treatments-xml/data/03/C6/3E/03C63E67680FFFA5FF000641FE3200B6.xml

<document id="086FF6594ADA037918DED8F01D1B4037" ID-DOI="10.5281/zenodo.5328899" ID-ISSN="0253-116X" ID-Zenodo-Dep="5328899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630160147779" checkinUser="carolina" docAuthor="Casale, A., D. W &amp; P, Antalya" docDate="2012" docId="03C63E67680FFFA5FF000641FE3200B6" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1111-1127.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Laemostenus (Laemostenus) phoenicius Casale &amp; D.W &amp; P 2012, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1116" masterDocId="FFFF461F680CFFA0FFB2033BFFE2035B" masterDocTitle="New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)" masterLastPageNumber="1127" masterPageNumber="1111" pageNumber="1114" updateTime="1699169291185" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8074F8CBAF1ED9578DEC1476D7392842">New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="C42101D6EB9628B9EB0B19C4E2E47FB7">D. W</mods:namePart>
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<taxonomicName id="4C6FF4F2680FFFA3FF000641FDB706D4" ID-CoL="8MKK3" authority="Casale &amp; D.W &amp; P, 2012" authorityName="Casale &amp; D.W &amp; P" authorityYear="2012" box="[178,597,1401,1423]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1114" phylum="Arthropoda" rank="species" species="phoenicius" status="sp. nov." subGenus="Laemostenus">
<emphasis id="B91B5363680FFFA3FF000641FEDE06D4" bold="true" box="[178,316,1402,1423]" italics="true" pageId="3" pageNumber="1114">Laemostenus</emphasis>
(
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) 
<emphasis id="B91B5363680FFFA3FE510642FDB706D4" bold="true" box="[483,597,1401,1423]" italics="true" pageId="3" pageNumber="1114">phoenicius</emphasis>
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<materialsCitation id="3B07852C680FFFA3FF000692FE67068E" collectingDate="1997-04-13" collectionCode="T" collectorName="Heinz" country="Lebanon" elevation="800" location="Mayfouq" pageId="3" pageNumber="1114" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED7E17B4680FFFA3FF000692FF2206E6" box="[178,192,1449,1469]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="1114">T</collectionCode>
y p e m a t e r i a l: 
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, labelled: &quot; 
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: 
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<quantity id="4C972294680FFFA3FC940692FC8206E6" box="[806,864,1449,1469]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="1114" unit="m" value="800.0">
<elevation id="00426842680FFFA3FC940692FC8206E6" box="[806,864,1449,1469]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="1114" unit="m" value="800.0">800 m</elevation>
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4/ 
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<collectingDate id="EF955059680FFFA3FCC80692FC3F06E6" box="[890,989,1449,1469]" pageId="3" pageNumber="1114" value="1997-04-13">13/IV/1997</collectingDate>
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(ne. Jbaїl) 
<collectorName id="269AEAA7680FFFA3FF6D06FAFEF1068E" box="[223,275,1473,1493]" pageId="3" pageNumber="1114">Heinz</collectorName>
leg. &quot; (cHE)
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. 
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<typeStatus id="54D431D3680FFFA3FE3C06FAFE04068E" box="[398,486,1473,1493]" pageId="3" pageNumber="1114" type="paratype">Paratypes</typeStatus>
: 1 with same data but handwritten on backside of the label: 11.IV. (cHE)
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; 
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; 
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1, 2: &quot; 
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E Saida 
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ZW. 
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u. 
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5/ 
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<collectingDate id="EF955059680FFFA3FE8406CAFE7A055E" box="[310,408,1521,1541]" pageId="3" pageNumber="1114" value="2003-11-17">17.XI.2003</collectingDate>
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<collectorName id="269AEAA7680FFFA3FE2F06CAFE33055E" box="[413,465,1521,1541]" pageId="3" pageNumber="1114">Heinz</collectorName>
leg.&quot; (cCa, cHE)
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.
