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<document ID-DOI="http://doi.org/10.5281/zenodo.5720521" ID-GBIF-Dataset="92918ae4-0a1b-4294-862d-e726ecc071d1" ID-ISBN="978-84-96553-77-4" ID-Zenodo-Dep="5720521" checkinTime="1633654429924" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2011" docId="03C6E250FFBEF059FACAF5871E9BFA66" docLanguage="en" docName="hbmw_2_Moschidae_0336.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 2 Hoofed Mammals, Barcelona: Lynx Edicions" docTitle="Moschus berezovskii Flerov 1929" docType="treatment" docVersion="10" lastPageNumber="348" masterDocId="FFFF9A28FFBAF05CFFB1FFC61969FFC9" masterDocTitle="Moschidae" masterLastPageNumber="348" masterPageNumber="336" pageNumber="347" updateTime="1658336495735" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Moschidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2011</mods:dateIssued>
<mods:dateOther type="pubDate">2011-08-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 2 Hoofed Mammals</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>336</mods:start>
<mods:end>348</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.5720521</mods:identifier>
<mods:identifier type="GBIF-Dataset">92918ae4-0a1b-4294-862d-e726ecc071d1</mods:identifier>
<mods:identifier type="ISBN">978-84-96553-77-4</mods:identifier>
<mods:identifier type="Zenodo-Dep">5720521</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5720551" ID-GBIF-Taxon="190573076" ID-Zenodo-Dep="5720551" LSID="urn:lsid:plazi:treatment:03C6E250FFBEF059FACAF5871E9BFA66" httpUri="http://treatment.plazi.org/id/03C6E250FFBEF059FACAF5871E9BFA66" lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347">
<subSubSection box="[1403,1434,2625,2667]" pageId="4" pageNumber="347" type="multiple">
<paragraph blockId="4.[1400,2400,2625,2709]" box="[1403,1434,2625,2667]" pageId="4" pageNumber="347">
<heading box="[1403,1434,2625,2667]" pageId="4" pageNumber="347">
<figureCitation box="[1403,1434,2625,2667]" captionStart="On" captionStartId="2.[112,144,3394,3415]" captionTargetBox="[12,2754,13,3632]" captionTargetPageId="1" captionText="On following pages 3 Hrmalayan Musk-deer (Moschus Ieuoogaston, 4 Kashmır Musk-deer (Moschus cuprousl. 5 Black Musk-deer (Moschus fuscus) 6 Anhuı Musk-deer (Moschus ınhuıensısl, 7 Forest Musk-deer (Moschus berezovskıı)" figureDoi="http://doi.org/10.5281/zenodo.6514072" httpUri="https://zenodo.org/record/6514072/files/figure.png" pageId="4" pageNumber="347">7.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1451,1810,2625,2667]" pageId="4" pageNumber="347" type="vernacular_names">
<paragraph blockId="4.[1400,2400,2625,2709]" box="[1451,1810,2625,2667]" pageId="4" pageNumber="347">
<heading box="[1451,1810,2625,2667]" pageId="4" pageNumber="347">
<vernacularName box="[1451,1810,2625,2667]" pageId="4" pageNumber="347">Forest Musk-deer</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1861,2230,2625,2667]" pageId="4" pageNumber="347" type="nomenclature">
<paragraph blockId="4.[1400,2400,2625,2709]" box="[1861,2230,2625,2667]" pageId="4" pageNumber="347">
<heading box="[1861,2230,2625,2667]" pageId="4" pageNumber="347">
<taxonomicName authorityName="Flerov" authorityYear="1929" box="[1861,2230,2625,2667]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="347" phylum="Chordata" rank="species" species="berezovskii">
<emphasis box="[1861,2230,2625,2667]" italics="true" pageId="4" pageNumber="347">Moschus berezovskii</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1402,2399,2685,2706]" pageId="4" pageNumber="347" type="vernacular_names">
<paragraph blockId="4.[1400,2400,2625,2709]" box="[1402,2399,2685,2706]" pageId="4" pageNumber="347">
<heading box="[1402,2399,2685,2706]" pageId="4" pageNumber="347">
<emphasis bold="true" box="[1402,1478,2685,2706]" pageId="4" pageNumber="347">French:</emphasis>
<vernacularName box="[1488,1687,2685,2706]" pageId="4" pageNumber="347">Porte-musc de forét</vernacularName>
/
<emphasis bold="true" box="[1708,1799,2685,2706]" pageId="4" pageNumber="347">German:</emphasis>
<vernacularName box="[1809,1996,2685,2706]" pageId="4" pageNumber="347">China-Moschustier</vernacularName>
/
<emphasis bold="true" box="[2017,2108,2685,2706]" pageId="4" pageNumber="347">Spanish:</emphasis>
