treatments-xml/data/03/DD/87/03DD87C8FFFFFFD03488AD798BB9DD58.xml

<document ID-DOI="10.5733/afin.056.0219" ID-GBIF-Dataset="30750ff4-b11b-493f-8ee6-6880dc9390c7" ID-ISSN="2305-2562" ID-Zenodo-Dep="7914328" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683585685456" checkinUser="felipe" docAuthor="Harvey, Mark S. &amp; Mahnert, Volker" docDate="2015" docId="03DD87C8FFFFFFD03488AD798BB9DD58" docLanguage="en" docName="AfrInvertebr.56.2.491-497.pdf" docOrigin="African Invertebrates 56 (2)" docSource="http://www.bioone.org/doi/10.5733/afin.056.0219" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Sphaerowithius ansieae Harvey &amp; Mahnert 2015, sp. n." docType="treatment" docVersion="4" lastPageNumber="496" masterDocId="FFE4FFB0FFFEFFD53563A87A8A49D96C" masterDocTitle="A new species of Sphaerowithius (Pseudoscorpiones, Withiidae) from Namibia" masterLastPageNumber="491" masterPageNumber="491" pageNumber="492" updateTime="1683635464251" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Sphaerowithius (Pseudoscorpiones, Withiidae) from Namibia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mahnert, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-08-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>491</mods:start>
<mods:end>491</mods:end>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.056.0219</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.056.0219</mods:identifier>
<mods:identifier type="GBIF-Dataset">30750ff4-b11b-493f-8ee6-6880dc9390c7</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7914328</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7914336" ID-Zenodo-Dep="7914336" LSID="urn:lsid:plazi:treatment:03DD87C8FFFFFFD03488AD798BB9DD58" httpUri="http://treatment.plazi.org/id/03DD87C8FFFFFFD03488AD798BB9DD58" lastPageId="5" lastPageNumber="496" pageId="1" pageNumber="492">
<subSubSection box="[491,832,1283,1311]" pageId="1" pageNumber="492" type="nomenclature">
<paragraph blockId="1.[181,1142,1283,1677]" box="[491,832,1283,1311]" pageId="1" pageNumber="492">
<heading box="[491,832,1283,1311]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="492" reason="2">
<taxonomicName ID-CoL="4YSWD" authority="Harvey &amp; Mahnert, 2015" authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[491,765,1283,1310]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
<emphasis bold="true" box="[491,765,1283,1310]" italics="true" pageId="1" pageNumber="492">Sphaerowithius ansieae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[772,832,1285,1311]" pageId="1" pageNumber="492" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="492" type="description">
<paragraph blockId="1.[181,1142,1283,1677]" box="[613,710,1324,1350]" pageId="1" pageNumber="492">
<figureCitation box="[613,682,1324,1350]" captionStart="Fig" captionStartId="2.[276,309,721,743]" captionTargetBox="[151,1096,204,704]" captionTargetId="figure-325@2.[151,1096,204,704]" captionTargetPageId="2" captionText="Fig. 1. Sphaerowithius ansieae sp. n., holotype male: (A) dorsal; (B) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914330" httpUri="https://zenodo.org/record/7914330/files/figure.png" pageId="1" pageNumber="492">Figs 1</figureCitation>
, 
<figureCitation box="[696,710,1324,1350]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="1" pageNumber="492">2</figureCitation>
</paragraph>
<paragraph blockId="1.[181,1142,1283,1677]" box="[181,931,1360,1382]" pageId="1" pageNumber="492">
<taxonomicName authority="Beier, 1953" authorityName="Beier" authorityYear="1953" box="[181,544,1360,1382]" class="Arachnida" family="Withiidae" genus="Xenowithius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis box="[181,430,1360,1381]" italics="true" pageId="1" pageNumber="492">Xenowithius transvaalensis</emphasis>
Beier, 1953
</taxonomicName>
: 
<bibRefCitation author="MAHNERT, V." box="[553,684,1360,1382]" pageId="1" pageNumber="492" pagination="539 - 561" refId="ref3707" refString="MAHNERT, V. 1975. Pseudoskorpione der Insel Reunion und von T. F. A. I. (Djibouti). Revue Suisse de Zoologie 82: 539 - 561." type="journal article" year="1975">Mahnert 1975</bibRefCitation>
: fig. 6b (misidentification).
</paragraph>
</subSubSection>
<subSubSection box="[181,926,1384,1406]" pageId="1" pageNumber="492" type="reference_group">
<paragraph blockId="1.[181,1142,1283,1677]" box="[181,926,1384,1406]" pageId="1" pageNumber="492">
<treatmentCitationGroup box="[181,926,1384,1406]" pageId="1" pageNumber="492">
<taxonomicName authority="(Beier, 1953)" baseAuthorityName="Beier" baseAuthorityYear="1953" box="[181,573,1384,1406]" class="Arachnida" family="Withiidae" genus="Nannowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis box="[181,444,1384,1405]" italics="true" pageId="1" pageNumber="492">Nannowithius transvaalensis</emphasis>
(Beier, 1953)
</taxonomicName>
: 
<treatmentCitation author="STRINATI, P." box="[582,745,1384,1406]" page="104" pageId="1" pageNumber="492" year="1977">
<bibRefCitation author="STRINATI, P." box="[582,745,1384,1406]" pageId="1" pageNumber="492" pagination="103 - 104" refId="ref3764" refString="STRINATI, P. 1977. Recherches biospeologiques au sud-ouest Africain. In: Comunicacions del 6 e simposium d'Espeleologia (Bioespeleologia), Terrassa, desembre del 1977 / Escola Catalana d'Espeleologia, Centre Excursionista de Terrassa, Seccio d'Investigacions Subterranies. Terrassa: Centre Excursionista, DL, pp. 103 - 104." type="book chapter" year="1977">Strinati 1977: 104</bibRefCitation>
</treatmentCitation>
(misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="496" pageId="1" pageNumber="492" type="etymology">
<paragraph blockId="1.[181,1142,1283,1677]" pageId="1" pageNumber="492">Etymology: This species is named for our friend Ansie Dippenaar-Schoeman for her outstanding contributions to arachnology.</paragraph>
<paragraph blockId="1.[181,1142,1283,1677]" pageId="1" pageNumber="492">
Diagnosis: 
<taxonomicName authority="Harvey &amp; Mahnert, 2015" authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[308,574,1492,1518]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
<emphasis box="[308,574,1492,1518]" italics="true" pageId="1" pageNumber="492">Sphaerowithius ansieae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[581,641,1492,1518]" pageId="1" pageNumber="492" rank="species">sp. n.</taxonomicNameLabel>
