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<document id="D5A97808B2C395A5A4E90DA1DEBD8A5A" ID-CLB-Dataset="24558" ID-DOI="10.11646/zootaxa.4712.1.2" ID-GBIF-Dataset="19829187-393b-46c5-8e83-c132a5742fb3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586816" ID-ZooBank="6C18DE3D-8B51-473D-B528-98337BF5730E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824202715" checkinUser="plazi" docAuthor="Perreault, Ray T. &amp; Buckeridge, John S." docDate="2019" docId="03F87725FFAC89025CA9FAAB9359FC61" docLanguage="en" docName="zootaxa.4712.1.2.pdf" docOrigin="Zootaxa 4712 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Verruca gailgoedertae Perreault &amp; Buckeridge 2019, sp. nov." docType="treatment" docUuid="D313492D-00E1-4A32-B048-4E7327A3224A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="50" masterDocId="FFC10F5DFFA889045C3EFF909770FFE2" masterDocTitle="Paleogene Verrucidae (Cirripedia: Verrucomorpha) of North America, with descriptions of three new species" masterLastPageNumber="50" masterPageNumber="34" pageNumber="34" updateTime="1698775395024" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="772F69C08CDEED124FDA202C12795CFE">Paleogene Verrucidae (Cirripedia: Verrucomorpha) of North America, with descriptions of three new species</mods:title>
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<mods:namePart id="70B3492031AC3DA71B760E76A2009AF9">Perreault, Ray T.</mods:namePart>
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<mods:namePart id="48AD89AAC98CAFD66EC2951BA4F59353">Buckeridge, John S.</mods:namePart>
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<mods:title id="C493ACCD865B02B341CAA8D851D5FFF1">Zootaxa</mods:title>
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<mods:date id="3246C8E46DD9CF40954FF61835E93B99">2019</mods:date>
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<mods:identifier id="98EE34EA7EA2AF85A9BA139E8235466C" type="DOI">10.11646/zootaxa.4712.1.2</mods:identifier>
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<mods:identifier id="9DAFF44BA2BCF44C5EA08F835EC1A81B" type="Zenodo-Dep">3586816</mods:identifier>
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<treatment id="03F87725FFAC89025CA9FAAB9359FC61" ID-DOI="http://doi.org/10.5281/zenodo.5940769" ID-GBIF-Taxon="160797954" ID-Zenodo-Dep="5940769" LSID="urn:lsid:zoobank.org:act:D313492D-00E1-4A32-B048-4E7327A3224A" httpUri="http://treatment.plazi.org/id/03F87725FFAC89025CA9FAAB9359FC61" lastPageId="6" pageId="4">
<subSubSection id="C34B95B8FFAC89005CA9FAAB944BFAB7" box="[151,827,1338,1365]" pageId="4" type="nomenclature">
<paragraph id="8BEEC633FFAC89005CA9FAAB944BFAB7" blockId="4.[151,959,1338,1436]" box="[151,827,1338,1365]" pageId="4">
<heading id="D0A6715FFFAC89005CA9FAAB944BFAB7" bold="true" box="[151,827,1338,1365]" fontSize="11" level="1" pageId="4" reason="1">
<emphasis id="B9251A21FFAC89005CA9FAAB944BFAB7" bold="true" box="[151,827,1338,1365]" pageId="4">
<taxonomicName id="4C51BDB0FFAC89005CA9FAAB95A5FAB6" authority="Perreault &amp; Buckeridge" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[151,725,1338,1365]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="4" phylum="Arthropoda" rank="species" species="gailgoedertae" status="sp. nov.">
<emphasis id="B9251A21FFAC89005CA9FAAB96D2FAB7" bold="true" box="[151,418,1338,1365]" italics="true" pageId="4">Verruca gailgoedertae</emphasis>
