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<document id="BEDCA71FF0F77E9F961948BB2A9E61AD" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBD3E111B3CD2F9BEFCDDFB53" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola tishechkini Park and Carlton, new species" docType="treatment" docVersion="24" lastPageNumber="88" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="88" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="27A9990A6486905093E694EF8A214987">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="0F020CF16F7E9E664441585DBF55E495">Park, J. - S.</mods:namePart>
<mods:affiliation id="CAC585514E8811083D71F94DEB80CCE6">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="C2F7241F714857E66EE875795070C253">Carlton, C. E.</mods:namePart>
<mods:affiliation id="4C54570F8B4AA56D703614E7A748D24C">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="1D823B9B4999936162F21CC175AFFB82">The Coleopterists Bulletin</mods:title>
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<mods:date id="AB2D1BBC87D5F20E5CC870ADE9780228">2014</mods:date>
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<mods:title id="17E1DFC402B6FAA425B26FF23BCAA6CF">mo 13</mods:title>
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<mods:number id="71E71AF6949AEDF4E5F70AA851EFA6F8">2014-12-01</mods:number>
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<treatment id="03D82A3EBD3E111B3CD2F9BEFCDDFB53" ID-DOI="http://doi.org/10.5281/zenodo.5765181" ID-GBIF-Taxon="191245053" ID-Zenodo-Dep="5765181" LSID="urn:lsid:zoobank.org:act:5B96BFB4-14DA-4397-9B7A-730F53FB6C5F" httpUri="http://treatment.plazi.org/id/03D82A3EBD3E111B3CD2F9BEFCDDFB53" lastPageNumber="88" pageId="87" pageNumber="88">
<subSubSection id="C36BC8A3BD3E111B3CD2F9BEFDABF9F8" box="[151,637,1613,1634]" pageId="87" pageNumber="88" type="nomenclature">
<paragraph id="8BCE9B28BD3E111B3CD2F9BEFDABF9F8" blockId="87.[151,637,1613,1660]" box="[151,637,1613,1634]" pageId="87" pageNumber="88">
<heading id="D0862C44BD3E111B3CD2F9BEFDABF9F8" box="[151,637,1613,1634]" centered="true" fontSize="10" level="2" pageId="87" pageNumber="88" reason="2">
<emphasis id="B905473ABD3E111B3CD2F9BEFDABF9F8" bold="true" box="[151,637,1613,1634]" pageId="87" pageNumber="88">
<taxonomicName id="4C71E0ABBD3E111B3CD2F9BEFDD7F9F8" ID-CoL="4TZ43" authority="Park and Carlton" authorityName="Park and Carlton" box="[151,513,1613,1634]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tishechkini" status="sp. nov.">
<emphasis id="B905473ABD3E111B3CD2F9BEFE9EF9F8" bold="true" box="[151,328,1613,1634]" italics="true" pageId="87" pageNumber="88">Sagola tishechkini</emphasis>
Park and Carlton
</taxonomicName>
,
<taxonomicNameLabel id="A236FA41BD3E111B3E49F9BEFDABF9F8" box="[524,637,1613,1634]" pageId="87" pageNumber="88" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3D6EF994FE3CF9E6" box="[299,490,1639,1660]" pageId="87" pageNumber="88" type="description">
<paragraph id="8BCE9B28BD3E111B3D6EF994FE3CF9E6" blockId="87.[151,637,1613,1660]" box="[299,490,1639,1660]" pageId="87" pageNumber="88">
(
<figureCitation id="134A87ADBD3E111B3D77F994FE69F9E6" box="[306,447,1639,1660]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Figs. 49b, 49d</figureCitation>
,
<figureCitation id="134A87ADBD3E111B3D8EF994FE35F9E6" box="[459,483,1639,1660]" captionStart="Fig" captionStartId="88.[731,763,1662,1681]" captionTargetBox="[712,1189,953,1639]" captionTargetId="figure-327@88.[709,1190,948,1642]" captionTargetPageId="88" captionText="Fig. 51. Known collection localities of New Zealand Sagola species. S. gilae: black circle; S. tishechkini: triangle; S. owensae: black square." figureDoi="http://doi.org/10.5281/zenodo.4907874" httpUri="https://zenodo.org/record/4907874/files/figure.png" pageId="87" pageNumber="88">51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3CEAF961FB47FEBD" pageId="87" pageNumber="88" type="materials_examined">
<paragraph id="8BCE9B28BD3E111B3CEAF961FDABF93D" blockId="87.[151,637,1682,1730]" box="[175,637,1682,1703]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3CEAF961FDABF93D" bold="true" box="[175,637,1682,1703]" pageId="87" pageNumber="88">
Type Material.
