478DCB124159C2D434D8FDA0F3E1" authorityName="Benson" authorityYear="1860" box="[466,555,189,213]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB124159C2D434D8FDA0F3E1" box="[466,555,189,213]" italics="true" pageId="3" pageNumber="184">Sophina</emphasis>
</taxonomicName>
(
<taxonomicName id="4C74478DCB124159C13934DBFD14F3E1" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[575,671,189,213]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="species" species="calias">
<emphasis id="B900E01CCB124159C13934DBFD14F3E1" box="[575,671,189,213]" italics="true" pageId="3" pageNumber="184">S. calias</emphasis>
</taxonomicName>
and
<taxonomicName id="4C74478DCB124159C1DA34DBFF72F3C1" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="species" species="schistostelis">
<emphasis id="B900E01CCB124159C1DA34DBFF72F3C1" italics="true" pageId="3" pageNumber="184">S. schistostelis</emphasis>
</taxonomicName>
) by the absence of an amatorial organ and the gametolytic sac with three distinct portions, including thick wall, robust tube at proximal portion and slender tube at middle portion leading to bulbous and thin walled head of distal portion. The penis is also larger and stouter than in
<taxonomicName id="4C74478DCB124159C3CE351BFEADF2A1" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[200,294,381,405]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="species" species="calias">
<emphasis id="B900E01CCB124159C3CE351BFEADF2A1" box="[200,294,381,405]" italics="true" pageId="3" pageNumber="184">S. calias</emphasis>
</taxonomicName>
and
<taxonomicName id="4C74478DCB124159C266351BFE75F2A1" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[352,510,381,405]" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="species" species="schistostelis">
<emphasis id="B900E01CCB124159C266351BFE75F2A1" box="[352,510,381,405]" italics="true" pageId="3" pageNumber="184">S. schistostelis</emphasis>
</taxonomicName>
(
<tableCitation id="C6F609B5CB124159C1093518FDEEF2A1" box="[527,613,381,405]" captionStart="Table 4" captionStartId="5.[112,167,658,681]" captionText="Table 4. Comparison of known genital system and radula of Sophina, Chalepotaxis, and Aenigmatoconcha species. Characters in brackets were used by Stolizcka (1871), Blanford &amp; Godwin- Austen (1908), and Páll-Gergely et al. (2016)." pageId="3" pageNumber="184">Table 4</tableCitation>
and
<figureCitation id="134F208BCB124159C1993518FD69F2A1" box="[671,738,381,405]" captionStart="Fig" captionStartId="6.[135,170,1183,1205]" captionTargetBox="[135,1473,189,1168]" captionTargetId="figure-7@6.[135,1499,189,1168]" captionTargetPageId="6" captionText="Fig. 5. Aenigmatoconcha clivicola, new species, genital system of paratype (NHMSU-0014) and schematic drawing. Abbreviations: ag, albumen gland; at, atrium; ep, epiphallus; erc, epiphallic retractor caecum; fo, free oviduct; gs, gametolytic sac; hd, hermaphroditic duct; p, penis; pr, penial retractor; pro, prostate gland; ut, uterus; v, vagina; vd, vas deferens. Photograph and illustration: Kitti Tanmuangpak." figureDoi="http://doi.org/10.5281/zenodo.5355949" httpUri="https://zenodo.org/record/5355949/files/figure.png" pageId="3" pageNumber="184">Fig. 5</figureCitation>
).
<taxonomicName id="4C74478DCB124159C37735F8FEB2F281" authorityName="Tumpeesuwan &amp; Tumpeesuwan" authorityYear="2017" box="[113,313,413,437]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB124159C37735F8FEB2F281" box="[113,313,413,437]" italics="true" pageId="3" pageNumber="184">Aenigmatoconcha</emphasis>
</taxonomicName>
has more characters in common with
<taxonomicName id="4C74478DCB124159C37735D8FE89F2E1" authorityName="Ancey" authorityYear="1887" box="[113,258,445,469]" class="Gastropoda" family="Helicarionidae" genus="Chalepotaxis" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="184" phylum="Mollusca" rank="genus">
<emphasis id="B900E01CCB124159C37735D8FE89F2E1" box="[113,258,445,469]" italics="true" pageId="3" pageNumber="184">Chalepotaxis</emphasis>
</taxonomicName>
, including a radula with V-shaped rows of teeth, the absence of an amatorial organ and three distinct portions of gametolytic sac (
<bibRefCitation id="EFE541FFCB124159C2AF3598FD31F121" author="Pall-Gergely B &amp; Feher Z &amp; Othani JU &amp; Asami T" box="[425,698,509,533]" pageId="3" pageNumber="184" refId="ref4159" refString="Pall-Gergely B, Feher Z, Othani JU &amp; Asami T (2016) An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylomatophora: Helicarionidae). Molluscan Research, DOI: 10.1080 / 13235818.2016.1234996." type="book" year="2016">Páll-Gergely et al., 2016</bibRefCitation>
), but it differs in shell characters, radula teeth without smaller upper part, and short and stout penis without penial sheath.
</paragraph>
</subSubSection>
</treatment>
</document>
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