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<document id="921106F34A2CE6B6EBB728740B769AD7" ID-DOI="10.5281/zenodo.5328899" ID-ISSN="0253-116X" ID-Zenodo-Dep="5328899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630160147779" checkinUser="carolina" docAuthor="Casale, A., D. W &amp; P, Antalya" docDate="2012" docId="03C63E67680AFFA7FF000705FE84057F" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1111-1127.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Laemostenus (Antisphodrus) skalei Casale &amp; D.W &amp; P 2012, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1118" masterDocId="FFFF461F680CFFA0FFB2033BFFE2035B" masterDocTitle="New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)" masterLastPageNumber="1127" masterPageNumber="1111" pageNumber="1117" updateTime="1699169291185" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9B7C4E5ED25F1D01AD7DFB93C434E707">New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="3EF8788A53438D15B11DF513D4F56BC4">Casale, A.</mods:namePart>
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<mods:namePart id="F4A7F14DAFEC8CCBA0082115DBE2212B">D. W</mods:namePart>
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<mods:namePart id="18D6CC987860F0E04CB9276A2A2CDBEE">P, Antalya</mods:namePart>
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<mods:title id="DE266D44CC153F8433B89427B3F0292E">Linzer biologische Beiträge</mods:title>
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<mods:date id="30DD1B7FDF9DA0EFDE2465C97D61207D">2012</mods:date>
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<mods:number id="0A7F977AA07AB522CE325F35F55C9D60">2012-12-28</mods:number>
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<mods:identifier id="D438BD0300F45DD4BB465CF10136786F" type="DOI">10.5281/zenodo.5328899</mods:identifier>
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<treatment id="03C63E67680AFFA7FF000705FE84057F" LSID="urn:lsid:plazi:treatment:03C63E67680AFFA7FF000705FE84057F" httpUri="http://treatment.plazi.org/id/03C63E67680AFFA7FF000705FE84057F" lastPageId="7" lastPageNumber="1118" pageId="6" pageNumber="1117">
<subSubSection id="C375DCFA680AFFA6FF000705FBEF07C2" pageId="6" pageNumber="1117" type="nomenclature">
<paragraph id="8BD08F71680AFFA6FF000705FD97070F" blockId="6.[178,629,1085,1108]" box="[178,629,1085,1108]" pageId="6" pageNumber="1117">
<heading id="D098381D680AFFA6FF000705FD97070F" box="[178,629,1085,1108]" fontSize="8" level="2" pageId="6" pageNumber="1117" reason="2">
<emphasis id="B91B5363680AFFA6FF000705FD97070F" bold="true" box="[178,629,1085,1108]" pageId="6" pageNumber="1117">
<taxonomicName id="4C6FF4F2680AFFA6FF000705FDC00708" ID-CoL="8MDZZ" authority="Casale &amp; D.W &amp; P, 2012" authorityName="Casale &amp; D.W &amp; P" authorityYear="2012" box="[178,546,1085,1108]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1117" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="B91B5363680AFFA6FF000705FEDE0708" bold="true" box="[178,316,1086,1107]" italics="true" pageId="6" pageNumber="1117">Laemostenus</emphasis>
(
<emphasis id="B91B5363680AFFA6FEF80706FE340708" bold="true" box="[330,470,1085,1107]" italics="true" pageId="6" pageNumber="1117">Antisphodrus</emphasis>
)
<emphasis id="B91B5363680AFFA6FE540706FDC00708" bold="true" box="[486,546,1085,1107]" italics="true" pageId="6" pageNumber="1117">skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228EE18680AFFA6FD9A0705FD97070F" box="[552,629,1086,1108]" pageId="6" pageNumber="1117" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD08F71680AFFA6FF000756FBEF07C2" blockId="6.[178,1086,1133,1177]" pageId="6" pageNumber="1117">
<materialsCitation id="3B07852C680AFFA6FF000756FBEF07C2" collectingDate="2010-05-02" collectionCode="T" collectorName="A. Skale" elevation="880" latitude="36.52722" location="Kheiroud Kenar forest" longLatPrecision="19" longitude="51.646114" municipality="Now Shahr" pageId="6" pageNumber="1117" specimenCount="1" stateProvince="Mazandaran" typeStatus="holotype">
<collectionCode id="ED7E17B4680AFFA6FF000756FF2207DA" box="[178,192,1133,1153]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="1117">T</collectionCode>
y p e m a t e r i a l:
<typeStatus id="54D431D3680AFFA6FE060756FDE707DA" box="[436,517,1133,1153]" pageId="6" pageNumber="1117" type="holotype">Holotype</typeStatus>
, labelled: &quot;N-Iran: Prov.
