treatments-xml/data/03/ED/2C/03ED2C23FF941264E1A9F8A1C724FAD4.xml
2024-06-21 12:22:17 +02:00

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<document id="338FEDEEB5EB7CBE721FEB3A3010FC48" ID-CLB-Dataset="22255" ID-DOI="10.11646/zootaxa.4772.3.8" ID-GBIF-Dataset="c7693ffd-e157-4dfe-ad25-dd8c054b07e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819768" ID-ZooBank="D166A1EF-943F-44BC-86E6-15682456F2AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175862959" checkinUser="plazi" docAuthor="Xu, Deliang, Dietrich, Christopher H., Webb, M. D. &amp; Zhang, Yalin" docDate="2020" docId="03ED2C23FF941264E1A9F8A1C724FAD4" docLanguage="en" docName="zootaxa.4772.3.8.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kutara Distant 1908" docType="treatment" docVersion="8" lastPageNumber="586" masterDocId="FFD4545BFF941265E13EFF9CC642FF82" masterDocTitle="A new species of the Oriental leafhopper genus Kutara Distant (Hemiptera: Cicadellidae: Deltocephalinae) from Vietnam" masterLastPageNumber="592" masterPageNumber="585" pageNumber="585" updateTime="1698839123959" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="221945CE16B9E4AA2DFB25673390D674">A new species of the Oriental leafhopper genus Kutara Distant (Hemiptera: Cicadellidae: Deltocephalinae) from Vietnam</mods:title>
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<mods:namePart id="C6046FF0386AD23E1E305FF5F4AD4B0B">Xu, Deliang</mods:namePart>
<mods:affiliation id="F21DDF79DE10DCB3FF43555CADD872F6">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, China</mods:affiliation>
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<mods:namePart id="024894B86A9F50B0C07C5B0B1BBF4F17">Dietrich, Christopher H.</mods:namePart>
<mods:affiliation id="7A55B43E0DBD33D2B81667A89BEDCD7C">Illinois Natural History Survey, Prairie Research Institute, University of Illinois, 1816 S. Oak Street, Champaign, Illinois 61820, USA</mods:affiliation>
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<mods:namePart id="AE5FF60164AA30F034B75E5C4A1681EA">Webb, M. D.</mods:namePart>
<mods:affiliation id="3F8BB5F312C6A19857924BCDE783615F">The Natural History Museum, Cromwell Road, South Kensington, London, SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="0F806C42CE36A07C753932A424D012CA">Zhang, Yalin</mods:namePart>
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<treatment id="03ED2C23FF941264E1A9F8A1C724FAD4" ID-DOI="http://doi.org/10.5281/zenodo.3843717" ID-GBIF-Taxon="164035944" ID-Zenodo-Dep="3843717" LSID="urn:lsid:plazi:treatment:03ED2C23FF941264E1A9F8A1C724FAD4" httpUri="http://treatment.plazi.org/id/03ED2C23FF941264E1A9F8A1C724FAD4" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585">
<subSubSection id="C35ECEBEFF941265E1A9F8A1C70CF8D4" box="[151,334,1852,1879]" pageId="0" pageNumber="585" type="nomenclature">
<paragraph id="8BFB9D35FF941265E1A9F8A1C70CF8D4" blockId="0.[151,334,1852,1879]" box="[151,334,1852,1879]" pageId="0" pageNumber="585">
<heading id="D0B32A59FF941265E1A9F8A1C70CF8D4" bold="true" box="[151,334,1852,1879]" fontSize="11" level="1" pageId="0" pageNumber="585" reason="1">
<taxonomicName id="4C44E6B6FF941265E1A9F8A1C70CF8D4" ID-CoL="8R5D6" authority="Distant" authorityName="Distant" authorityYear="1908" box="[151,334,1852,1879]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF941265E1A9F8A1C70CF8D4" bold="true" box="[151,334,1852,1879]" pageId="0" pageNumber="585">
<emphasis id="B9304127FF941265E1A9F8A1C6AEF8D5" bold="true" box="[151,236,1853,1879]" italics="true" pageId="0" pageNumber="585">Kutara</emphasis>
Distant
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35ECEBEFF941265E1A9F81DC214F81B" box="[151,1110,1920,1945]" pageId="0" pageNumber="585" type="reference_group">
<paragraph id="8BFB9D35FF941265E1A9F81DC214F81B" blockId="0.[151,1110,1920,1945]" box="[151,1110,1920,1945]" pageId="0" pageNumber="585">
<treatmentCitationGroup id="AB54BA1BFF941265E1A9F81DC214F81B" box="[151,1110,1920,1945]" pageId="0" pageNumber="585">