<materialsCitation id="3B199175BD3E111B3D11F961FDABF93D" ID-GBIF-Occurrence="3415153974" box="[340,637,1682,1703]" country="New Zealand" location="NEW ZEALAND" pageId="87" pageNumber="88" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CA258ABD3E111B3D11F961FE65F93D" box="[340,435,1682,1703]" pageId="87" pageNumber="88" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F366DBB8BD3E111B3DFBF961FDAFF93D" box="[446,633,1682,1703]" name="New Zealand" pageId="87" pageNumber="88">NEW ZEALAND</collectingCountry>
:
</materialsCitation>
</emphasis>
</paragraph>
<paragraph id="8BCE9B28BD3E111B3CD2F95FFB47FEBD" blockId="87.[151,637,1682,1730]" lastBlockId="87.[685,1172,194,1225]" pageId="87" pageNumber="88">
<materialsCitation id="3B199175BD3E111B3CD2F95FFDABF95B" ID-GBIF-Occurrence="3415154543" box="[151,637,1708,1730]" collectionCode="NZAC" location="Gisborne" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Gisborne">
<emphasis id="B905473ABD3E111B3CD2F95FFEDCF95B" bold="true" box="[151,266,1708,1729]" pageId="87" pageNumber="88">
<collectingRegion id="49B555CABD3E111B3CD2F95FFED2F95B" box="[151,260,1708,1729]" country="New Zealand" name="Gisborne" pageId="87" pageNumber="88">Gisborne</collectingRegion>
:
</emphasis>
<specimenCount id="9D7750A1BD3E111B3D50F95FFEE3F958" box="[277,309,1708,1730]" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
(
<collectionCode id="ED6003EDBD3E111B3D0DF95FFE43F95B" box="[328,405,1708,1729]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="87" pageNumber="88" type="Other institutional type">NZAC</collectionCode>
), aedeagus dissected
</materialsCitation>
<materialsCitation id="3B199175BD3E111B3EE8FF31FB5DFEBD" ID-GBIF-Occurrence="3415154524" collectingDate="1972-01-17" collectionCode="HOLO-" collectorName="G. W. Ramsay Litter" country="New Zealand" location="Lake Waikaremoana" pageId="87" pageNumber="88" specimenCount="1" typeStatus="holotype">
and mounted in balsam on a clear plastic card, “
<emphasis id="B905473ABD3E111B3EFCFF2EFC6EFF68" bold="true" box="[697,952,221,242]" pageId="87" pageNumber="88">
<collectingCountry id="F366DBB8BD3E111B3EFCFF2EFCABFF68" box="[697,893,221,242]" name="New Zealand" pageId="87" pageNumber="88">NEW ZEALAND</collectingCountry>
: GB:
</emphasis>
<location id="8EAECDF3BD3E111B3F87FF2EFB42FF68" LSID="urn:lsid:plazi:treatment:03D82A3EBD3E111B3CD2F9BEFCDDFB53:8EAECDF3BD3E111B3F87FF2EFB42FF68" box="[962,1172,221,242]" country="New Zealand" name="Lake Waikaremoana" pageId="87" pageNumber="88">Lake Waikaremoana</location>
<date id="FFCFBDE8BD3E111B3EE8FF04FCC4FE96" box="[685,786,247,268]" pageId="87" pageNumber="88" value="1972-01-17">
<collectingDate id="EF8B4400BD3E111B3EE8FF04FCC4FE96" box="[685,786,247,268]" pageId="87" pageNumber="88" value="1972-01-17">17 I 1972</collectingDate>
</date>
,
<collectorName id="2684FEFEBD3E111B3F5AFF04FC33FE96" box="[799,997,247,268]" pageId="87" pageNumber="88">G.W. Ramsay Litter</collectorName>
72/21”, “
<collectionCode id="ED6003EDBD3E111B380CFF04FD31FEBD" pageId="87" pageNumber="88">
<typeStatus id="54CA258ABD3E111B380CFF04FD31FEBD" pageId="87" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
</collectionCode>