<collectingRegion id="49AB4193680AFFA6FC930756FC6D07DA" box="[801,911,1133,1153]" country="Iran" name="Mazandaran" pageId="6" pageNumber="1117">Mazandaran</collectingRegion>
, vic.
<collectingMunicipality id="6BB4150B680AFFA6FC7F0756FBD807DA" box="[973,1082,1133,1153]" pageId="6" pageNumber="1117">Now Shahr</collectingMunicipality>
,
<location id="8EB0D9AA680AFFA6FF6D07BEFE4007C2" LSID="urn:lsid:plazi:treatment:03C63E67680AFFA7FF000705FE84057F:8EB0D9AA680AFFA6FF6D07BEFE4007C2" box="[223,418,1157,1177]" latitude="36.52722" longLatPrecision="19" longitude="51.646114" municipality="Now Shahr" name="Kheiroud Kenar forest" pageId="6" pageNumber="1117" stateProvince="Mazandaran">Kheiroud Kenar forest</location>
<geoCoordinate id="EE5BE9B6680AFFA6FE1507BEFDF107C2" box="[423,531,1157,1177]" degrees="36" direction="north" minutes="31" orientation="latitude" pageId="6" pageNumber="1117" precision="15" seconds="38" value="36.52722">36°31’38&quot;N</geoCoordinate>
,
<geoCoordinate id="EE5BE9B6680AFFA6FDA907BEFD6607C2" box="[539,644,1157,1177]" degrees="51" direction="east" minutes="38" orientation="longitude" pageId="6" pageNumber="1117" precision="15" seconds="46" value="51.646114">51°38’46&quot;E</geoCoordinate>
,
<quantity id="4C972294680AFFA6FD3E07BEFD2707C2" box="[652,709,1157,1177]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="1117" unit="m" value="880.0">
<elevation id="00426842680AFFA6FD3E07BEFD2707C2" box="[652,709,1157,1177]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="1117" unit="m" value="880.0">880 m</elevation>
</quantity>
,
<date id="FFD1A9B1680AFFA6FD7C07BEFCFD07C2" box="[718,799,1157,1177]" pageId="6" pageNumber="1117" value="2010-05-02">
<collectingDate id="EF955059680AFFA6FD7C07BEFCFD07C2" box="[718,799,1157,1177]" pageId="6" pageNumber="1117" value="2010-05-02">2.V.2010</collectingDate>
</date>
leg.
<collectorName id="269AEAA7680AFFA6FCFB07BEFC7607C2" box="[841,916,1157,1177]" pageId="6" pageNumber="1117">A. Skale</collectorName>
(43)&quot; (cWR).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C375DCFA680AFFA7FF000793FE84057F" lastPageId="7" lastPageNumber="1118" pageId="6" pageNumber="1117" type="description">
<paragraph id="8BD08F71680AFFA6FF000793FC9B0780" blockId="6.[178,1085,1192,1243]" pageId="6" pageNumber="1117">
D i a g n o s i s: Amedium-sized
<taxonomicName id="4C6FF4F2680AFFA6FDAE0792FCDC07E6" authority="" box="[540,830,1192,1214]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1117" phylum="Arthropoda" rank="subGenus" subGenus="Antisphodrus">
<emphasis id="B91B5363680AFFA6FDAE0792FD4207E5" box="[540,672,1193,1214]" italics="true" pageId="6" pageNumber="1117">Laemostenus</emphasis>
(
<emphasis id="B91B5363680AFFA6FD1D0793FCD407E5" box="[687,822,1192,1214]" italics="true" pageId="6" pageNumber="1117">Antisphodrus</emphasis>
)
</taxonomicName>
species sharing the main features of
<emphasis id="B91B5363680AFFA6FE9607FDFEDB0780" box="[292,313,1222,1243]" italics="true" pageId="6" pageNumber="1117">L.</emphasis>
<taxonomicName id="4C6FF4F2680AFFA6FE8D07FEFDAC0780" authority="(MORVAN)" baseAuthorityName="MORVAN" baseAuthorityYear="1974" box="[319,590,1221,1243]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1117" phylum="Arthropoda" rank="species" species="esfandiarii" subGenus="Antisphodrus">
(
<emphasis id="B91B5363680AFFA6FEF507FEFEB40780" box="[327,342,1221,1243]" italics="true" pageId="6" pageNumber="1117">A</emphasis>
.)