<treatmentCitation id="0AE5BB24FF941265E1A9F81DC7E6F81B" author="Distant, W. L." box="[151,420,1920,1945]" page="308" pageId="0" pageNumber="585" year="1908">
<taxonomicName id="4C44E6B6FF941265E1A9F81DC7E6F81B" ID-CoL="8R5D6" authority="Distant, 1908: 308" authorityName="Distant" authorityPageNumber="308" authorityYear="1908" box="[151,420,1920,1945]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF941265E1A9F81DC69DF81A" box="[151,223,1921,1944]" italics="true" pageId="0" pageNumber="585">Kutara</emphasis>
<bibRefCitation id="EFD5E0C4FF941265E1DBF81CC7E6F81B" author="Distant, W. L." box="[229,420,1920,1945]" pageId="0" pageNumber="585" refId="ref4519" refString="Distant, W. L. (1908) The fauna of British India, including Ceylon and Burma. Rhynchota. IV. Homoptera and appendix (Pt.). Taylor &amp; Francis, London, 501 pp." type="book" year="1908">Distant, 1908: 308</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54FF2397FF941265E093F81DC79DF81B" box="[429,479,1921,1945]" pageId="0" pageNumber="585">Type</typeStatus>
species:
<taxonomicName id="4C44E6B6FF941265E303F81DC51EF81B" ID-CoL="3RGG6" authority="Distant" authorityName="Distant" authorityYear="1908" box="[573,860,1920,1945]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="585" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B9304127FF941265E303F81DC548F81A" box="[573,778,1920,1944]" italics="true" pageId="0" pageNumber="585">Kutara brunnescens</emphasis>
Distant
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35ECEBEFF941264E1A9F85AC540FBB5" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585" type="diagnosis">
<paragraph id="8BFB9D35FF941264E1A9F85AC688FDE1" blockId="0.[151,1436,1990,2017]" lastBlockId="1.[151,1437,152,1366]" lastPageId="1" lastPageNumber="586" pageId="0" pageNumber="585">
<emphasis id="B9304127FF941265E1A9F85AC748F863" bold="true" box="[151,266,1990,2017]" pageId="0" pageNumber="585">Diagnosis</emphasis>
. Moderately robust leafhoppers with coloration ochraceous to brown with a black transverse spot ante- riorly on crown; antennal pedicel dark brown. Head wider than pronotum with two transverse carina on anterior margin with a sulcus between; anterior tentorial branches L-shaped; crown of similar length throughout width, posteriorly elevated, anterior and posterior margin nearly parallel; ocelli situated on sulcus on anterior margin of head, distant from eyes; face rather flat and broad; frontoclypeal suture present; anteclypeus with apex slightly broader than base, sides concave; lora conspicuously broader than anteclypeus; gena prominent; antennae long, more than half body length, arising near upper corners of eyes; antennal ledge strong and oblique. Pronotum with transverse striations and dense punctures, anterior margin arched, lateral margin short, hind margin concave. Scutellum similar in length to pronotum with transverse depression distinct. Forewing with four apical cells and three subapical cells, inner subapical cell closed, cross vein present between claval suture and A1 and between A1 and A2, appendix narrow. Fore femur with anteromedial (AM1) seta present and more additional proximal macrosetae; intercalary (IC) row with fine setae apically, anteroventral (AV) row with short or longish microsetae on ventral margin. Protibia dorsal surface rounded with 4+4 or more dorsal setae (
<figureCitation id="137F81B0FF951264E23CFDB8C51EFDBD" box="[770,860,548,575]" captionStart="FIGURE 1" captionStartId="4.[151,250,1932,1957]" captionTargetBox="[197,1400,196,1876]" captionTargetId="figure@4.[184,1402,181,1906]" captionTargetPageId="4" captionText="FIGURE 1. Kutara trispinosa Xu &amp; Zhang sp. nov., male; A, Habitus, dorsal view; B, Head, pronotum and scutellum, dorsal view; C, Face, ventral view; D, Head, dorsoanterior view; E, Lateral view; F, Male abdomen, ventral view; G, Fore femur and tibia, anterior view; H, Apex of hind femur, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3825502" httpUri="https://zenodo.org/record/3825502/files/figure.png" pageId="1" pageNumber="586">Fig. 1G</figureCitation>
). Metafemur with apical setal formula 2+2+1 (