<taxonomicName id="4C71E0ABBD3E111B3EB5FEE2FB57FEBD" authority="Park and Carlton 2013" authorityName="Park and Carlton" authorityYear="2013" box="[752,1153,273,295]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B905473ABD3E111B3EB5FEE2FC74FEBC" bold="true" box="[752,930,273,294]" italics="true" pageId="87" pageNumber="88">Sagola tishechkini</emphasis>
Park and Carlton 2013
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3E80FEDFFB99FEED" pageId="87" pageNumber="88" type="etymology">
<paragraph id="8BCE9B28BD3E111B3E80FEDFFB99FEED" blockId="87.[685,1172,194,1225]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3E80FEDFFCEEFEDB" bold="true" box="[709,824,300,321]" pageId="87" pageNumber="88">Etymology.</emphasis>
This species is named for LSAM alumnus, histerid specialist, and enthusiastic supporter of this study, Alexey K. Tishechkin.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3E80FE8FFBA8FD8C" pageId="87" pageNumber="88" type="diagnosis">
<paragraph id="8BCE9B28BD3E111B3E80FE8FFBA8FD8C" blockId="87.[685,1172,194,1225]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3E80FE8FFCFCFE0B" bold="true" box="[709,810,380,401]" pageId="87" pageNumber="88">Diagnosis.</emphasis>
This species can be distinguished from the other species of the
<taxonomicName id="4C71E0ABBD3E111B3F92FE64FBD0FE36" box="[983,1030,407,428]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="gilae">
<emphasis id="B905473ABD3E111B3F92FE64FBD0FE36" box="[983,1030,407,428]" italics="true" pageId="87" pageNumber="88">gilae</emphasis>
</taxonomicName>
species-group by the following combination of characters: body smaller, length
<quantity id="4C8936CDBD3E111B3F7FFE3FFC52FE7B" box="[826,900,460,481]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="87" pageNumber="88" unit="mm" value="1.8">1.8 mm</quantity>
; frontal rostrum bluntly triangular; gular depression with small process from anterior middle; shape of genitalia unique to species.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3E80FDE8FCB1FBE3" pageId="87" pageNumber="88" type="description">
<paragraph id="8BCE9B28BD3E111B3E80FDE8FCB1FBE3" blockId="87.[685,1172,194,1225]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3E80FDE8FC4EFDAA" bold="true" box="[709,920,539,560]" pageId="87" pageNumber="88">Description of Male.</emphasis>
Length
<quantity id="4C8936CDBD3E111B3FBFFDE8FB99FDAA" box="[1018,1103,539,560]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="87" pageNumber="88" unit="mm" value="1.8">1.8 mm</quantity>
. Body brown, antennae, elytra, legs, and maxillary palpi paler (
<figureCitation id="134A87ADBD3E111B3EABFDA2FC90FDFC" box="[750,838,593,614]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49b</figureCitation>
).
<emphasis id="B905473ABD3E111B3F1FFDA2FC4EFDFC" bold="true" box="[858,920,593,614]" pageId="87" pageNumber="88">Head:</emphasis>
As long as wide, widest across eyes (
<figureCitation id="134A87ADBD3E111B3F6BFD98FC53FD1A" box="[814,901,619,640]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49b</figureCitation>
). Gular region transversely depressed with small process medially and bearing dense setae. Antennomere 1 approximately 2.5 times longer than wide with dull surface, 2 longer than wide, 3 subquadrate, 45 longer than wide, 610 subquadrate. Frontal rostrum bluntly triangular. Frontal sulcus deep, reaching posterior end of eye. Anterior frontal fovea oval and partially covered by frontal rostrum, posterior frontal fovea oval. Eye prominent, one-third length of temple.