Perreault &amp; Buckeridge
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFAC89005EE3FAAB944BFAB7" box="[733,827,1339,1365]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAC89005CA9FACE94CFFA7E" pageId="4" type="description">
<paragraph id="8BEEC633FFAC89005CA9FACE9628FA9B" blockId="4.[151,959,1338,1436]" box="[151,344,1374,1401]" pageId="4">
(
<figureCitation id="136ADAB6FFAC89005CA1FACE979EFA9B" box="[159,238,1374,1401]" captionStart="PLATE 1" captionStartId="5.[151,234,1141,1166]" captionTargetBox="[185,1374,182,1097]" captionTargetId="figure@5.[160,1427,179,1114]" captionTargetPageId="5" captionText="PLATE 1. Verruca gailgoedertae, sp. nov. a. Paratype NMV P332716, carina (exterior); b. paratype NMV P332716, carina (interior); c. paratype NMV P332719, rostrum (exterior); d. paratype NMV P332719, rostrum (interior); e. paratype NMV P332718, fixed scutum (exterior); f. paratype NMV P332718, fixed scutum (interior); g. holotype LACMIP 14759, movable tergum (exterior); h. holotype LACMIP 14759, movable tergum (interior); i. paratype NMV P332717, fixed tergum (exterior); j. paratype NMV P332717, fixed tergum (interior); k. paratype, fixed scutum (exterior); l. paratype fixed scutum (interior). All specimens are from the Lower Middle Eocene Crescent-McIntosh transitional zone, Location (CSUN 1573), Doty Hills, Washington, USA." figureDoi="http://doi.org/10.5281/zenodo.3586818" httpUri="https://zenodo.org/record/3586818/files/figure.png" pageId="4">Plate 1</figureCitation>
, figs al)
</paragraph>
<paragraph id="8BEEC633FFAC89005CA9FA1294CFFA7E" blockId="4.[151,959,1338,1436]" box="[151,959,1410,1436]" pageId="4">
<uri id="FFC0CA31FFAC89005CA9FA1294CFFA7E" box="[151,959,1410,1436]" pageId="4">
urn:lsid:zoobank.org:act:
<uuid id="FFF7FCE6FFAC89005D97FA1294CFFA7E" box="[425,959,1410,1436]" pageId="4">D313492D-00E1-4A32-B048-4E7327A3224A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAC89005CA9FA54923DF9C5" pageId="4" type="diagnosis">
<paragraph id="8BEEC633FFAC89005CA9FA54923DF9C5" blockId="4.[151,1437,1476,2040]" pageId="4">
<emphasis id="B9251A21FFAC89005CA9FA54967FFA3D" bold="true" box="[151,271,1476,1503]" pageId="4">Diagnosis.</emphasis>
Shell with moderately rugose, irregular external ribbing crossed by fine growth lines; all plates perforated by fine punctae; first and second articulating ribs on movable tergum similar in width; rostrum and carina articulating with up to five ribs; fixed scutal radio-alar wing extending out a distance equal to basal width of fixed scutum.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAC89005CF9F9A0945CF91C" pageId="4" type="materials_examined">
<paragraph id="8BEEC633FFAC89005CF9F9A09605F9A8" blockId="4.[151,1437,1476,2040]" box="[199,373,1583,1610]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F9A09605F9A8" bold="true" box="[199,373,1583,1610]" pageId="4">Type material.</emphasis>
</paragraph>
<paragraph id="8BEEC633FFAC89005CF9F9C494FBF98D" blockId="4.[151,1437,1476,2040]" box="[199,907,1619,1647]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F9C4964BF98C" bold="true" box="[199,315,1620,1646]" pageId="4">
<typeStatus id="54EA7891FFAC89005CF9F9C49646F98C" box="[199,310,1620,1646]" pageId="4" type="holotype">Holotype</typeStatus>
.
</emphasis>
LACMIP 14759, a movable tergum (pl. 1, figs g, h).