<emphasis id="B91B5363680AFFA6FEDE07FEFE3B0780" box="[364,473,1221,1243]" italics="true" pageId="6" pageNumber="1117">esfandiarii</emphasis>
(MORVAN)
</taxonomicName>
(see above). Habitus
<figureCitation id="135493F4680AFFA6FC9807FEFC9B0780" box="[810,889,1221,1243]" captionStart="Figs 19-20" captionStartId="15.[178,218,1533,1553]" captionTargetBox="[233,1026,222,1506]" captionTargetId="figure-67@15.[209,1053,212,1520]" captionTargetPageId="15" captionText="Figs 19-20: Laemostenus, habitus. (19) L. esfandiarii (MORVAN) (Kheiroud Kenar forest). (20) L. skalei nov.sp., holotype. Fig. 21: biotope of the type locality of L. skalei nov.sp., (Kheiroud Kenar forest) (photograph: André Skale)." figureDoi="http://doi.org/10.5281/zenodo.5328903" httpUri="https://zenodo.org/record/5328903/files/figure.png" pageId="6" pageNumber="1117">Fig. 20.</figureCitation>
</paragraph>
<paragraph id="8BD08F71680AFFA6FF0007D0FCC50645" blockId="6.[178,1085,1258,1310]" pageId="6" pageNumber="1117">
E t y m o l o g y: Cordially dedicated to our friend and colleague André Skale (Hof/Saale,
<collectingCountry id="F378CFE1680AFFA6FE9E0633FE6C0645" box="[300,398,1288,1310]" name="Germany" pageId="6" pageNumber="1117">Germany</collectingCountry>
), well known specialist in water beetles.
</paragraph>
<paragraph id="8BD08F71680AFFA6FF000615FC280618" blockId="6.[178,970,1325,1347]" box="[178,970,1325,1347]" pageId="6" pageNumber="1117">
D e s c r i p t i o n: Body size
<quantity id="4C972294680AFFA6FE480616FDB60618" box="[506,596,1325,1347]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="1117" unit="mm" value="12.5">12.5 mm</quantity>
(TL) and 14.0 mm (L), respectively.
</paragraph>
<paragraph id="8BD08F71680AFFA6FF000669FE9806DE" blockId="6.[178,1086,1362,1413]" pageId="6" pageNumber="1117">Colour: Dorsal surface black, very shiny, polished; labrum, palpomeres, antennae and legs brown reddish.</paragraph>
<paragraph id="8BD08F71680AFFA6FF0006AEFD460693" blockId="6.[178,1085,1429,1480]" pageId="6" pageNumber="1117">Microsculpure: Head and pronotum with hardly distinguishable transversal microlines, elytra with almost vanished isodiametric meshes.</paragraph>
<paragraph id="8BD08F71680AFFA6FF0006ECFBDD051E" blockId="6.[178,1087,1495,1605]" pageId="6" pageNumber="1117">Head wide, robust; tempora swollen, moderately oblique, convergent backwards; neck constriction evident; eyes relatively large, as long as 2/3 of tempora, not prominent outside; frontal impressions rather deep but short, wide, deep, reaching the level of the anterior supra-orbital setiferous punctures; antennae relatively short, exceeding by three</paragraph>
<paragraph id="8BD08F71680BFFA7FF0003E0FE4A0256" blockId="7.[178,1085,219,270]" pageId="7" pageNumber="1118">antennomeres the base of pronotum; antennomere 3 without accessory setae in addition to the apical fixed setae.</paragraph>
<paragraph id="8BD08F71680BFFA7FF000225FCCA029E" blockId="7.[178,1086,285,453]" pageId="7" pageNumber="1118">Pronotum cordate, slightly transverse (PL/PW: 0.90), its maximum width at the anterior third; lateral sides narrowly beaded, deeply sinuate and constricted to the basolateral angles, which are acute, evident; anterolateral angles rounded, slightly prominent; basal margin beaded at sides only. Disc moderately convex; median furrow superficial; basal impressions narrow, elongate, each with a few, deep punctures extended to the lateral furrows; anterolateral and basolateral setiferous pores present.</paragraph>
<paragraph id="8BD08F71680BFFA7FF0002EEFD2002B1" blockId="7.[178,706,469,491]" box="[178,706,469,491]" pageId="7" pageNumber="1118">Mesosternum not denticulate in front of mesocoxae.</paragraph>
<paragraph id="8BD08F71680BFFA7FF0002C1FDD001DE" blockId="7.[178,1086,506,645]" pageId="7" pageNumber="1118">Elytra elongate-ovate, relatively narrow (EL/EW: 1.60), convex, but with a wide depression on disc extended on intervals 1-4. Base narrow, oblique at sides; basal ridge weakly concave; humeri rounded, humeral tooth vanished. Striae very deep and deeply punctuate; intervals flat. Chaetotaxy: Basal pore present; umbilicate series with 16 setiferous punctures; 1-2 setae at apex of stria 7.</paragraph>
<paragraph id="8BD08F71680BFFA7FF0001AEFC870066" blockId="7.[178,1085,660,829]" pageId="7" pageNumber="1118">
Legs moderately long and slender; profemora with ventral side flat, its external and internal sides smooth, without marginal setae; mesotibiae straight; metatibiae each with some apical short, yellow reddish setae; metatarsomeres elongate and narrow, with dorsal pubescence long, scarce, not strigose; protarsomeres 1-3 dilated and with ventral, biseriate adhesive vestiture (male
<typeStatus id="54D431D3680BFFA7FDAA0031FD97007B" box="[536,629,778,800]" pageId="7" pageNumber="1118" type="holotype">holotype</typeStatus>
). Tarsal claws with a series of small teeth, reduced in number (3-4), along the basal half of the internal margin.