differs from all other species of the genus in being substantially larger, e.g. the pedipalpal femur length is 
<quantity box="[883,1049,1523,1550]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" metricValueMax="1.05" metricValueMin="0.99" pageId="1" pageNumber="492" unit="mm" value="1.02" valueMax="1.05" valueMin="0.99">0.99–1.05 mm</quantity>
(males) and 1.00– 
<quantity box="[295,396,1555,1582]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="1" pageNumber="492" unit="mm" value="1.09">1.09 mm</quantity>
(females), whereas in 
<taxonomicName baseAuthorityName="Beier" baseAuthorityYear="1966" box="[655,743,1555,1581]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="vafer">
<emphasis box="[655,743,1555,1581]" italics="true" pageId="1" pageNumber="492">S. vafer</emphasis>
</taxonomicName>
it is 
<quantity box="[801,903,1555,1582]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="492" unit="mm" value="0.58">0.58 mm</quantity>
(male) and 
<quantity box="[1041,1142,1555,1582]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="492" unit="mm" value="0.62">0.62 mm</quantity>
(female), in 
<taxonomicName baseAuthorityName="Beier" baseAuthorityYear="1962" box="[316,472,1587,1613]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[316,472,1587,1613]" italics="true" pageId="1" pageNumber="492">S. basilewskyi</emphasis>
</taxonomicName>
it is 
<quantity box="[523,622,1587,1613]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="492" unit="mm" value="0.66">0.66 mm</quantity>
(male) and 
<quantity box="[752,850,1587,1613]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="492" unit="mm" value="0.71">0.71 mm</quantity>
(female), in 
<taxonomicName baseAuthorityName="Ellingsen" baseAuthorityYear="1910" box="[990,1141,1587,1613]" class="Arachnida" family="Cheliferidae" genus="Chelifer" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="perpusillus">
<emphasis box="[990,1141,1587,1613]" italics="true" pageId="1" pageNumber="492">S. perpusillus</emphasis>
</taxonomicName>
it is 
<quantity box="[238,409,1619,1645]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="1" pageNumber="492" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38–0.43 mm</quantity>
(female), in 
<taxonomicName baseAuthorityName="Beier" baseAuthorityYear="1972" box="[566,681,1619,1645]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis box="[566,681,1619,1645]" italics="true" pageId="1" pageNumber="492">S. saegeri</emphasis>
</taxonomicName>
it is 
<quantity box="[748,919,1618,1645]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.1" metricValueMin="5.5" pageId="1" pageNumber="492" unit="mm" value="0.58" valueMax="0.61" valueMin="0.55">0.55–0.61 mm</quantity>
(female) and in 
<taxonomicName baseAuthorityName="Beier" baseAuthorityYear="1966" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="492" phylum="Arthropoda" rank="species" species="salomonensis">
<emphasis italics="true" pageId="1" pageNumber="492">S. salomonensis</emphasis>
</taxonomicName>
it is 
<quantity box="[388,488,1651,1677]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="492" unit="mm" value="0.42">0.42 mm</quantity>
(male).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7914330" ID-Zenodo-Dep="7914330" box="[276,972,721,743]" httpUri="https://zenodo.org/record/7914330/files/figure.png" pageId="2" pageNumber="493" startId="2.[276,309,721,743]" targetBox="[151,1096,204,704]" targetPageId="2" targetType="figure">
<paragraph blockId="2.[276,972,721,743]" box="[276,972,721,743]" pageId="2" pageNumber="493">
Fig. 1. 
<taxonomicName authority="Harvey &amp; Mahnert, 2015" authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[340,555,721,743]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="493" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
<emphasis box="[340,555,721,743]" italics="true" pageId="2" pageNumber="493">Sphaerowithius ansieae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[561,611,722,743]" pageId="2" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
, holotype male: (A) dorsal; (B) ventral.
</paragraph>
</caption>
<paragraph blockId="2.[144,283,796,822]" box="[144,283,796,822]" pageId="2" pageNumber="493">Description:</paragraph>
<paragraph blockId="2.[144,223,839,866]" box="[144,223,839,866]" pageId="2" pageNumber="493">
<emphasis box="[144,216,839,865]" italics="true" pageId="2" pageNumber="493">Adults</emphasis>
:
</paragraph>
<paragraph blockId="2.[144,1103,882,940]" pageId="2" pageNumber="493">
<emphasis box="[144,223,882,908]" italics="true" pageId="2" pageNumber="493">Colour</emphasis>
. Pedipalps and carapace red-brown, tergites slightly paler, remainder of sclerotised portions pale yellow-brown (
<figureCitation box="[528,594,914,940]" captionStart="Fig" captionStartId="2.[276,309,721,743]" captionTargetBox="[151,1096,204,704]" captionTargetId="figure-325@2.[151,1096,204,704]" captionTargetPageId="2" captionText="Fig. 1. Sphaerowithius ansieae sp. n., holotype male: (A) dorsal; (B) ventral." figureDoi="http://doi.org/10.5281/zenodo.7914330" httpUri="https://zenodo.org/record/7914330/files/figure.png" pageId="2" pageNumber="493">Fig. 1</figureCitation>
).
</paragraph>
<paragraph blockId="2.[144,1104,957,1079]" pageId="2" pageNumber="493">
<emphasis box="[144,254,957,983]" italics="true" pageId="2" pageNumber="493">Chelicera</emphasis>
. With 5 setae on hand, 
<emphasis box="[530,555,957,983]" italics="true" pageId="2" pageNumber="493">bs</emphasis>
and 
<emphasis box="[614,650,957,983]" italics="true" pageId="2" pageNumber="493">sbs</emphasis>
slightlY dentate; movable finger with 1 subdistal seta; galea of 
<specimenCount box="[403,424,989,1015]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
and 
<specimenCount box="[477,498,989,1015]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
with 4 small terminal and 2 small medial rami; rallum of 4 blades, the most distal blade with several serrations on leading edge, other blades smooth; serrula exterior with 17 (
<specimenCount box="[519,542,1053,1079]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
, 
<specimenCount box="[554,573,1053,1079]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) blades; lamina exterior present.