<emphasis id="B905473ABD3E111B3F46FC86FC8AFC10" bold="true" box="[771,860,885,906]" pageId="87" pageNumber="88">Thorax:</emphasis>
Prosternum as long as wide, widest at midpoint. Elytra rectangular (
<figureCitation id="134A87ADBD3E111B3877FC7CFB5FFC3E" box="[1074,1161,911,932]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49b</figureCitation>
). Meso- and metathorax trapezoidal, longer than wide.
<emphasis id="B905473ABD3E111B3EA8FC37FC8DFC43" bold="true" box="[749,859,964,985]" pageId="87" pageNumber="88">Abdomen:</emphasis>
Tergite IV without patches of microtrichia.
<emphasis id="B905473ABD3E111B3F79FC2CFC7BFC6E" bold="true" box="[828,941,991,1012]" pageId="87" pageNumber="88">Aedeagus:</emphasis>
Median lobe straight and slender (
<figureCitation id="134A87ADBD3E111B3F6EFC09FC57FB95" box="[811,897,1018,1039]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49d</figureCitation>
). Phallobase of median lobe asymmetrical and rounded (
<figureCitation id="134A87ADBD3E111B3F81FBE7FBCAFBB3" box="[964,1052,1044,1065]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49d</figureCitation>
). Parameres asymmetrical, left divided and apical lobe of major lobe arrow head-shaped, right shallowly divided (
<figureCitation id="134A87ADBD3E111B3F40FB97FC8AFBE3" box="[773,860,1124,1145]" captionStart="Fig" captionStartId="86.[197,229,1662,1681]" captionTargetBox="[195,1181,676,1640]" captionTargetId="figure-182@86.[178,1188,661,1652]" captionTargetPageId="86" captionText="Fig. 49. Sagola species in the gilae species-group. Habitus, dorsal views: a) S. gilae; b) S. tishechkini. Aedeagi, dorsal views: c) S. gilae, d) S. tishechkini. S. gilae, male head: e) Dorsal surface, f) Ventral surface. Scale bars ab = 1 mm, cf = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907870" httpUri="https://zenodo.org/record/4907870/files/figure.png" pageId="87" pageNumber="88">Fig. 49d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BD3E111B3E80FB8CFCDDFB53" pageId="87" pageNumber="88" type="distribution">
<paragraph id="8BCE9B28BD3E111B3E80FB8CFBB1FB0E" blockId="87.[685,1172,194,1225]" box="[709,1127,1151,1172]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3E80FB8CFC9EFB0E" bold="true" box="[709,840,1151,1172]" pageId="87" pageNumber="88">Distribution.</emphasis>
<collectingRegion id="49B555CABD3E111B3F10FB8CFC7AFB0E" box="[853,940,1151,1172]" country="New Zealand" name="Gisborne" pageId="87" pageNumber="88">Gisborne</collectingRegion>
(
<figureCitation id="134A87ADBD3E111B3FFEFB8CFBD3FB0E" box="[955,1029,1151,1172]" captionStart="Fig" captionStartId="88.[731,763,1662,1681]" captionTargetBox="[712,1189,953,1639]" captionTargetId="figure-327@88.[709,1190,948,1642]" captionTargetPageId="88" captionText="Fig. 51. Known collection localities of New Zealand Sagola species. S. gilae: black circle; S. tishechkini: triangle; S. owensae: black square." figureDoi="http://doi.org/10.5281/zenodo.4907874" httpUri="https://zenodo.org/record/4907874/files/figure.png" pageId="87" pageNumber="88">Fig. 51</figureCitation>
: triangle).
</paragraph>
<paragraph id="8BCE9B28BD3E111B3E80FB6AFCDDFB53" blockId="87.[685,1172,194,1225]" pageId="87" pageNumber="88">
<emphasis id="B905473ABD3E111B3E80FB6AFCCCFB34" bold="true" box="[709,794,1177,1198]" pageId="87" pageNumber="88">Habitat.</emphasis>
The
<typeStatus id="54CA258ABD3E111B3F11FB6AFC71FB34" box="[852,935,1177,1198]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
was collected by sifting leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>