<figureCitation id="137F81B0FF951264E451FDB8C6FEFDE1" captionStart="FIGURE 1" captionStartId="4.[151,250,1932,1957]" captionTargetBox="[197,1400,196,1876]" captionTargetId="figure@4.[184,1402,181,1906]" captionTargetPageId="4" captionText="FIGURE 1. Kutara trispinosa Xu &amp; Zhang sp. nov., male; A, Habitus, dorsal view; B, Head, pronotum and scutellum, dorsal view; C, Face, ventral view; D, Head, dorsoanterior view; E, Lateral view; F, Male abdomen, ventral view; G, Fore femur and tibia, anterior view; H, Apex of hind femur, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3825502" httpUri="https://zenodo.org/record/3825502/files/figure.png" pageId="1" pageNumber="586">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="8BFB9D35FF951264E1F9FDF0C4BAFC48" blockId="1.[151,1437,152,1366]" pageId="1" pageNumber="586">
Male genitalia: pygofer basolateral membranous cleft present; with an internal posterior ledge; lateral lobe caudally truncate or broad to narrowly rounded, without apical process or with a ventrocaudal process, with several macrosetae and microsetae. Valve triangular or pentagonal. Subgenital plate triangular without macrosetae, with fine hair-like microsetae arising from lateral margin, irregularly arranged. Style with apical process of variable length and shape, robust; preapical lobe well developed or not. Connective usually Y-shaped, stem similar in length to arms, sometimes bifid distally, ventral margin angled at mid-length in lateral view. Aedeagus with dorsal apodeme well developed; shaft slender and elongate, curved dorsad with a subapical prolongation arising from ventral margin exceeding apex of shaft, with or without a pair of apical processes, rarely with pair of basal processes (
<taxonomicName id="4C44E6B6FF951264E425FCF5C754FC25" authority="Kuoh" authorityName="Kuoh" authorityYear="1992" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="nigrifasciata">
<emphasis id="B9304127FF951264E425FCF5C36CFC00" box="[1307,1326,873,898]" italics="true" pageId="1" pageNumber="586">K</emphasis>
.
<emphasis id="B9304127FF951264E402FCF5C68DFC24" italics="true" pageId="1" pageNumber="586">nigrifasciata</emphasis>
Kuoh
</taxonomicName>
) or lateral spines (
<taxonomicName id="4C44E6B6FF951264E0DAFC11C556FC24" authority="Zhang &amp; Chen" authorityName="Zhang &amp; Chen" authorityYear="1997" box="[484,788,908,935]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="B9304127FF951264E0DAFC11C7B5FC24" box="[484,503,909,934]" italics="true" pageId="1" pageNumber="586">K</emphasis>
.
<emphasis id="B9304127FF951264E33AFC11C42AFC25" box="[516,616,909,935]" italics="true" pageId="1" pageNumber="586">spinifera</emphasis>
Zhang &amp; Chen
</taxonomicName>
and
<taxonomicName id="4C44E6B6FF951264E277FC11C200FC25" authority="Kuoh" authorityName="Kuoh" authorityYear="1992" box="[841,1090,908,935]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="nigrifasciata">
<emphasis id="B9304127FF951264E277FC11C5BAFC25" box="[841,1016,909,935]" italics="true" pageId="1" pageNumber="586">K. nigrifasciata</emphasis>
Kuoh
</taxonomicName>
); gonopore subapical on dorsal side; anal tube large, sclerotized laterally and dorsally.
</paragraph>
<paragraph id="8BFB9D35FF951264E1F9FC48C540FBB5" blockId="1.[151,1437,152,1366]" pageId="1" pageNumber="586">Female genitalia: pygofer with numerous setae; ovipositor conspicuously surpassing apex of pygofer; first valvulae with strigate texture dorsally; second valvulae with tooth-like prominence at mid-length of dorsal margin, with dorsal teeth restricted to approximately apical 1/3.</paragraph>
</subSubSection>
<subSubSection id="C35ECEBEFF951264E1F9FBDCC517FBD8" box="[199,853,1088,1115]" pageId="1" pageNumber="586" type="distribution">
<paragraph id="8BFB9D35FF951264E1F9FBDCC517FBD8" blockId="1.[151,1437,152,1366]" box="[199,853,1088,1115]" pageId="1" pageNumber="586">
<emphasis id="B9304127FF951264E1F9FBDCC71BFBD9" bold="true" box="[199,345,1088,1115]" pageId="1" pageNumber="586">Distribution</emphasis>
. Palaearctic, Oriental and Australian Region.