</paragraph>
<paragraph blockId="2.[144,1105,1096,1602]" pageId="2" pageNumber="493">
<emphasis box="[144,244,1096,1122]" italics="true" pageId="2" pageNumber="493">Pedipalp</emphasis>
(
<figureCitation box="[257,345,1096,1122]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="2" pageNumber="493">Fig. 2D</figureCitation>
). Trochanter, femur, patella and chelal hand coarsely granulate, chelal fingers smooth; setae generallY clavate and denticulate; segments long and slender, trochanter 1.41–1.43 (
<specimenCount box="[391,414,1160,1186]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 1.96–2.05 (
<specimenCount box="[566,585,1160,1186]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
), femur 5.60–5.68 (
<specimenCount box="[812,835,1160,1186]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 5.46–5.60 (
<specimenCount box="[987,1006,1160,1186]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
), patella 4.66–4.98 (
<specimenCount box="[279,302,1192,1218]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 4.20–4.62 (
<specimenCount box="[464,483,1192,1218]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
), chela (with pedicel) 4.82–5.09 (
<specimenCount box="[883,906,1192,1218]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 4.44–4.61 (
<specimenCount box="[1068,1087,1192,1218]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
), chela (without pedicel) 4.52–4.78 (
<specimenCount box="[538,561,1224,1250]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 4.15–4.34 (
<specimenCount box="[714,733,1224,1250]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
), hand 2.35–2.45 (
<specimenCount box="[948,971,1224,1250]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 2.11–2.19 (
<specimenCount box="[154,172,1256,1282]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) × longer than broad, movable finger 0.97–1.00 (
<specimenCount box="[737,760,1256,1282]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 0.96–0.99 (
<specimenCount box="[916,934,1256,1282]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) × longer than hand (without pedicel). Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria (
<figureCitation box="[385,478,1320,1346]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="2" pageNumber="493">Fig. 2A</figureCitation>
): 
<emphasis box="[498,524,1320,1346]" italics="true" pageId="2" pageNumber="493">eb</emphasis>
and 
<emphasis box="[581,618,1320,1346]" italics="true" pageId="2" pageNumber="493">esb</emphasis>
situated basally; 
<emphasis box="[818,849,1321,1346]" italics="true" pageId="2" pageNumber="493">est</emphasis>
situated sub-medially, opposite 
<emphasis box="[244,277,1352,1378]" italics="true" pageId="2" pageNumber="493">isb</emphasis>
; 
<emphasis box="[291,311,1353,1378]" italics="true" pageId="2" pageNumber="493">et</emphasis>
situated sub-distally; 
<emphasis box="[554,576,1352,1378]" italics="true" pageId="2" pageNumber="493">ib</emphasis>
and 
<emphasis box="[627,653,1353,1378]" italics="true" pageId="2" pageNumber="493">ist</emphasis>
situated basally; 
<emphasis box="[845,878,1352,1378]" italics="true" pageId="2" pageNumber="493">isb</emphasis>
situated medially; 
<emphasis box="[1088,1104,1353,1378]" italics="true" pageId="2" pageNumber="493">it</emphasis>
situated sub-distally; 
<emphasis box="[381,395,1384,1410]" italics="true" pageId="2" pageNumber="493">b</emphasis>
and 
<emphasis box="[447,472,1384,1410]" italics="true" pageId="2" pageNumber="493">sb</emphasis>
situated near one another; 
<emphasis box="[769,788,1385,1410]" italics="true" pageId="2" pageNumber="493">st</emphasis>
substantially closer to 
<emphasis box="[1042,1050,1385,1410]" italics="true" pageId="2" pageNumber="493">t</emphasis>
than to 
<emphasis box="[173,198,1416,1442]" italics="true" pageId="2" pageNumber="493">sb</emphasis>
. Venom apparatus present in both chelal fingers, venom ducts long, terminating in nodus ramosus between 
<emphasis box="[452,472,1449,1474]" italics="true" pageId="2" pageNumber="493">et</emphasis>
and 
<emphasis box="[528,559,1449,1474]" italics="true" pageId="2" pageNumber="493">est</emphasis>
in fixed finger and distal to 
<emphasis box="[882,890,1449,1474]" italics="true" pageId="2" pageNumber="493">t</emphasis>
in movable finger. Retrolateral side of fixed finger with 11 sense-spots; retrolateral side of movable finger with 15 sense-spots; prolateral sides without sense-spots. Chelal teeth verY small; fixed finger with 59 (
<specimenCount box="[319,342,1544,1570]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 58 (
<specimenCount box="[411,429,1544,1570]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) teeth; movable finger with 65 (
<specimenCount box="[797,820,1544,1570]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 55 (
<specimenCount box="[888,906,1544,1570]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) teeth; accessorY teeth absent.
</paragraph>
<paragraph blockId="2.[144,1103,1619,1677]" lastBlockId="4.[144,1104,204,567]" lastPageId="4" lastPageNumber="495" pageId="2" pageNumber="493">
<emphasis box="[144,254,1619,1645]" italics="true" pageId="2" pageNumber="493">Carapace</emphasis>
(
<figureCitation box="[272,364,1619,1645]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="2" pageNumber="493">Fig. 2C</figureCitation>
). 1.22–1.28 (
<specimenCount box="[517,540,1619,1645]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 1.17–1.26 (
<specimenCount box="[697,715,1619,1645]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) × longer than broad; posteriorlY widened; with 2 verY small non-corneate eYe-spots; with 70 (
<specimenCount box="[811,834,1651,1677]" pageId="2" pageNumber="493" type="male">♂</specimenCount>
), 61 (
<specimenCount box="[897,915,1651,1677]" pageId="2" pageNumber="493" type="female">♀</specimenCount>
) setae, including 31 (
<specimenCount box="[187,210,204,230]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 27 (
<specimenCount box="[276,294,204,230]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
) in anterior zone (with 4 near anterior margin), 32 (
<specimenCount box="[870,893,204,230]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 28 (
<specimenCount box="[958,976,204,230]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
) in median zone, and 7 (
<specimenCount box="[283,306,235,261]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 6 (
<specimenCount box="[355,373,235,261]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
) in posterior zone; with 2 deeplY incised furrows, posterior furrow closer to posterior margin than to median furrow.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7914334" ID-Zenodo-Dep="7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="3" pageNumber="494" startId="3.[181,214,1558,1580]" targetBox="[206,1141,224,1472]" targetPageId="3" targetType="figure">
<paragraph blockId="3.[181,1143,1558,1676]" pageId="3" pageNumber="494">
Fig. 2. 