</paragraph>
<paragraph id="8BEEC633FFAC89005CF9F9E89338F954" blockId="4.[151,1437,1476,2040]" pageId="4">
<materialsCitation id="3B39CC6EFFAC89005CF9F9E89456F970" ID-GBIF-Occurrence="2521263194" box="[199,806,1655,1682]" collectionCode="NMV" pageId="4" specimenCode="P332716" specimenCount="1" typeStatus="paratype">
<emphasis id="B9251A21FFAC89005CF9F9E89636F970" bold="true" box="[199,326,1656,1682]" pageId="4">
<typeStatus id="54EA7891FFAC89005CF9F9E89631F970" box="[199,321,1656,1682]" pageId="4" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED405EF6FFAC89005D71F9E896E0F970" box="[335,400,1656,1682]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">NMV</collectionCode>
<specimenCode id="DBF76E48FFAC89005DA7F9E7968FF973" box="[409,511,1655,1681]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">P332716</specimenCode>
, a carina (pl. 1, figs a, b)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAC89005F0AF9E8923DF970" ID-GBIF-Occurrence="2521263199" box="[820,1357,1655,1682]" collectionCode="NMV" pageId="4" specimenCode="P332717" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAC89005F0AF9E89405F970" box="[820,885,1656,1682]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">NMV</collectionCode>
<specimenCode id="DBF76E48FFAC89005F40F9E89495F970" box="[894,997,1656,1682]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">P332717</specimenCode>
, a fixed tergum (pl. 1, figs i, j)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAC89005965F9E8952FF954" ID-GBIF-Occurrence="2521263193" collectionCode="NMV" pageId="4" specimenCode="P332718" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAC89005965F9E892ECF970" box="[1371,1436,1656,1682]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">NMV</collectionCode>
<specimenCode id="DBF76E48FFAC89005CA9F90B978EF957" box="[151,254,1691,1717]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">P332718</specimenCode>
, a fixed scutum (pl. 1, figs e, f)
</materialsCitation>
;
<materialsCitation id="3B39CC6EFFAC89005E55F90C9334F954" ID-GBIF-Occurrence="2521263195" box="[619,1092,1691,1718]" collectionCode="NMV" pageId="4" specimenCode="P332719" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED405EF6FFAC89005E55F90C95DCF954" box="[619,684,1692,1718]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">NMV</collectionCode>
<specimenCode id="DBF76E48FFAC89005E8CF90B9469F957" box="[690,793,1691,1717]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="4" type="Museum">P332719</specimenCode>
, a rostrum (pl. 1, figs c, d)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEEC633FFAC89005CF9F92F95F5F93B" blockId="4.[151,1437,1476,2040]" box="[199,645,1726,1753]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F92F9615F93B" bold="true" box="[199,357,1726,1753]" pageId="4">Type locality.</emphasis>
LACMIP locality 16851.
</paragraph>
<paragraph id="8BEEC633FFAC89005CF9F972945CF91C" blockId="4.[151,1437,1476,2040]" box="[199,812,1762,1790]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F97296CAF91F" bold="true" box="[199,442,1762,1789]" pageId="4">Stratigraphic range.</emphasis>
Basal middle Eocene (Lutetian).
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAC89005CF9F89696FDF88A" pageId="4" type="distribution">
<paragraph id="8BEEC633FFAC89005CF9F8969311F8C3" blockId="4.[151,1437,1476,2040]" box="[199,1121,1798,1825]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F8969611F8C3" bold="true" box="[199,353,1798,1825]" pageId="4">Distribution.</emphasis>
Crescent/McIntosh transitional zone, Doty Hills,
<collectingRegion id="499508D1FFAC89005FB0F8969366F8C2" box="[910,1046,1798,1824]" country="United States of America" name="Washington" pageId="4">Washington</collectingRegion>
,
<collectingCountry id="F34686A3FFAC8900581FF897932DF8C3" box="[1057,1117,1799,1825]" name="United States of America" pageId="4">USA</collectingCountry>
.