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<paragraph id="8BD08F71680FFFA3FF00052FFBDF051D" blockId="3.[178,1086,1555,1606]" pageId="3" pageNumber="1114">
D i a g n o s i s: A small sized 
<taxonomicName id="4C6FF4F2680FFFA3FD9B052FFCB50572" authority="(Laemostenus)" box="[553,855,1555,1577]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1114" phylum="Arthropoda" rank="subGenus" subGenus="Laemostenus">
<emphasis id="B91B5363680FFFA3FD9B052FFD4F0572" box="[553,685,1556,1577]" italics="true" pageId="3" pageNumber="1114">Laemostenus</emphasis>
(
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)
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species of the 
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<emphasis id="B91B5363680FFFA3FB9A052FFECE051D" italics="true" pageId="3" pageNumber="1114">L. quadricollis</emphasis>
</taxonomicName>
species group (sensu 
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), mostly characterised by the uniformly
</paragraph>
<paragraph id="8BD08F716808FFA4FF0003E0FC290255" blockId="4.[178,1086,219,270]" pageId="4" pageNumber="1115">
reddish colour with brown-bluish disc of elytra, parallel-sided tempora, small eyes, and the tarsal claws with reduced, small teeth on the internal side. Habitus 
<figureCitation id="135493F46808FFA4FCCE03C3FC290255" box="[892,971,248,270]" captionStart="Figs 22-25" captionStartId="16.[178,218,1506,1526]" captionTargetBox="[266,976,232,1475]" captionTargetId="figure-64@16.[178,1085,212,1495]" captionTargetPageId="16" captionText="Figs 22-25: Laemostenus, habitus. (22) L. croyi nov.sp., paratype. (23) L. phoenicius nov.sp., paratype (Maїfouq). (24) L. eggeri nov.sp., paratype. (25) L. bellicosus nov.sp., holotype (various magnifications)." figureDoi="http://doi.org/10.5281/zenodo.5328905" httpUri="https://zenodo.org/record/5328905/files/figure.png" pageId="4" pageNumber="1115">Fig. 23.</figureCitation>
</paragraph>
<paragraph id="8BD08F716808FFA4FF000225FD6C0236" blockId="4.[178,1085,285,366]" pageId="4" pageNumber="1115">
E t y m o l o g y: Referring on the 
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locality of the new species, 
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, from which the Phoenician people originated and extended to one of the most important and ancient civilizations in the Mediterranean area.
</paragraph>
<paragraph id="8BD08F716808FFA4FF000245FF0A02EB" blockId="4.[178,1085,381,432]" pageId="4" pageNumber="1115">
D e s c r i p t i o n: Body small: TL: 11.5-12.0 (
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) mm; L: 12.5-13.0 (
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) mm).
</paragraph>
<paragraph id="8BD08F716808FFA4FF000284FDC5014B" blockId="4.[178,1085,447,528]" pageId="4" pageNumber="1115">Colour: Uniformly reddish, markedly depigmented; disc of elytra (except the sutural interval) darker, brownish with slight bluish-violet reflection at oblique light. Surface moderately shining, elytra sericeous.</paragraph>
<paragraph id="8BD08F716808FFA4FF000124FE3C0108" blockId="4.[178,1085,543,595]" pageId="4" pageNumber="1115">Microsculpure: Head and pronotum with shallow, transversal microlines, elytra with distinct, isodiametric meshes.</paragraph>
<paragraph id="8BD08F716808FFA4FF000159FD40018B" blockId="4.[178,1085,610,720]" pageId="4" pageNumber="1115">Head elongate but robust, thickened; dorsal surface smooth; frons convex, tempora parallel-sided; eyes small, much shorter than tempora, slightly prominent laterally; frontal impressions small, short, slightly impressed; antennae long, if stretched backwards exceeding by four antennomeres the elytral base.</paragraph>
<paragraph id="8BD08F716808FFA4FF0001DBFCB100FF" blockId="4.[178,1087,735,933]" pageId="4" pageNumber="1115">Pronotum subquadrate cordiform (PL/PW: 0.97), its lateral sides slightly reflexed in the posterior half, briefly sinuate anteriorly to the basolateral angles, which are rectangular; anterolateral angles acutely prominent; base moderately oblique at sides, superficially beaded. Disc depressed, smooth, with sparse and shallow transverse wrinkles close to the median furrow; lateral furrow deeper in the anterior half; basal impressions wide, moderately deep, each with a few, deep punctures extended to the basal area and the lateral furrows; anterolateral and basolateral setiferous punctures present.</paragraph>