<taxonomicName authority="Harvey &amp; Mahnert, 2015" authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[244,458,1558,1580]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
<emphasis box="[244,458,1558,1580]" italics="true" pageId="3" pageNumber="494">Sphaerowithius ansieae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[463,513,1559,1580]" pageId="3" pageNumber="494" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela, dorsal, protonymph paratype; (C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme; la, lateral apodemes; pvd, postero-ventral diverticulum; vd, ventral diverticulum. Scale bars: (A, C–F) =0.5 mm, (B, G) =0.2 mm, (H, I) = 0.1 mm.
</paragraph>
</caption>
<paragraph blockId="4.[144,1104,204,567]" pageId="4" pageNumber="495">
<emphasis box="[144,289,309,335]" italics="true" pageId="4" pageNumber="495">Coxal region</emphasis>
. Maxilla with 2 apical setae and 1 very small internal, sub-oral seta, plus 20 (
<specimenCount box="[188,211,341,367]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 25 (
<specimenCount box="[279,297,341,367]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
) other setae; maxilla without rugose area; chaetotaxY of coxae I–IV: 
<specimenCount box="[1076,1099,341,367]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
, 10: 10: 14: 23, 
<specimenCount box="[314,335,373,399]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
, 8: 9: 10: 20.
</paragraph>
<paragraph blockId="4.[144,1104,204,567]" pageId="4" pageNumber="495">
<emphasis box="[144,197,415,440]" italics="true" pageId="4" pageNumber="495">Legs</emphasis>
(
<figureCitation box="[210,327,414,440]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="4" pageNumber="495">Figs 2E, F</figureCitation>
). Long and slender; junction between femora and patellae I and II only slightlY oblique; femur + patella of leg IV 4.25 (
<specimenCount box="[680,703,446,472]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 4.56 (
<specimenCount box="[790,808,446,472]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
) × longer than broad; legs III and IV with slightly shortened, acuminate, tarsal tactile seta, situated sub-distally, TS = 0.78 (
<specimenCount box="[277,300,510,536]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
), 0.75 (
<specimenCount box="[392,411,510,536]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
); subterminal tarsal setae simple, arcuate and acute; arolium slightly shorter than claws.
</paragraph>
<paragraph blockId="4.[144,1105,583,895]" pageId="4" pageNumber="495">
<emphasis box="[144,248,583,609]" italics="true" pageId="4" pageNumber="495">Abdomen</emphasis>
. Tergites I–X and sternites VI–VIII with faint medial suture. Tergal chaetotaxy: 
<specimenCount box="[144,167,615,641]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
, 6: 8: 9: 12: 11: 13: 16: 16: 15: 16: 10 (including 2 tactile setae): 2; 
<specimenCount box="[942,963,615,641]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
, 6: 6: 9: 10: 10: 12: 13: 14: 14: 14: 8: 2; mostly uniseriate, but some tergites with a few setae placed anteriorlY; all setae foliate. Sternal chaetotaxY: 
<specimenCount box="[675,698,679,705]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
, 8: (2) 11 (2): (1) 12 [0] (1): 18: 16: 16: 15 + 
<emphasis box="[242,276,711,736]" italics="true" pageId="4" pageNumber="495">ca.</emphasis>
26 
<emphasis box="[316,349,710,736]" italics="true" pageId="4" pageNumber="495">gls</emphasis>
: 14 + 
<emphasis box="[420,453,711,736]" italics="true" pageId="4" pageNumber="495">ca.</emphasis>
20 
<emphasis box="[494,527,710,736]" italics="true" pageId="4" pageNumber="495">gls</emphasis>
: 12: 7 (including 2 tactile setae): 2; 
<specimenCount box="[927,948,711,737]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
, 14: (2) 9 (2): (1) 6 (1): 16: 15: 13: 12: 12: 10: 6 (including 2 tactile setae): 2; 
<specimenCount box="[838,859,742,768]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
without internal setae in the genital atrium; sternites VIII–IX of 
<specimenCount box="[607,628,774,800]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
with small patches of glandular setae (
<figureCitation captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="4" pageNumber="495">Fig. 2G</figureCitation>
), set in a depression that extends posteriorly over the segmental membrane, glandular setae small and conical; sternites of 
<specimenCount box="[539,560,838,864]" pageId="4" pageNumber="495" type="female">♀</specimenCount>
without glandular setae; all other setae uniseriate and acuminate; 
<specimenCount box="[321,342,869,895]" pageId="4" pageNumber="495" type="male">♂</specimenCount>
without paired invaginations on anterior margins of sternites.
</paragraph>
<paragraph blockId="4.[144,1103,911,1032]" pageId="4" pageNumber="495">
<emphasis box="[144,253,911,937]" italics="true" pageId="4" pageNumber="495">Genitalia</emphasis>
. Male with rounded dorsal apodemes, lateral apodemes not elongated, postero-ventral diverticulum (?) with small lobes (
<figureCitation box="[723,816,942,969]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="4" pageNumber="495">Fig. 2H</figureCitation>
); female with 2 spheroid spermathecal receptacula, with 1 ovoid median cribriform plate and a pair of curved lateral apodemes (
<figureCitation box="[348,427,1006,1032]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="4" pageNumber="495">Fig. 2I</figureCitation>
).
</paragraph>
<paragraph blockId="4.[144,1104,1048,1296]" pageId="4" pageNumber="495">
<emphasis box="[144,400,1048,1074]" italics="true" pageId="4" pageNumber="495">Male dimensions (mm)</emphasis>
. 