</paragraph>
<paragraph id="8BEEC633FFAC89005CF9F8B996FDF88A" blockId="4.[151,1437,1476,2040]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F8B996CDF8A6" bold="true" box="[199,445,1833,1860]" pageId="4">Additional material.</emphasis>
25 loose plates, including carinae, rostra, movable and fixed terga, fixed scuta; most plates are incomplete.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAC89025CF9F8E19521FE69" lastPageId="6" pageId="4" type="description">
<paragraph id="8BEEC633FFAC89005CF9F8E19277F815" blockId="4.[151,1437,1476,2040]" pageId="4">
<emphasis id="B9251A21FFAC89005CF9F8E19629F86E" bold="true" box="[199,345,1905,1932]" pageId="4">Description.</emphasis>
Wall plates with exterior having moderately rugose longitudinal ribbing crossed by irregular, fine growth lines; all plates perforated by fine punctae. Carina highly asymmetrical, with strong tilt to fixed side; fixed tergal margin with two to three articular ribs, very short, indistinct; rostral margin with four to five articular ribs, top two long, lower very short, smooth; interior with prominent dependent sheath widest at apex of plate.
</paragraph>
<caption id="DF2E96BBFFAD89015CA9FBE59247FAA0" ID-DOI="http://doi.org/10.5281/zenodo.3586818" ID-Zenodo-Dep="3586818" httpUri="https://zenodo.org/record/3586818/files/figure.png" pageId="5" startId="5.[151,234,1141,1166]" targetBox="[185,1374,182,1097]" targetPageId="5">
<paragraph id="8BEEC633FFAD89015CA9FBE59247FAA0" blockId="5.[151,1437,1141,1346]" pageId="5">
<emphasis id="B9251A21FFAD89015CA9FBE59673FB6C" bold="true" box="[151,259,1141,1166]" pageId="5">PLATE 1.</emphasis>
<taxonomicName id="4C51BDB0FFAD89015D37FBE6969CFB6C" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[265,492,1142,1166]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="5" phylum="Arthropoda" rank="species" species="gailgoedertae" status="sp. nov.">
<emphasis id="B9251A21FFAD89015D37FBE6969CFB6C" box="[265,492,1142,1166]" italics="true" pageId="5">Verruca gailgoedertae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A216A75AFFAD89015DC6FBE69536FB6F" box="[504,582,1142,1165]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
a. Paratype NMV P332716, carina (exterior); b. paratype NMV P332716, carina (interior); c. paratype NMV P332719, rostrum (exterior); d. paratype NMV P332719, rostrum (interior); e. paratype NMV P332718, fixed scutum (exterior); f. paratype NMV P332718, fixed scutum (interior); g. holotype LACMIP 14759, movable tergum (exterior); h. holotype LACMIP 14759, movable tergum (interior); i. paratype NMV P332717, fixed tergum (exterior); j. paratype NMV P332717, fixed tergum (interior); k. paratype, fixed scutum (exterior); l. paratype fixed scutum (interior). All specimens are from the Lower Middle Eocene Crescent-McIntosh transitional zone, Location (CSUN 1573), Doty Hills, Washington, USA.
</paragraph>
</caption>
<paragraph id="8BEEC633FFAD89015CF9FAE19549FA30" blockId="5.[151,1437,1392,2030]" pageId="5">Rostrum rounded, fixed scutal margin with two very short, indistinct articular ribs; carinal margin with four ribs, long; ornamented by growth ridges and vague radial ridges; interior with very short sheath, dependent full length; interior otherwise featureless.</paragraph>
<paragraph id="8BEEC633FFAD89015CF9FA4C9240F964" blockId="5.[151,1437,1392,2030]" pageId="5">Fixed scutum convex, elongated horizontally; fixed tergal margin convex, without ribs; articular wing very long, triangular, separated from plate by wide sulcus, sharply edged on plate; upper margin of wing slightly concave; up to three articular ribs below wing; ornamented by regularly spaced growth lines on wing only, and by low radial ribs, especially on basal portions of plate; interior concave, with bordering ridge on wing; adductor plate high, erect, apical part highest, flat, with two ridges extending from corners to apex of plate; myophore absent.</paragraph>