</paragraph>
</subSubSection>
<subSubSection id="C35ECEBEFF951264E1F9FBF9C724FAD4" pageId="1" pageNumber="586" type="discussion">
<paragraph id="8BFB9D35FF951264E1F9FBF9C724FAD4" blockId="1.[151,1437,152,1366]" pageId="1" pageNumber="586">
<emphasis id="B9304127FF951264E1F9FBF9C779FBFD" bold="true" box="[199,315,1125,1151]" pageId="1" pageNumber="586">Remarks.</emphasis>
<taxonomicName id="4C44E6B6FF951264E07AFBF9C7D1FBFC" authorityName="Distant" authorityYear="1908" box="[324,403,1125,1150]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E07AFBF9C7D1FBFC" box="[324,403,1125,1150]" italics="true" pageId="1" pageNumber="586">Kutara</emphasis>
</taxonomicName>
, together with
<taxonomicName id="4C44E6B6FF951264E37FFBF8C53BFBFD" authority="Kwon &amp; Lee" authorityName="Kwon &amp; Lee" authorityYear="1979" box="[577,889,1124,1151]" class="Insecta" family="Cicadellidae" genus="Drabescoides" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E37FFBF8C499FBFC" box="[577,731,1124,1150]" italics="true" pageId="1" pageNumber="586">Drabescoides</emphasis>
Kwon &amp; Lee
</taxonomicName>
and
<taxonomicName id="4C44E6B6FF951264E28DFBF8C36CFBFC" authority="Lu, Webb &amp; Zhang" authorityName="Lu, Webb &amp; Zhang" authorityYear="2018" box="[947,1326,1124,1151]" class="Insecta" family="Cicadellidae" genus="Papuakutara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E28DFBF8C207FBFC" box="[947,1093,1124,1150]" italics="true" pageId="1" pageNumber="586">Papuakutara</emphasis>
Lu, Webb &amp; Zhang
</taxonomicName>
, form the
<taxonomicName id="4C44E6B6FF951264E1A9FB15C6A4FB20" authorityName="Distant" authorityYear="1908" box="[151,230,1161,1186]" class="Insecta" family="Cicadellidae" genus="Kutara" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E1A9FB15C6A4FB20" box="[151,230,1161,1186]" italics="true" pageId="1" pageNumber="586">Kutara</emphasis>
</taxonomicName>
group. The group is distinguished by the head marking with a transverse medial spot or band and shape of the head (
<figureCitation id="137F81B0FF951264E03BFB30C71EFB45" box="[261,348,1196,1223]" captionStart="FIGURE 1" captionStartId="4.[151,250,1932,1957]" captionTargetBox="[197,1400,196,1876]" captionTargetId="figure@4.[184,1402,181,1906]" captionTargetPageId="4" captionText="FIGURE 1. Kutara trispinosa Xu &amp; Zhang sp. nov., male; A, Habitus, dorsal view; B, Head, pronotum and scutellum, dorsal view; C, Face, ventral view; D, Head, dorsoanterior view; E, Lateral view; F, Male abdomen, ventral view; G, Fore femur and tibia, anterior view; H, Apex of hind femur, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3825502" httpUri="https://zenodo.org/record/3825502/files/figure.png" pageId="1" pageNumber="586">Fig. 1A</figureCitation>
) and short vertex with posterior margin elevated (
<figureCitation id="137F81B0FF951264E2AFFB30C5A5FB45" box="[913,999,1196,1223]" captionStart="FIGURE 1" captionStartId="4.[151,250,1932,1957]" captionTargetBox="[197,1400,196,1876]" captionTargetId="figure@4.[184,1402,181,1906]" captionTargetPageId="4" captionText="FIGURE 1. Kutara trispinosa Xu &amp; Zhang sp. nov., male; A, Habitus, dorsal view; B, Head, pronotum and scutellum, dorsal view; C, Face, ventral view; D, Head, dorsoanterior view; E, Lateral view; F, Male abdomen, ventral view; G, Fore femur and tibia, anterior view; H, Apex of hind femur, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3825502" httpUri="https://zenodo.org/record/3825502/files/figure.png" pageId="1" pageNumber="586">Fig. 1E</figureCitation>
). It can be distinguished from
<taxonomicName id="4C44E6B6FF951264E47EFB30C69DFB68" authorityName="Kwon &amp; Lee" authorityYear="1979" class="Insecta" family="Cicadellidae" genus="Drabescoides" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E47EFB30C69DFB68" italics="true" pageId="1" pageNumber="586">Drabescoides</emphasis>
</taxonomicName>
by the narrower stem of the connective and male pygofer side without posterior process and from
<taxonomicName id="4C44E6B6FF951264E474FB4DC6A2FA8C" authorityName="Lu, Webb &amp; Zhang" authorityYear="2018" class="Insecta" family="Cicadellidae" genus="Papuakutara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B9304127FF951264E474FB4DC6A2FA8C" italics="true" pageId="1" pageNumber="586">Papuakutara</emphasis>
</taxonomicName>
in having a cross vein between the claval veins of the forewing, and the connective ventral margin angled at mid-length in lateral view and aedeagal shaft usually extended beyond the gonopore and without a basal process in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>