<paragraph id="8BD08F716808FFA4FF00008FFD9A0764" blockId="4.[178,1085,948,1087]" pageId="4" pageNumber="1115">Elytra ovate, short (EL/EW: 1.70), widened in the posterior third, subconvex but depressed on the disc. Base narrow, almost straight; basal ridge incavate; humeral tooth absent, shoulders obtusely rounded but evident. Striae very deep, shallowly punctured; intervals flat, smooth. Chaetotaxy: Basal pore present; umbilicate series with 17-18 setiferous punctures; 1-2 setae at apex of stria 7.</paragraph>
<paragraph id="8BD08F716808FFA4FF000774FCB2064F" blockId="4.[178,1085,1102,1300]" pageId="4" pageNumber="1115">Legs long and slender; profemora on ventral side longitudinally shallowly concave for entire length, its outer side with an oblique series of setae, one of them reaching the apical third of the outer margin, which is smooth; mesotibiae straight; metatibiae each with apical group of short, sparse yellow-reddish setae. Tarsomeres narrow, thin, with dorsal pubescence short and sparse. Males with fore tarsomeres 1-3 moderately dilated and with ventral, biseriate adhesive vestiture. Tarsal claws with a series of small teeth, reduced in number (4), along the basal half of the internal margin.</paragraph>
<paragraph id="8BD08F716808FFA4FF000618FDD806CA" blockId="4.[178,1086,1315,1425]" pageId="4" pageNumber="1115">Male genitalia and abdominal segment IX in male as in Figs. 2-5. Aedeagus small; median lobe slightly curved, its apex short, in dorsal aspect slightly asymmetric, subtruncate and emarginate distally; right paramere very short, subtruncate apically; left paramere with reduced apical membranous lobe.</paragraph>
<paragraph id="8BD08F716808FFA4FF00069BFE1F06ED" blockId="4.[178,509,1440,1462]" box="[178,509,1440,1462]" pageId="4" pageNumber="1115">Female genitalia: Not examined.</paragraph>
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D i s t r i b u t i o n a n d h a b i t a t: Known so far from two localities in 
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, in a range of altitude from 
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a.s.l. The few specimens of the 
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series were collected in different localities and in two different years. The adaptive features of this species indicate a subterranean way of life, confirmed by the scarcity of individuals and its living condition. The localities are far apart, but of the same geological character: karstic limestone rock terraces with fissures. Grass grows on the horizontal surface up to the fissures which are filled with humus and detritus and very humid, at least after the thaw (in Maїfouq snow was still lying on the ground). The specimens were captured in traps on the horizontal surface and each placed near to a fissure. Today the biotope at Maїfouq is completely destroyed being now used for waste disposal. The locality near Saida is thought to still exist (W. Heinz, pers. comm.).
</paragraph>
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C o m p a r i s o n s: Close to 
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<emphasis id="B91B53636809FFA5FDBB028BFDF5029E" box="[521,535,432,453]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
. (
<emphasis id="B91B53636809FFA5FD83028BFDA4029E" box="[561,582,432,453]" italics="true" pageId="5" pageNumber="1116">L.</emphasis>
) 
<emphasis id="B91B53636809FFA5FDE80294FD3D029E" box="[602,735,431,453]" italics="true" pageId="5" pageNumber="1116">antonrichteri</emphasis>
<bibRefCitation id="EFFEF2806809FFA5FD590294FC9F029E" author="CASALE A" box="[747,893,431,453]" pageId="5" pageNumber="1116" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
</taxonomicName>
, from which it is distinguished by the elytral disc darker, brownish with slight bluish-violet reflection, the elytra shorter, ovate, and by elytral intervals flat and the tarsal claws with reduced but evident denticulation on the internal side in the basal half.