<vernacularName box="[2117,2179,2685,2706]" pageId="4" pageNumber="347">Ciervo</vernacularName>
almizclero de bosque
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="347" type="reference_group">
<paragraph blockId="4.[2017,2603,2759,3177]" box="[2017,2600,2759,2784]" pageId="4" pageNumber="347">
<emphasis bold="true" box="[2017,2171,2759,2784]" pageId="4" pageNumber="347">Taxonomy.</emphasis>
<taxonomicName authority="Flerov, 1929" authorityName="Flerov" authorityYear="1929" box="[2181,2596,2759,2784]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="347" phylum="Chordata" rank="species" species="berezovskii">Moschus berezovskii Flerov, 1929</taxonomicName>
,
</paragraph>
<paragraph blockId="4.[2017,2603,2759,3177]" box="[2018,2564,2798,2823]" pageId="4" pageNumber="347">
<materialsCitation ID-GBIF-Occurrence="3761237307" box="[2018,2564,2798,2823]" pageId="4" pageNumber="347">
Ho-tzi-how Pass, near Lungan,
<collectingRegion box="[2447,2560,2798,2823]" country="China" name="Sichuan" pageId="4" pageNumber="347">Sichuan</collectingRegion>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="347" type="discussion">
<paragraph blockId="4.[2017,2603,2759,3177]" pageId="4" pageNumber="347">
Flerov, when he described this species, regarded it as an enigmatic species of apparently restricted distribution in the alpine zone of
<collectingRegion box="[2129,2243,2947,2980]" country="China" name="Sichuan" pageId="4" pageNumber="347">Sichuan</collectingRegion>
. It was not until the 1960s that it was shown that this is not in fact an alpine zone species at all, but is a forest animal, widespread in the forested areas of south-eastern
<collectingCountry box="[2266,2352,3104,3137]" name="China" pageId="4" pageNumber="347">China</collectingCountry>
. Four subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.5720543" ID-Zenodo-Dep="5720543" httpUri="https://zenodo.org/record/5720543/files/figure.png" inLine="true" lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347" targetBox="[1400,1992,2761,3175]" targetPageId="4">
<paragraph blockId="4.[1402,2602,3188,3452]" box="[1402,1801,3188,3217]" pageId="4" pageNumber="347">
<emphasis bold="true" box="[1402,1801,3188,3217]" pageId="4" pageNumber="347">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="4.[1402,2602,3188,3452]" pageId="4" pageNumber="347">
<taxonomicName authority="Flerov, 1929" authorityName="Flerov" authorityYear="1929" box="[1403,1785,3230,3255]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="347" phylum="Chordata" rank="subSpecies" species="berezovskii" subSpecies="berezovskii">M.b.berezovskiiFlerov,1929—SNingxia,SGansu,SEQuinghai,SShaanxi,WHenan,Sichuan,andSXizang;between500and2500m.</taxonomicName>
</paragraph>
<paragraph blockId="4.[1402,2602,3188,3452]" box="[1404,2482,3305,3334]" pageId="4" pageNumber="347">
<taxonomicName authority="Wang &amp; Li, 1993" authorityName="Wang &amp; Li" authorityYear="1993" box="[1404,1867,3305,3334]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="347" phylum="Chordata" rank="subSpecies" species="berezovskii" subSpecies="bijiangensis">M.b.bijiangensisWang&amp;Li,1993—NWYunnantoSEXizang(ZayuCounty).</taxonomicName>
</paragraph>
<paragraph blockId="4.[1402,2602,3188,3452]" pageId="4" pageNumber="347">
<taxonomicName authority="Dao, 1969" authorityName="Dao" authorityYear="1969" box="[1404,1759,3340,3373]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="347" phylum="Chordata" rank="subSpecies" species="berezovskii" subSpecies="caobangis">M. b. caobangis Dao, 1969</taxonomicName>
— S Yunnan (Mile County &amp; Nujiang Langcang), Guangxi, Guangdong, and N
<collectingCountry box="[1677,1795,3379,3412]" name="Vietnam" pageId="4" pageNumber="347">Vietnam</collectingCountry>
(
<collectingRegion box="[1818,2091,3379,3412]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="347">Cao Bang Province</collectingRegion>
); between
<quantity box="[2240,2307,3379,3412]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="347" unit="m" value="50.0">50 m</quantity>
and
<quantity box="[2377,2463,3379,3412]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="347" unit="m" value="400.0">400 m</quantity>
. Presence uncertain in N
<collectingCountry box="[1618,1684,3423,3452]" name="Laos" pageId="4" pageNumber="347">Laos</collectingCountry>
.