<typeStatus box="[414,521,1048,1074]" pageId="4" pageNumber="495">Holotype</typeStatus>
, followed by other male 
<typeStatus box="[799,895,1048,1074]" pageId="4" pageNumber="495">paratype</typeStatus>
(when measured): Body length 2.38 (2.43). Pedipalps: trochanter 
<geoCoordinate box="[710,774,1079,1105]" degrees="0.468" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.468">0.468</geoCoordinate>
/ 
<geoCoordinate box="[782,847,1079,1105]" degrees="0.242" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.242">0.242</geoCoordinate>
(
<geoCoordinate box="[867,934,1079,1106]" degrees="0.435" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.435">0.435</geoCoordinate>
/ 
<geoCoordinate box="[943,1011,1079,1106]" degrees="0.228" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.228">0.228</geoCoordinate>
), femur 
<geoCoordinate box="[144,206,1111,1138]" degrees="1.045" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="1.045">1.045</geoCoordinate>
/ 
<geoCoordinate box="[214,279,1111,1138]" degrees="0.184" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.184">0.184</geoCoordinate>
(
<geoCoordinate box="[298,365,1111,1137]" degrees="0.992" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.992">0.992</geoCoordinate>
/ 
<geoCoordinate box="[374,441,1111,1137]" degrees="0.184" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.184">0.184</geoCoordinate>
), patella 
<geoCoordinate box="[550,614,1111,1137]" degrees="0.899" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.899">0.899</geoCoordinate>
/ 
<geoCoordinate box="[623,686,1111,1137]" degrees="0.193" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.193">0.193</geoCoordinate>
(
<geoCoordinate box="[705,771,1111,1137]" degrees="0.877" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.877">0.877</geoCoordinate>
/ 
<geoCoordinate box="[780,847,1111,1137]" degrees="0.176" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.176">0.176</geoCoordinate>
), chela (with pedicel) 
<geoCoordinate box="[144,206,1143,1169]" degrees="1.486" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="1.486">1.486</geoCoordinate>
/ 
<geoCoordinate box="[214,278,1143,1169]" degrees="0.308" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.308">0.308</geoCoordinate>
(
<geoCoordinate box="[294,358,1143,1169]" degrees="1.424" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="1.424">1.424</geoCoordinate>
/ 
<geoCoordinate box="[367,434,1143,1169]" degrees="0.280" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.28">0.280</geoCoordinate>
), chela (without pedicel) 1.392 (1.338), hand length 0.725 (0.685), movable finger length 0.704 (0.686). Chelicera 
<geoCoordinate box="[784,848,1175,1201]" degrees="0.242" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.242">0.242</geoCoordinate>
/ 
<geoCoordinate box="[856,920,1175,1201]" degrees="0.106" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.106">0.106</geoCoordinate>
, movable finger length 0.173. Carapace 
<geoCoordinate box="[414,477,1206,1232]" degrees="0.888" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.888">0.888</geoCoordinate>
/ 
<geoCoordinate box="[485,547,1206,1232]" degrees="0.693" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.693">0.693</geoCoordinate>
(
<geoCoordinate box="[564,629,1206,1233]" degrees="0.851" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.851">0.851</geoCoordinate>
/ 
<geoCoordinate box="[638,704,1206,1233]" degrees="0.698" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.698">0.698</geoCoordinate>
); eye diameter 0.039. Leg I: femur 
<geoCoordinate box="[144,208,1238,1264]" degrees="0.200" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.2">0.200</geoCoordinate>
/ 
<geoCoordinate box="[216,280,1238,1264]" degrees="0.140" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.14">0.140</geoCoordinate>
, patella 
<geoCoordinate box="[373,437,1238,1264]" degrees="0.378" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.378">0.378</geoCoordinate>
/ 
<geoCoordinate box="[445,509,1238,1264]" degrees="0.130" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.13">0.130</geoCoordinate>
, tibia 
<geoCoordinate box="[577,641,1238,1265]" degrees="0.452" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.452">0.452</geoCoordinate>
/ 
<geoCoordinate box="[650,714,1238,1265]" degrees="0.079" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.079">0.079</geoCoordinate>
, tarsus 
<geoCoordinate box="[797,862,1238,1265]" degrees="0.410" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.41">0.410</geoCoordinate>
/ 
<geoCoordinate box="[870,934,1238,1265]" degrees="0.058" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.058">0.058</geoCoordinate>
. Leg IV: femur + patella 
<geoCoordinate box="[248,312,1270,1296]" degrees="0.722" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.722">0.722</geoCoordinate>
/ 
<geoCoordinate box="[321,385,1270,1296]" degrees="0.170" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.17">0.170</geoCoordinate>
, tibia 
<geoCoordinate box="[453,517,1270,1296]" degrees="0.630" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.63">0.630</geoCoordinate>
/ 
<geoCoordinate box="[525,589,1270,1296]" degrees="0.090" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.09">0.090</geoCoordinate>
, tarsus 
<geoCoordinate box="[673,737,1270,1296]" degrees="0.463" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.463">0.463</geoCoordinate>
/ 
<geoCoordinate box="[745,809,1270,1296]" degrees="0.068" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.068">0.068</geoCoordinate>
, TS length 0.360.
</paragraph>
<paragraph blockId="4.[144,1104,1312,1677]" pageId="4" pageNumber="495">
<emphasis box="[144,427,1312,1338]" italics="true" pageId="4" pageNumber="495">Female dimensions (mm)</emphasis>
. 