</paragraph>
<paragraph id="8BD08F716809FFA5FF000116FC5701E3" blockId="5.[178,1085,557,696]" pageId="5" pageNumber="1116">
R e m a r k s: The general depigmentation of integument, the reduced eyes, the slender and elongate appendages, and the reduced denticulation of tarsal claws, clearly indicate in this species marked adaptive features to a subterranean way of life, as in three other species of the 
<emphasis id="B91B53636809FFA5FEF701BFFE5D01C1" box="[325,447,644,666]" italics="true" pageId="5" pageNumber="1116">quadricollis</emphasis>
species group: 
<taxonomicName id="4C6FF4F26809FFA5FDD201BEFC1201C1" authority="SCHATZMAYR 1936" authorityName="SCHATZMAYR" authorityYear="1936" box="[608,1008,644,666]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="aegyptiacus" subGenus="Laemostenus">
<emphasis id="B91B53636809FFA5FDD201BEFD8F01C1" box="[608,621,645,666]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
. (
<emphasis id="B91B53636809FFA5FD3101BEFD7A01C1" box="[643,664,645,666]" italics="true" pageId="5" pageNumber="1116">L.</emphasis>
) 
<emphasis id="B91B53636809FFA5FD1A01BEFCC301C1" box="[680,801,645,666]" italics="true" pageId="5" pageNumber="1116">aegyptiacus</emphasis>
SCHATZMAYR 1936
</taxonomicName>
, 
<emphasis id="B91B53636809FFA5FC4E01BEFBE801C1" box="[1020,1034,645,666]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
. (
<emphasis id="B91B53636809FFA5FB9201BEFBCC01C1" box="[1056,1070,645,666]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
.) 
<emphasis id="B91B53636809FFA5FF000199FF0C01E3" box="[178,238,674,696]" italics="true" pageId="5" pageNumber="1116">heinzi</emphasis>
<bibRefCitation id="EFFEF2806809FFA5FF470199FE9D01E3" author="CASALE A" box="[245,383,674,696]" pageId="5" pageNumber="1116" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
and 
<taxonomicName id="4C6FF4F26809FFA5FE020198FCEF01E3" authority="CASALE 1988" authorityName="CASALE" authorityYear="1988" box="[432,781,674,696]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="antonrichteri" subGenus="Laemostenus">
<emphasis id="B91B53636809FFA5FE020198FE5C01E3" box="[432,446,675,696]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
. (
<emphasis id="B91B53636809FFA5FE610198FE0301E3" box="[467,481,675,696]" italics="true" pageId="5" pageNumber="1116">L</emphasis>
.) 
<emphasis id="B91B53636809FFA5FE450199FD9E01E3" box="[503,636,674,696]" italics="true" pageId="5" pageNumber="1116">antonrichteri</emphasis>
<bibRefCitation id="EFFEF2806809FFA5FD300199FCEF01E3" author="CASALE A" box="[642,781,674,696]" pageId="5" pageNumber="1116" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFFEF2806809FFA5FCA90199FC4801E3" author="CASALE A" box="[795,938,674,696]" pageId="5" pageNumber="1116" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
).