</paragraph>
<paragraph blockId="5.[184,1390,278,1451]" pageId="5" pageNumber="348">
<taxonomicName authority="Wang &amp; Ma, 1993" authorityName="Wang &amp; Ma" authorityYear="1993" box="[190,684,278,311]" class="Mammalia" family="Moschidae" genus="Moschus" kingdom="Animalia" order="Artiodactyla" pageId="5" pageNumber="348" phylum="Chordata" rank="subSpecies" species="berezovskii" subSpecies="yunguiensis">M. b. yunguiensis Wang &amp; Ma, 1993</taxonomicName>
— from the Yunnan-Guizhou plateau margins NE to W
<collectingRegion box="[264,351,321,350]" country="China" name="Hubei" pageId="5" pageNumber="348">Hubei</collectingRegion>
(Yichang).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="description">
<paragraph blockId="5.[184,1390,278,1451]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[188,442,366,391]" pageId="5" pageNumber="348">Descriptive notes.</emphasis>
Head-body 70-80 cm, tail 3-4 cm, height at shoulder less than
<quantity box="[190,276,396,429]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="348" unit="cm" value="50.0">50 cm</quantity>
; weight 6-9 kg. Skull length is 14-15 cm in the nominotypical subspecies, but only 13.2-14.
<quantity box="[378,448,436,469]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="348" unit="cm" value="5.0">5 cm</quantity>
in subspecies bijiangensis and yunguiensis, and only 12:5-13-5 mm in caobangis, the smallest of all musk-deer. Hair bases are short, gray-white; the throat has three wide longitudinal stripes, white to orange in color, the lateral pair running from the jaw angles to the brisket. The central stripe is shorter, and all three may sometimes be broken. The haunch is yellower, but the rump is nearly black; the limbs are dark down the front surface. The underside is yellow. The ears are orange at the base, black at the tip, and white inside. The hair is 73-87 mm long on the withers, 45-68 mm on the rump, varying according to season. The facial skeleton is relatively short; the lachrymalis shorter than itis high. The limbs are relatively short: metacarpal length 73-89 mm, metatarsal 103-120 mm. The metatarsal is 135-139% of the metacarpal, so the hindlimb is relatively longer than in most other taxa. The metapodials are relatively short (the metatarsal only 80-81% of the skull length, the metacarpal only 59%) and robust, the width of the lower head of the metacarpal being more than 18% ofits length, of the metatarsal more than 15-5%. M. b. caobangis is very pale in color, limbs dark only from knee and hocks downward, but buttocks and ear tips are black. Hair shorter, 21-30 mm on withers, 38-47 mm on rump. M. b. byiangensis 1s paler than nominotypical M. b. berezouskii, fulvous-brown, with a grayish-white ventral surface; toothrow averaging slightly shorter. M. b. yunguiensis is somewhat smaller than nominotypical M. b. berezouski.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="biology_ecology">
<paragraph blockId="5.[184,1390,278,1451]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[186,297,1187,1216]" pageId="5" pageNumber="348">Habitat.</emphasis>
Lives predominantly in forests with mixed coniferous and broadleaved trees, usually below
<quantity box="[380,483,1222,1255]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="348" unit="m" value="3500.0">3500 m</quantity>
, and often almost at sea level. In pure broadleafforest its population density is lower, and in shrub lowerstill; it goes into bamboo forest, but does not go into grassy areas or the “dwarf bush zone.” It is always found on relatively steep slopes, and does not like slopes under 10°. On northern slopes,it is found in forests between
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and
<quantity box="[492,596,1379,1412]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="348" unit="m" value="2100.0">2100 m</quantity>
, whereas on south-facing slopes,its highest density is in woodlands between
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and
<quantity box="[645,749,1418,1451]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="5" pageNumber="348" unit="m" value="2450.0">2450 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="food_feeding">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1461,1720,282,315]" pageId="5" pageNumber="348">Food and Feeding.</emphasis>