<paragraph id="8BEEC633FFAD89015CF9F9009626F8D8" blockId="5.[151,1437,1392,2030]" pageId="5">Fixed tergum higher than wide, slightly bowed, plate flared slightly basally; fixed scutal margin flat, ala nonprotuberant, with indentation below; carinal margin with protruding ala extending from umbo, with indentation separating ala from four vague articular ribs below; apical margin slightly concave, umbo removed from apex; exterior ornamented by low, fine radial ridges, especially basally; interior very concave, apical shelf nearly horizontal, wide, dependent.</paragraph>
<paragraph id="8BEEC633FFAD89015CF9F8D495B3F80C" blockId="5.[151,1437,1392,2030]" pageId="5">Movable scutum slightly convex, moderately curved tergally; apico-basal ridge very narrow, protruding; first articular rib extending to level of basi-occludent angle, nearly as wide as fixed scutal section; second articular rib centrally placed, protruding by its own width, sharply pointed; occludent margin strongly and smoothly convex; ornamented by regularly spaced, fine growth lines; interior with two small teeth present on movable tergal margin; movable tergal margin with internal growth lines.</paragraph>
<paragraph id="8BEEC633FFAE89025CF9FF099521FE69" blockId="6.[151,1437,153,899]" pageId="6">Movable tergum slightly wider than high, flat to slightly bowed; apico-basal ridge curved slightly toward carinal margin, flat, with slightly overhanging margins, merging into plate near apex, growth lines straight, perpendicular to carinal margin; carinal portion triangular, fixed tergal margin convex, ornamented by fine regularly spaced growth lines and two to three faint radial incised lines; first articular ridge short, nearly width of second, with smooth area between ridge and second articular ridge; movable scutal margin indented at point of contact with smooth area; interior concave, with thickened occludent surfaces, bearing faint growth ridges, and shallow furrow coinciding with apico-basal ridge in lower half of plate.</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CF9FE049559FD11" pageId="6" type="discussion">
<paragraph id="8BEEC633FFAE89025CF9FE0493D6FD81" blockId="6.[151,1437,153,899]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FE04963CFE4D" bold="true" box="[199,332,404,431]" pageId="6">Discussion.</emphasis>
<taxonomicName id="4C51BDB0FFAE89025D69FE069527FE4D" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[343,599,405,431]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="gailgoedertae" status="sp. nov.">
<emphasis id="B9251A21FFAE89025D69FE069527FE4D" box="[343,599,405,431]" italics="true" pageId="6">Verruca gailgoedertae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFAE89025E5FFE0595CCFE4D" box="[609,700,405,431]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
may be distinguished from most other congeners by its strong rugose ribbing. The presence of punctae presently distinguishes it from
<taxonomicName id="4C51BDB0FFAE89025FF0FE2A9307FE31" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[974,1143,441,467]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="alabamensis" status="sp. nov.">
<emphasis id="B9251A21FFAE89025FF0FE2A9307FE31" box="[974,1143,441,467]" italics="true" pageId="6">V. alabamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFAE890258BEFE2993ABFE31" box="[1152,1243,441,467]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C51BDB0FFAE890258D6FE2A923EFE31" authorityName="Buckeridge" authorityYear="2011" box="[1256,1358,442,467]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="sauria">