<typeStatus box="[442,543,1312,1338]" pageId="4" pageNumber="495">Paratype</typeStatus>
, followed by other 
<typeStatus box="[761,857,1312,1338]" pageId="4" pageNumber="495">paratype</typeStatus>
females (when measured): Body length 2.38 (2.51–3.22). Pedipalps: trochanter 
<geoCoordinate box="[803,866,1343,1370]" degrees="0.435" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.435">0.435</geoCoordinate>
/ 
<geoCoordinate box="[874,937,1343,1370]" degrees="0.218" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.218">0.218</geoCoordinate>
(0.448–0.481/ 0.219–0.246),femur 
<geoCoordinate box="[370,431,1375,1401]" degrees="0.996" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.996">0.996</geoCoordinate>
/ 
<geoCoordinate box="[439,499,1375,1401]" degrees="0.178" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.178">0.178</geoCoordinate>
(1.005–1.091/0.184–0.198), patella 
<geoCoordinate box="[887,947,1375,1402]" degrees="0.947" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.947">0.947</geoCoordinate>
/ 
<geoCoordinate box="[955,1015,1375,1402]" degrees="0.205" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.205">0.205</geoCoordinate>
(0.886– 0.982/0.211–0.231), chela (with pedicel) 
<geoCoordinate box="[592,652,1407,1433]" degrees="1.446" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="1.446">1.446</geoCoordinate>
/ 
<geoCoordinate box="[660,723,1407,1433]" degrees="0.314" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.314">0.314</geoCoordinate>
(1.488–1.586/0.328–0.357), chela (without pedicel) 1.363 (
<geoCoordinate box="[417,481,1438,1465]" degrees="1.405" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="1.405">1.405</geoCoordinate>
<geoCoordinate box="[483,565,1439,1465]" degrees="1.482" direction="west" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="-1.482">–1.482</geoCoordinate>
), hand length 0.688 (
<geoCoordinate box="[795,861,1439,1465]" degrees="0.694" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.694">0.694</geoCoordinate>
<geoCoordinate box="[861,944,1439,1465]" degrees="0.770" direction="west" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="-0.77">–0.770</geoCoordinate>
), movable finger length 0.668 (
<geoCoordinate box="[345,410,1470,1497]" degrees="0.685" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.685">0.685</geoCoordinate>
<geoCoordinate box="[410,495,1470,1497]" degrees="0.736" direction="west" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="-0.736">–0.736</geoCoordinate>
). Chelicera 
<geoCoordinate box="[630,693,1470,1497]" degrees="0.255" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.255">0.255</geoCoordinate>
/ 
<geoCoordinate box="[701,765,1470,1497]" degrees="0.110" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.11">0.110</geoCoordinate>
, movable finger length 0.177. Carapace 
<geoCoordinate box="[256,319,1502,1528]" degrees="0.848" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.848">0.848</geoCoordinate>
/ 
<geoCoordinate box="[327,390,1502,1528]" degrees="0.688" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.688">0.688</geoCoordinate>
(0.880–0.904/0.718–0.752); eye diameter 0.026. Leg I: femur 
<geoCoordinate box="[144,208,1534,1561]" degrees="0.187" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.187">0.187</geoCoordinate>
/ 
<geoCoordinate box="[216,280,1534,1561]" degrees="0.125" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.125">0.125</geoCoordinate>
, patella 
<geoCoordinate box="[373,436,1534,1560]" degrees="0.403" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.403">0.403</geoCoordinate>
/ 
<geoCoordinate box="[445,510,1534,1560]" degrees="0.074" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.074">0.074</geoCoordinate>
, tibia 
<geoCoordinate box="[577,642,1534,1560]" degrees="0.414" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.414">0.414</geoCoordinate>
/ 
<geoCoordinate box="[650,713,1534,1560]" degrees="0.073" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.073">0.073</geoCoordinate>
, tarsus 
<geoCoordinate box="[797,862,1534,1561]" degrees="0.389" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.389">0.389</geoCoordinate>
/ 
<geoCoordinate box="[870,934,1534,1561]" degrees="0.057" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.057">0.057</geoCoordinate>
. Leg IV: femur + patella 
<geoCoordinate box="[248,312,1566,1593]" degrees="0.725" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.725">0.725</geoCoordinate>
/ 
<geoCoordinate box="[320,384,1566,1593]" degrees="0.159" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.159">0.159</geoCoordinate>
, tibia 
<geoCoordinate box="[453,517,1566,1592]" degrees="0.640" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.64">0.640</geoCoordinate>
/ 
<geoCoordinate box="[526,590,1566,1592]" degrees="0.088" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.088">0.088</geoCoordinate>
, tarsus 
<geoCoordinate box="[673,736,1566,1592]" degrees="0.481" direction="north" orientation="latitude" pageId="4" pageNumber="495" precision="55" value="0.481">0.481</geoCoordinate>
/ 
<geoCoordinate box="[745,810,1566,1592]" degrees="0.067" direction="east" orientation="longitude" pageId="4" pageNumber="495" precision="55" value="0.067">0.067</geoCoordinate>
, TS length 0.360.
</paragraph>
<paragraph blockId="4.[144,1104,1312,1677]" box="[144,289,1608,1635]" pageId="4" pageNumber="495">
<emphasis box="[144,281,1608,1634]" italics="true" pageId="4" pageNumber="495">Protonymph</emphasis>
:
</paragraph>
<paragraph blockId="4.[144,1104,1312,1677]" lastBlockId="5.[181,1142,204,1076]" lastPageId="5" lastPageNumber="496" pageId="4" pageNumber="495">
<emphasis box="[144,228,1650,1676]" italics="true" pageId="4" pageNumber="495">Colour.</emphasis>
Mostly very pale yellow-brown, with pedipalps and carapace pale red-brown. 
<emphasis box="[181,291,204,230]" italics="true" pageId="5" pageNumber="496">Chelicera</emphasis>
. With 4 setae on hand and no seta on movable finger; rallum with 4 blades. 
<emphasis box="[181,282,242,268]" italics="true" pageId="5" pageNumber="496">Pedipalp</emphasis>
. Trochanter 1.78, femur 2.97, patella 2.37, chela (with pedicel) 3.60, chela (without pedicel) 3.42, hand 1.76 × longer than broad, movable finger 0.96 × longer than hand (without pedicel). Fixed chelal finger with 3 trichobothria, movable chelal finger with 1 trichobothrium (
<figureCitation box="[515,604,338,364]" captionStart="Fig" captionStartId="3.[181,214,1558,1580]" captionTargetBox="[206,1141,224,1472]" captionTargetId="figure-87@3.[181,1141,215,1515]" captionTargetPageId="3" captionText="Fig.2. Sphaerowithius ansieae sp. n., holotype male, unless stated otherwise: (A) left chela, lateral; (B) left chela,dorsal, protonymph paratype;(C) carapace, dorsal; (D) right pedipalp, dorsal; (E) leg I; (F) leg IV; (G) sternites VIII and IX, ventral; (H) genitalia, ventral; (I) genitalia, ventral, female paratype. Abbreviations:da, dorsal apodeme;la, lateral apodemes;pvd,postero-ventral diverticulum;vd,ventral diverticulum. Scale bars: (A, C–F)=0.5 mm, (B, G)=0.2 mm, (H, I)=0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7914334" httpUri="https://zenodo.org/record/7914334/files/figure.png" pageId="5" pageNumber="496">Fig. 2B</figureCitation>
): onlY 
<emphasis box="[681,707,338,364]" italics="true" pageId="5" pageNumber="496">eb</emphasis>
, 
<emphasis box="[721,741,339,364]" italics="true" pageId="5" pageNumber="496">et</emphasis>
, 
<emphasis box="[755,781,339,364]" italics="true" pageId="5" pageNumber="496">ist</emphasis>
and 
<emphasis box="[836,844,339,364]" italics="true" pageId="5" pageNumber="496">t</emphasis>
present.