They feed on twigs, shoots, lichens, moss, grasses, and leaves. Their leaping ability allows them to forage on the lower branches oftrees.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="breeding">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1460,1594,360,393]" pageId="5" pageNumber="348">Breeding.</emphasis>
Dominant females (dominance is calculated by the frequency of one animal supplanting others) give birth from April 25 to May 25, low-ranking femaleslater, from May 15 to June 25. Birthweight is less than in other musk-deer, ranging from 350-558 g. Twinning is extremely high: 137 twin litters and 59 singletons are recorded. Juvenile mortality is high, about 50%, up to age two, but this differs according to the dominance rank of the mother. The survival rate of the offspring of dominant females is 100%; of females of the second rank, 56%; of the third rank, 43%; and of the fourth rank, 25%. In general, 30% of musk-deer survive to age 6, 20% to age 10%, just 10% to age 15.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="activity">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1458,1696,675,708]" pageId="5" pageNumber="348">Activity patterns.</emphasis>
They are most active between dusk and dawn, but alternately rest and feed.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="biology_ecology">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1460,2159,754,787]" pageId="5" pageNumber="348">Movements, Home range and Social organization.</emphasis>
The social organization, territoriality, etc., of this species are poorly known. The juvenile sex ratio is approximately 1:1, but after that males predominate until old age. The sex ratio in young adults (2-5-9 years) is 1-77:1, and in older adults (9-14-5 years) 2-8:1, whereas in aged individuals (14-5-20 years)itis 1:2. Young adults, at about 6-8 years of age, are the prime breeders. The mean generation length is 6-5-10 years.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="conservation">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1459,1810,990,1023]" pageId="5" pageNumber="348">Status and Conservation.</emphasis>
CITES Appendix II. Classified as Endangered on The [UCN Red List. By the early 1990s a decline from earlier levels was noticeable, and there had almost certainly been local extinctions; it remains relatively numerous only in western
<collectingRegion box="[1459,1570,1107,1140]" country="China" name="Sichuan" pageId="5" pageNumber="348">Sichuan</collectingRegion>
and north-west
<collectingRegion box="[1796,1908,1107,1140]" country="China" name="Yunnan" pageId="5" pageNumber="348">Yunnan</collectingRegion>
. Population sizes are very difficult to even estimate in musk-deer, but Prof Sheng Helin, from a lifetimes experience, suggested that in 1992 there would be about
<quantity box="[1781,1927,1187,1220]" metricMagnitude="3" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="348" unit="in" value="200000.0">200,000 in</quantity>
the whole of
<collectingCountry box="[2132,2218,1187,1220]" name="China" pageId="5" pageNumber="348">China</collectingCountry>
, split of course among at least five species, and it is evident that the population continues to decline. In
<collectingCountry box="[2480,2602,1225,1258]" name="Vietnam" pageId="5" pageNumber="348">Vietnam</collectingCountry>
, Do Tuoc estimated the population to have been some
<quantity box="[2176,2262,1265,1298]" metricMagnitude="0" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="348" unit="in" value="200.0">200 in</quantity>
the late 1990s, but here too there has been a decline.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="348" type="bibRefCitation_list">
<paragraph blockId="5.[1456,2665,282,1458]" pageId="5" pageNumber="348">
<emphasis bold="true" box="[1459,1612,1351,1376]" pageId="5" pageNumber="348">Bibliography.</emphasis>
Guo Jian et al. (2001), Hu Zhongjun et al. (2007), Wang Yingxiang &amp; Harris (2008), Wang Yingxlang et al. (1993), Wong Yu et al. (2006), Xu Zhenggiang &amp; Xu Hongfa (2002), Yang Qisen, Hu Jinchu &amp; Peng Jitai (1990), Yang Qisen, Meng Xiuxiang et al. (2003),
</paragraph>
</subSubSection>
</treatment>
</document>