<emphasis id="B9251A21FFAE890258D6FE2A923EFE31" box="[1256,1358,442,467]" italics="true" pageId="6">V. sauria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C51BDB0FFAE890259B6FE2A957CFE15" authority="Buckeridge &amp; Jagt, 2008" authorityName="Buckeridge &amp; Jagt" authorityYear="2008" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="B9251A21FFAE890259B6FE2A9793FE15" italics="true" pageId="6">V. punica</emphasis>
<bibRefCitation id="EFC0BBC2FFAE89025CD2FE4D957CFE15" author="Buckeridge, J. S. &amp; Jagt, J. W. M. &amp; Speijer, R. P." box="[236,524,477,503]" pageId="6" pagination="37 - 46" refId="ref10376" refString="Buckeridge, J. S., Jagt, J. W. M. &amp; Speijer, R. P. (2008) Verruca punica, a new species of verrucomorph barnacle (Crustacea, Cirripedia, Thoracica) from the lower Danian (Palaeocene) of Tunisia. Zootaxa, 1844 (1), 37 - 46. https: // doi. org / 10.11646 / zootaxa. 1844.1.3" type="journal article" year="2008">Buckeridge &amp; Jagt, 2008</bibRefCitation>
</taxonomicName>
. It can be distinguished from
<taxonomicName id="4C51BDB0FFAE89025F5EFE4E9491FE15" authority="Perreault &amp; Buckeridge, 2019" authorityName="Perreault &amp; Buckeridge" authorityYear="2019" box="[864,993,477,503]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="sorrellae" status="sp. nov.">
<emphasis id="B9251A21FFAE89025F5EFE4E9491FE15" box="[864,993,477,503]" italics="true" pageId="6">V. sorrellae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216A75AFFAE89025FD7FE4D9332FE15" box="[1001,1090,477,503]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
by its movable tergum, which has equal width first and second ribs: the shell would have been more depressed than
<taxonomicName id="4C51BDB0FFAE89025866FD9293B2FDF9" authorityName="Steinmann in Wilkins" authorityYear="1921" box="[1112,1218,514,539]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="rocana">
<emphasis id="B9251A21FFAE89025866FD9293B2FDF9" box="[1112,1218,514,539]" italics="true" pageId="6">V. rocana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C51BDB0FFAE890258C7FD929216FDF9" authorityName="Buckeridge in Buckeridge &amp; Finger" authorityYear="2001" box="[1273,1382,513,539]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="digitali">
<emphasis id="B9251A21FFAE890258C7FD929216FDF9" box="[1273,1382,513,539]" italics="true" pageId="6">V. digitali</emphasis>
</taxonomicName>
, and is more circular in outline than
<taxonomicName id="4C51BDB0FFAE89025DC1FDB69518FDDD" authorityName="Buckeridge &amp; Jagt" authorityYear="2008" box="[511,616,550,575]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="B9251A21FFAE89025DC1FDB69518FDDD" box="[511,616,550,575]" italics="true" pageId="6">V. punica</emphasis>
</taxonomicName>
. Although similar in size to
<taxonomicName id="4C51BDB0FFAE89025F90FDB6934EFDDD" authorityName="Buckeridge" authorityYear="1983" box="[942,1086,550,575]" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="tasmanica">
<emphasis id="B9251A21FFAE89025F90FDB6934EFDDD" box="[942,1086,550,575]" italics="true" pageId="6">V. tasmanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C51BDB0FFAE89025849FDB697A2FD81" authority="Buckeridge, 1983" authorityName="Buckeridge" authorityYear="1983" class="Maxillopoda" family="Verrucidae" genus="Verruca" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="nuciformis">
<emphasis id="B9251A21FFAE89025849FDB6927CFDDD" box="[1143,1292,549,575]" italics="true" pageId="6">V. nuciformis</emphasis>
<bibRefCitation id="EFC0BBC2FFAE8902592AFDB597A2FD81" author="Buckeridge, J. S." pageId="6" pagination="1 - 151" refId="ref10049" refString="Buckeridge, J. S. (1983) Fossil barnacles (Cirripedia: Thoracica) of New Zealand and Australia. New Zealand Geology Survey Paleontological Bulletin, 50, 1 - 151." type="journal article" year="1983">Buckeridge, 1983</bibRefCitation>
</taxonomicName>
, it has more than twice the number of interlocking ribs between the carina and rostrum.