</paragraph>
<paragraph id="8BD08F71680BFFA7FF000077FED900E1" blockId="7.[178,1085,844,954]" pageId="7" pageNumber="1118">Male genitalia (Figs. 6-9): Median lobe of aedeagus small, slightly curved; apex thickened, prominent on the dorsal side, obtusely angular in the middle in dorsal aspect; right paramere long and slender, rounded apically; left paramere with developed apical membranous lobe.</paragraph>
<paragraph id="8BD08F71680BFFA7FF0000F2FE300084" blockId="7.[178,466,969,991]" box="[178,466,969,991]" pageId="7" pageNumber="1118">Female genitalia: Unknown.</paragraph>
<paragraph id="8BD08F71680BFFA7FF0000D5FC2F07EF" blockId="7.[178,1087,1006,1204]" pageId="7" pageNumber="1118">
C o m p a r i s o n s: Very close to
<taxonomicName id="4C6FF4F2680BFFA7FD8000D4FC32075F" authority="(MORVAN)" baseAuthorityName="MORVAN" baseAuthorityYear="1974" box="[562,976,1006,1028]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="species" species="esfandiarii" subGenus="Antisphodrus">
<emphasis id="B91B5363680BFFA7FD8000D4FDA5075F" box="[562,583,1007,1028]" italics="true" pageId="7" pageNumber="1118">L.</emphasis>
(
<emphasis id="B91B5363680BFFA7FDE700D5FD3E075F" box="[597,732,1006,1028]" italics="true" pageId="7" pageNumber="1118">Antisphodrus</emphasis>
)
<emphasis id="B91B5363680BFFA7FD5E00D5FCBB075F" box="[748,857,1006,1028]" italics="true" pageId="7" pageNumber="1118">esfandiarii</emphasis>
(MORVAN)
</taxonomicName>
but markedly distinct by the wide, inflated head, with markedly swollen tempora and larger eyes; the wider, cordiform pronotum (PL/PW: 1.0 in
<taxonomicName id="4C6FF4F2680BFFA7FD2D0711FCC50764" baseAuthorityName="MORVAN" baseAuthorityYear="1974" box="[671,807,1065,1087]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="species" species="esfandiarii">
<emphasis id="B91B5363680BFFA7FD2D0711FCC50764" box="[671,807,1065,1087]" italics="true" pageId="7" pageNumber="1118">L. esfandiarii</emphasis>
</taxonomicName>
,
<quantity id="4C972294680BFFA7FC870712FC9C0764" box="[821,894,1065,1087]" metricMagnitude="-2" metricUnit="m" metricValue="2.286" pageId="7" pageNumber="1118" unit="in" value="0.9">0.90 in</quantity>
<taxonomicName id="4C6FF4F2680BFFA7FC350711FC3D0764" authority="Casale &amp; D.W &amp; P, 2012" authorityName="Casale &amp; D.W &amp; P" authorityYear="2012" box="[903,991,1065,1087]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="B91B5363680BFFA7FC350711FC3D0764" box="[903,991,1065,1087]" italics="true" pageId="7" pageNumber="1118">L. skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228EE18680BFFA7FC570711FBD30764" box="[997,1073,1066,1087]" pageId="7" pageNumber="1118" rank="species">nov.sp.</taxonomicNameLabel>
), markedly widened in the anterior third, with lateral sides curved in front and deeply sinuate in the posterior forth; the narrower, elongate, parallel-sided elytra (compare
<figureCitation id="135493F4680BFFA7FBA1075FFF1507CC" captionStart="Figs 19-20" captionStartId="15.[178,218,1533,1553]" captionTargetBox="[233,1026,222,1506]" captionTargetId="figure-67@15.[209,1053,212,1520]" captionTargetPageId="15" captionText="Figs 19-20: Laemostenus, habitus. (19) L. esfandiarii (MORVAN) (Kheiroud Kenar forest). (20) L. skalei nov.sp., holotype. Fig. 21: biotope of the type locality of L. skalei nov.sp., (Kheiroud Kenar forest) (photograph: André Skale)." figureDoi="http://doi.org/10.5281/zenodo.5328903" httpUri="https://zenodo.org/record/5328903/files/figure.png" pageId="7" pageNumber="1118">Figs 19, 20</figureCitation>
); and by the shape of the median lobe of the aedeagus (Fig. 6, 7), which is more thickened, less elongate, and wider at apex in dorsal aspect in the new species.