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
<emphasis box="[181,290,377,403]" italics="true" pageId="5" pageNumber="496">Carapace</emphasis>
. 1.30 × longer than broad; eyes absent; with 16 setae, including 4 near anterior margin and 4 near posterior margin; with 1 deep, median furrow.
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
<emphasis box="[181,287,448,474]" italics="true" pageId="5" pageNumber="496">Abdomen</emphasis>
. Tergal chaetotaxy: 2: 2: 4: 4: 4: 4: 4: 4: 4: 4: 2: 2. Sternal chaetotaxy: 0: (0) 2 (0): (1) 2 (1): 4: 4: 4: 4: 4: 4: 4: 2; glandular setae absent.
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
<emphasis box="[181,380,519,545]" italics="true" pageId="5" pageNumber="496">Dimensions (mm)</emphasis>
. Body length 1.24. Pedipalps: trochanter 
<geoCoordinate box="[848,912,518,544]" degrees="0.173" direction="north" orientation="latitude" pageId="5" pageNumber="496" precision="55" value="0.173">0.173</geoCoordinate>
/ 
<geoCoordinate box="[921,985,518,544]" degrees="0.097" direction="east" orientation="longitude" pageId="5" pageNumber="496" precision="55" value="0.097">0.097</geoCoordinate>
, femur 
<geoCoordinate box="[1071,1133,518,544]" degrees="0.288" direction="north" orientation="latitude" pageId="5" pageNumber="496" precision="55" value="0.288">0.288</geoCoordinate>
/ 
<geoCoordinate box="[181,247,550,576]" degrees="0.097" direction="east" orientation="longitude" pageId="5" pageNumber="496" precision="55" value="0.097">0.097</geoCoordinate>
, patella 
<geoCoordinate box="[344,408,550,577]" degrees="0.258" direction="north" orientation="latitude" pageId="5" pageNumber="496" precision="55" value="0.258">0.258</geoCoordinate>
/ 
<geoCoordinate box="[417,481,550,577]" degrees="0.109" direction="east" orientation="longitude" pageId="5" pageNumber="496" precision="55" value="0.109">0.109</geoCoordinate>
, chela (with pedicel) 
<geoCoordinate box="[729,793,550,577]" degrees="0.525" direction="north" orientation="latitude" pageId="5" pageNumber="496" precision="55" value="0.525">0.525</geoCoordinate>
/ 
<geoCoordinate box="[802,866,550,577]" degrees="0.146" direction="east" orientation="longitude" pageId="5" pageNumber="496" precision="55" value="0.146">0.146</geoCoordinate>
, chela (without pedicel) 0.499, hand length 0.257, movable finger length 0.248. Carapace 
<geoCoordinate box="[915,979,582,609]" degrees="0.506" direction="north" orientation="latitude" pageId="5" pageNumber="496" precision="55" value="0.506">0.506</geoCoordinate>
/ 
<geoCoordinate box="[988,1053,582,609]" degrees="0.390" direction="east" orientation="longitude" pageId="5" pageNumber="496" precision="55" value="0.39">0.390</geoCoordinate>
.
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
<materialsCitation collectingDate="1972-05-02" collectionCode="MHNG" collectorName="P. von Wrede &amp; P. Strinati" country="Namibia" latitude="-19.65" location="Otavi" longLatPrecision="1271" longitude="17.333334" municipality="Nosibhohle" pageId="5" pageNumber="496" specimenCount="1" specimenCount-male="1" stateProvince="Otjozondjupa" typeStatus="holotype">
<typeStatus box="[181,269,617,639]" pageId="5" pageNumber="496">Holotype</typeStatus>
: 
<collectingCountry box="[279,382,617,638]" name="Namibia" pageId="5" pageNumber="496">NAMIBIA</collectingCountry>
: 
<emphasis box="[392,522,617,638]" italics="true" pageId="5" pageNumber="496">
<collectingRegion box="[392,517,617,638]" country="Namibia" name="Otjozondjupa" pageId="5" pageNumber="496">Otjozondjupa</collectingRegion>
:
</emphasis>
<specimenCount box="[529,546,617,638]" pageId="5" pageNumber="496" type="male">♂</specimenCount>
<collectingMunicipality box="[553,660,617,639]" pageId="5" pageNumber="496">Nosibhöhle</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03DD87C8FFFFFFD03488AD798BB9DD58:8EAB6005FFFBFFD037FDAA138899DB13" box="[670,720,617,639]" country="Namibia" latitude="-19.65" longLatPrecision="1271" longitude="17.333334" municipality="Nosibhohle" name="Otavi" pageId="5" pageNumber="496" stateProvince="Otjozondjupa">Otavi</location>
(
<geoCoordinate box="[734,805,617,638]" degrees="19" direction="south" minutes="39" orientation="latitude" pageId="5" pageNumber="496" precision="925" value="-19.65">19°39'S</geoCoordinate>
<geoCoordinate box="[812,887,617,638]" degrees="17" direction="east" minutes="20" orientation="longitude" pageId="5" pageNumber="496" precision="925" value="17.333334">17°20'E</geoCoordinate>
), 
<date box="[904,985,617,638]" pageId="5" pageNumber="496" value="1972-05-02">
<collectingDate box="[904,985,617,638]" pageId="5" pageNumber="496" value="1972-05-02">2.v.1972</collectingDate>
</date>
, 
<collectorName box="[995,1117,617,639]" pageId="5" pageNumber="496">P. von Wrede</collectorName>
&amp; 
<collectorName box="[181,270,641,663]" pageId="5" pageNumber="496">P. Strinati</collectorName>
(
<collectionCode box="[282,357,641,662]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="496" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
<materialsCitation box="[181,997,671,693]" collectingDate="1972-05-02" collectionCode="MHNG" collectorName="P. von Wrede &amp; P. Strinati" country="Namibia" latitude="-19.65" location="Otavi" longLatPrecision="1271" longitude="17.333334" municipality="Nosibhohle" pageId="5" pageNumber="496" specimenCount="3" specimenCount-female="3" stateProvince="Otjozondjupa" typeStatus="paratype">