</paragraph>
<paragraph id="8BEEC633FFAE89025CA9FDFD9559FD11" blockId="6.[151,1437,153,899]" pageId="6">
On the basis of its lack of abrasion, this species is interpreted as having lived in quieter sublittoral environments. The
<typeStatus id="54EA7891FFAE89025CF4FD01978AFD49" box="[202,250,657,683]" pageId="6">type</typeStatus>
locality is interpreted as bathyal with introductions from shallower waters (
<bibRefCitation id="EFC0BBC2FFAE8902586DFD019200FD49" author="Squires, R. L. &amp; Goedert, J. L." box="[1107,1392,656,683]" pageId="6" pagination="254 - 269" refId="ref13114" refString="Squires, R. L. &amp; Goedert, J. L. (1995) New species of Middle Eocene gastropods from the northern Doty Hills, southwestern Washington. The Veliger, 38 (3), 254 - 269." type="journal article" year="1995">Squires &amp; Goedert, 1995</bibRefCitation>
) up to intertidal, and is thus a thanatocoenosis. The other two localities are interpreted as shallow shelf environments less disturbed than the
<typeStatus id="54EA7891FFAE89025DA6FD4996B8FD11" box="[408,456,729,755]" pageId="6">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CF9FD6D949CFCD9" pageId="6" type="etymology">
<paragraph id="8BEEC633FFAE89025CF9FD6D949CFCD9" blockId="6.[151,1437,153,899]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FD6D963EFCF5" bold="true" box="[199,334,765,791]" pageId="6">Etymology.</emphasis>
Named in honor of the late Gail Goedert, wife of James Goedert, who together discovered this and many other fossil localities, and aided
<collectingRegion id="499508D1FFAE89025E78FCB195BBFCD9" box="[582,715,801,827]" country="United States of America" name="Washington" pageId="6">Washington</collectingRegion>
paleontology immensely.
</paragraph>
</subSubSection>
<subSubSection id="C34B95B8FFAE89025CF9FCD59359FC61" pageId="6" type="materials_examined">
<paragraph id="8BEEC633FFAE89025CF9FCD59359FC61" blockId="6.[151,1437,153,899]" pageId="6">
<emphasis id="B9251A21FFAE89025CF9FCD596D6FCBD" bold="true" box="[199,422,836,863]" pageId="6">Type Repositories.</emphasis>
<materialsCitation id="3B39CC6EFFAE89025D90FCD59359FC61" ID-GBIF-Occurrence="2521263197" collectionCode="CA, NMV" country="Australia" county="Natural History Museum of Los Angeles County" location="Museum" municipality="Melbourne" pageId="6" specimenCode="P332716-9" specimenCount="2" stateProvince="Victoria" typeStatus="holotype">
<typeStatus id="54EA7891FFAE89025D90FCD59566FCBD" box="[430,534,837,863]" pageId="6" type="holotype">Holotype</typeStatus>
LACMIP 14759 held by the
<collectingCounty id="628FBEBFFFAE89025F57FCD592E8FCBD" box="[873,1432,837,863]" pageId="6">Natural History Museum of Los Angeles County</collectingCounty>
,
<collectionCode id="ED405EF6FFAE89025CA9FCF997B1FC61" box="[151,193,873,899]" country="USA" name="Chicago Academy of Sciences" pageId="6" type="Museum">CA</collectionCode>
;
<typeStatus id="54EA7891FFAE89025CF3FCF99648FC61" box="[205,312,873,899]" pageId="6" type="paratype">paratypes</typeStatus>
<collectionCode id="ED405EF6FFAE89025D01FCF996F0FC61" box="[319,384,873,899]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">NMV</collectionCode>
<specimenCode id="DBF76E48FFAE89025DB9FCF99572FC61" box="[391,514,873,899]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" type="Museum">P332716-9</specimenCode>
held by
<location id="8E8E90E8FFAE89025E5DFCF995B5FC61" LSID="urn:lsid:plazi:treatment:03F87725FFAC89025CA9FAAB9359FC61:8E8E90E8FFAE89025E5DFCF995B5FC61" box="[611,709,873,899]" country="Australia" county="Natural History Museum of Los Angeles County" municipality="Melbourne" name="Museum" pageId="6" stateProvince="Victoria">Museum</location>
<collectingRegion id="499508D1FFAE89025EF2FCF99459FC61" box="[716,809,873,899]" country="Australia" name="Victoria" pageId="6">Victoria</collectingRegion>
,
<collectingMunicipality id="6B8A5C49FFAE89025F0AFCF994C2FC61" box="[820,946,873,899]" pageId="6">Melbourne</collectingMunicipality>
,
<collectingCountry id="F34686A3FFAE89025F85FCF99355FC61" box="[955,1061,873,899]" name="Australia" pageId="6">Australia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>