</paragraph>
<paragraph id="8BD08F71680BFFA7FF0007FFFC9306FD" blockId="7.[178,1085,1219,1446]" pageId="7" pageNumber="1118">
D i s t r i b u t i o n a n d h a b i t a t: An epigean, forest dwelling species, known so far only from the
<typeStatus id="54D431D3680BFFA7FE3907DAFE5507AD" box="[395,439,1249,1270]" pageId="7" pageNumber="1118">type</typeStatus>
locality in the province
<collectingRegion id="49AB4193680BFFA7FD0A07DAFCD707AD" box="[696,821,1249,1271]" country="Iran" name="Mazandaran" pageId="7" pageNumber="1118">Māzandarān</collectingRegion>
in northern
<collectingCountry id="F378CFE1680BFFA7FC0507DAFC0207AD" box="[951,992,1249,1270]" name="Iran" pageId="7" pageNumber="1118">Iran</collectingCountry>
south of the town Now Shahr. This locality is situated in the Kheiroud Kenar forest, an old deciduous forest, partly of primary character (Fig. 21). The specimen was collected along a brook called &quot;Kheiroud Kenar stream&quot; on its bank or near to it under stones or wood (A. Skale, pers. comm.). Interestingly, in the same locality
<specimenCount id="9D6944F8680BFFA7FCBA066CFC700637" box="[776,914,1366,1388]" pageId="7" pageNumber="1118" type="generic">one specimen</specimenCount>
of
<taxonomicName id="4C6FF4F2680BFFA7FC06066CFEC206D2" authority="(MORVAN)" baseAuthorityName="MORVAN" baseAuthorityYear="1974" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="species" species="esfandiarii">
<emphasis id="B91B5363680BFFA7FC06066CFC200637" box="[948,962,1367,1388]" italics="true" pageId="7" pageNumber="1118">L</emphasis>
.
<emphasis id="B91B5363680BFFA7FC62066DFBDF0637" box="[976,1085,1366,1388]" italics="true" pageId="7" pageNumber="1118">esfandiarii</emphasis>
(MORVAN)
</taxonomicName>
was also collected. This syntopic condition is an additional proof of the specific separation of the two species of the
<taxonomicName id="4C6FF4F2680BFFA7FDE306AAFD3B06FD" baseAuthorityName="MORVAN" baseAuthorityYear="1974" box="[593,729,1424,1446]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="species" species="esfandiarii">
<emphasis id="B91B5363680BFFA7FDE306AAFD3B06FD" box="[593,729,1424,1446]" italics="true" pageId="7" pageNumber="1118">L. esfandiarii</emphasis>
</taxonomicName>
species group.
</paragraph>
<paragraph id="8BD08F71680BFFA7FF00068DFE84057F" blockId="7.[178,1085,1462,1572]" pageId="7" pageNumber="1118">
R e m a r k s: The discovery of this new, highly characterized species in a rather well investigated area, confirms the high diversity of
<taxonomicName id="4C6FF4F2680BFFA7FD0306E8FCFB06B2" authorityName="Laporte" authorityYear="1834" box="[689,793,1491,1513]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1118" phylum="Arthropoda" rank="subTribe" subTribe="Sphodrina">Sphodrina</taxonomicName>
species in the Iranian territory, highly interesting from the biogeographical point of view (see, for instance, LOHAJ &amp; CASALE 2011).
</paragraph>
</subSubSection>
</treatment>
</document>
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