</paragraph>
<paragraph id="8BD08F716809FFA5FF0001FCFE3900D6" blockId="5.[178,1085,711,909]" pageId="5" pageNumber="1116">
A preliminary, superficial examination of the individuals of this new 
<taxonomicName id="4C6FF4F26809FFA5FC3401F3FF3B01A1" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91B53636809FFA5FC3401F3FBE80186" box="[902,1034,712,733]" italics="true" pageId="5" pageNumber="1116">Laemostenus</emphasis>
species
</taxonomicName>
induced one of the authors (A.C.) to suspect the re-discovery of the enigmatic 
<taxonomicName id="4C6FF4F26809FFA5FF000039FCD70043" authority="(LA BRULERIE 1875)" baseAuthorityName="LA BRULERIE" baseAuthorityYear="1875" box="[178,821,770,792]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="libanensis" subGenus="Antisphodrus">
<emphasis id="B91B53636809FFA5FF000039FED4004C" box="[178,310,770,791]" italics="true" pageId="5" pageNumber="1116">Laemostenus</emphasis>
(
<emphasis id="B91B53636809FFA5FEF80039FE330043" box="[330,465,770,792]" italics="true" pageId="5" pageNumber="1116">Antisphodrus</emphasis>
) 
<emphasis id="B91B53636809FFA5FE570039FDA90043" box="[485,587,770,792]" italics="true" pageId="5" pageNumber="1116">libanensis</emphasis>
(LA BRÛLERIE 1875)
</taxonomicName>
, described from remains collected at high altitude (
<quantity id="4C9722946809FFA5FE770024FDFA006E" box="[453,536,799,821]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="1116" unit="m" value="2600.0">2600 m</quantity>
) in the Djebel Sannin, under a big stone, and never found again (
<bibRefCitation id="EFFEF2806809FFA5FEF10007FE3A0009" author="CASALE A" box="[323,472,828,850]" pageId="5" pageNumber="1116" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
). A subsequent, careful examination of these individuals showed that they were not 
<taxonomicName id="4C6FF4F26809FFA5FE740061FD940034" baseAuthorityName="LA BRULERIE" baseAuthorityYear="1875" box="[454,630,857,879]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="libanensis" subGenus="Antisphodrus">
<emphasis id="B91B53636809FFA5FE740061FE390034" box="[454,475,858,879]" italics="true" pageId="5" pageNumber="1116">L.</emphasis>
(
<emphasis id="B91B53636809FFA5FE5B0062FDE20034" box="[489,512,857,879]" italics="true" pageId="5" pageNumber="1116">A.</emphasis>
) 
<emphasis id="B91B53636809FFA5FDBD0062FD940034" box="[527,630,857,879]" italics="true" pageId="5" pageNumber="1116">libanensis</emphasis>
</taxonomicName>
but a new species here described, belonging to 
<taxonomicName id="4C6FF4F26809FFA5FF7E0043FEB200D6" box="[204,336,888,909]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91B53636809FFA5FF7E0043FEB200D6" box="[204,336,888,909]" italics="true" pageId="5" pageNumber="1116">Laemostenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228EE186809FFA5FEE4004CFE3500D6" box="[342,471,887,909]" pageId="5" pageNumber="1116" sensu="stricto">sensu stricto</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8BD08F716809FFA5FF0000A7FE3200B6" blockId="5.[178,1085,924,1005]" pageId="5" pageNumber="1116">
The 
<taxonomicName id="4C6FF4F26809FFA5FF5000A6FDE100E9" authority="(Laemostenus)" box="[226,515,924,946]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="subGenus" subGenus="Laemostenus">
<emphasis id="B91B53636809FFA5FF5000A6FE8400E9" box="[226,358,925,946]" italics="true" pageId="5" pageNumber="1116">Laemostenus</emphasis>
(
<emphasis id="B91B53636809FFA5FECA00A6FE1E00E9" box="[376,508,925,946]" italics="true" pageId="5" pageNumber="1116">Laemostenus</emphasis>
)
</taxonomicName>
species of the 
<taxonomicName id="4C6FF4F26809FFA5FD1700A6FCDE00E9" baseAuthorityName="REDTENBACHER" baseAuthorityYear="1843" box="[677,828,924,946]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1116" phylum="Arthropoda" rank="species" species="quadricollis">
<emphasis id="B91B53636809FFA5FD1700A6FCDE00E9" box="[677,828,924,946]" italics="true" pageId="5" pageNumber="1116">L. quadricollis</emphasis>
</taxonomicName>
species group, all localized in reduced range of distribution in the Near and Middle East, can be identified by the following operative key.
</paragraph>
</subSubSection>
</treatment>
</document>