<typeStatus box="[181,272,672,693]" pageId="5" pageNumber="496">Paratypes</typeStatus>
: 
<collectingCountry box="[280,383,671,692]" name="Namibia" pageId="5" pageNumber="496">NAMIBIA</collectingCountry>
: 
<emphasis box="[389,520,671,692]" italics="true" pageId="5" pageNumber="496">
<collectingRegion box="[389,514,671,692]" country="Namibia" name="Otjozondjupa" pageId="5" pageNumber="496">Otjozondjupa</collectingRegion>
:
</emphasis>
<specimenCount box="[524,552,671,692]" pageId="5" pageNumber="496" type="female">3♀</specimenCount>
, 1 protonYmph, collected with holotYpe (
<collectionCode box="[916,990,671,692]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="496" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1972-04-30" collectionCode="MHNG" collectorName="P. von Wrede &amp; P. Strinati" country="Namibia" latitude="-19.65" longLatPrecision="1271" longitude="17.333334" pageId="5" pageNumber="496" specimenCount="1" specimenCount-male="1" stateProvince="Otjozondjupa" typeStatus="paratype">
<specimenCount box="[1005,1032,671,692]" pageId="5" pageNumber="496" type="male">1♂</specimenCount>
Albathöhle, Otavi (
<geoCoordinate box="[245,316,695,716]" degrees="19" direction="south" minutes="39" orientation="latitude" pageId="5" pageNumber="496" precision="925" value="-19.65">19°39'S</geoCoordinate>
<geoCoordinate box="[322,397,695,716]" degrees="17" direction="east" minutes="20" orientation="longitude" pageId="5" pageNumber="496" precision="925" value="17.333334">17°20'E</geoCoordinate>
), 
<date box="[413,511,695,717]" pageId="5" pageNumber="496" value="1972-04-30">
<collectingDate box="[413,511,695,717]" pageId="5" pageNumber="496" value="1972-04-30">30.iv.1972</collectingDate>
</date>
, 
<collectorName box="[520,641,695,717]" pageId="5" pageNumber="496">P. von Wrede</collectorName>
&amp; 
<collectorName box="[669,757,695,717]" pageId="5" pageNumber="496">P. Strinati</collectorName>
(
<collectionCode box="[769,844,695,716]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="496" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="5.[181,1142,204,1076]" pageId="5" pageNumber="496">
Remarks: 
<taxonomicName authority="Harvey &amp; Mahnert, 2015" authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[298,566,730,756]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
<emphasis box="[298,566,730,756]" italics="true" pageId="5" pageNumber="496">Sphaerowithius ansieae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[575,637,731,757]" pageId="5" pageNumber="496" rank="species">sp. n.</taxonomicNameLabel>
is known only from two caves in the Otavi region of northern 
<collectingCountry box="[407,508,762,788]" name="Namibia" pageId="5" pageNumber="496">Namibia</collectingCountry>
, from Nosibhöhle and Albathöhle, and the specimens were first recorded bY 
<bibRefCitation author="STRINATI, P." box="[448,618,794,820]" pageId="5" pageNumber="496" pagination="103 - 104" refId="ref3764" refString="STRINATI, P. 1977. Recherches biospeologiques au sud-ouest Africain. In: Comunicacions del 6 e simposium d'Espeleologia (Bioespeleologia), Terrassa, desembre del 1977 / Escola Catalana d'Espeleologia, Centre Excursionista de Terrassa, Seccio d'Investigacions Subterranies. Terrassa: Centre Excursionista, DL, pp. 103 - 104." type="book chapter" year="1977">Strinati (1977)</bibRefCitation>
. He also mentioned a specimen from nearbY Ghaubhöhle, but this specimen has not been available for studY. These three caves, along with Aigamashöhle, were described bY 
<bibRefCitation author="STRINATI, P." box="[713,883,858,884]" pageId="5" pageNumber="496" pagination="103 - 104" refId="ref3764" refString="STRINATI, P. 1977. Recherches biospeologiques au sud-ouest Africain. In: Comunicacions del 6 e simposium d'Espeleologia (Bioespeleologia), Terrassa, desembre del 1977 / Escola Catalana d'Espeleologia, Centre Excursionista de Terrassa, Seccio d'Investigacions Subterranies. Terrassa: Centre Excursionista, DL, pp. 103 - 104." type="book chapter" year="1977">Strinati (1977)</bibRefCitation>
, who also listed other taxa collected in the caves including beetles, hemipterans, orthopterans, flies, isopods, spiders, mites, harvestmen, earthworms, and molluscs, as well as the pseudoscorpion 
<taxonomicName authority="Beier, 1970" authorityName="Beier" authorityYear="1970" box="[181,558,954,980]" class="Arachnida" family="Cheiridiidae" genus="Cheiridium" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="capense">
<emphasis box="[181,411,954,980]" italics="true" pageId="5" pageNumber="496">Cheiridium capense</emphasis>
Beier, 1970
</taxonomicName>
, which was previously known only from the 
<typeStatus box="[1093,1141,955,981]" pageId="5" pageNumber="496">type</typeStatus>
locality in 
<collectingCountry box="[304,448,986,1012]" name="South Africa" pageId="5" pageNumber="496">South Africa</collectingCountry>
(Beier 1970). The specimens of 
<taxonomicName authorityName="Harvey &amp; Mahnert" authorityYear="2015" box="[822,937,987,1013]" class="Arachnida" family="Withiidae" genus="Sphaerowithius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="ansieae">
<emphasis box="[822,937,987,1013]" italics="true" pageId="5" pageNumber="496">S. ansieae</emphasis>
</taxonomicName>
show some slight modifications to an obligate cavernicolous existence including extremelY small eYes and long, slender pedipalps.
</paragraph>
</subSubSection>
</treatment>
</document>