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<document id="8A4C64E2B294920F09302894D01B86FF" ID-CLB-Dataset="39271" ID-DOI="10.11646/zootaxa.4058.1.2" ID-GBIF-Dataset="c03b8685-e7b1-423d-8db3-8a8c99ee2202" ID-ISSN="1175-5326" ID-Zenodo-Dep="239870" ID-ZooBank="B316C84C-7415-449F-9ABF-5DD600556844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461357852012" checkinUser="plazi" docAuthor="Tselikh, Ekaterina" docDate="2015" docId="03AD5B42FFB5FFC76893FB11FB61EF85" docLanguage="en" docName="zt04058p065.pdf" docOrigin="Zootaxa 4058 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spaniopus Walker 1833" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FF94233AFFB4FFC56804FF8CFFCBEE70" masterDocTitle="Review of the world species of Spaniopus Walker, 1833 (Hymenoptera: Pteromalidae), with description of a new species from the Russian Far East" masterLastPageNumber="65" masterPageNumber="41" pageNumber="42" updateTime="1698661609092" updateUser="ExternalLinkService">
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<mods:title id="FB59F702F7363F34418E7B5429A5C536">Review of the world species of Spaniopus Walker, 1833 (Hymenoptera: Pteromalidae), with description of a new species from the Russian Far East</mods:title>
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<treatment id="03AD5B42FFB5FFC76893FB11FB61EF85" ID-DOI="http://doi.org/10.5281/zenodo.6113666" ID-GBIF-Taxon="119647671" ID-Zenodo-Dep="6113666" LSID="urn:lsid:plazi:treatment:03AD5B42FFB5FFC76893FB11FB61EF85" httpUri="http://treatment.plazi.org/id/03AD5B42FFB5FFC76893FB11FB61EF85" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42">
<subSubSection id="C31EB9DFFFB5FFC46893FB11FE0AEAC7" box="[151,449,1181,1207]" pageId="1" pageNumber="42" type="nomenclature">
<paragraph id="8BBBEA54FFB5FFC46893FB11FE0AEAC7" blockId="1.[151,449,1181,1207]" box="[151,449,1181,1207]" pageId="1" pageNumber="42">
<heading id="D0F35D38FFB5FFC46893FB11FE0AEAC7" bold="true" box="[151,449,1181,1207]" fontSize="11" level="1" pageId="1" pageNumber="42" reason="1">
<taxonomicName id="4C0491D7FFB5FFC46893FB11FE0AEAC7" ID-CoL="7JXM" ID-ENA="337978" authority="Walker, 1833" authorityName="Walker" authorityYear="1833" box="[151,449,1181,1207]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB5FFC46893FB11FE0AEAC7" bold="true" box="[151,449,1181,1207]" pageId="1" pageNumber="42">
<emphasis id="B9703646FFB5FFC46893FB11FEDDEAC7" bold="true" box="[151,278,1181,1207]" italics="true" pageId="1" pageNumber="42">Spaniopus</emphasis>
<bibRefCitation id="EF9597A5FFB5FFC46919FB11FE0AEAC7" author="Walker" box="[285,449,1181,1207]" pageId="1" pageNumber="42" refString="Walker, F. (1833) Monographia Chalciditum. (Continued.). Entomological Magazine, 1 (5), 455 - 466." type="journal article" year="1833">Walker, 1833</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31EB9DFFFB5FFC46893FB68FAF5EBE3" pageId="1" pageNumber="42" type="reference_group">
<paragraph id="8BBBEA54FFB5FFC46893FB68FEF2EB68" blockId="1.[151,1436,1252,1427]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="AB14CD7AFFB5FFC46893FB68FEF2EB68" pageId="1" pageNumber="42">
<taxonomicName id="4C0491D7FFB5FFC46893FB68FE10EA8A" ID-CoL="7JXM" authority="Walker, 1833: 466" authorityName="Walker" authorityPageNumber="466" authorityYear="1833" box="[151,475,1252,1274]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB5FFC46893FB68FECAEA89" box="[151,257,1252,1273]" italics="true" pageId="1" pageNumber="42">Spaniopus</emphasis>
<treatmentCitation id="0AA5CC45FFB5FFC4690AFB68FE10EA8A" author="Walker" box="[270,475,1252,1274]" page="466" pageId="1" pageNumber="42" year="1833">
<bibRefCitation id="EF9597A5FFB5FFC4690AFB68FE6BEA8A" author="Walker" box="[270,416,1252,1274]" pageId="1" pageNumber="42" refString="Walker, F. (1833) Monographia Chalciditum. (Continued.). Entomological Magazine, 1 (5), 455 - 466." type="journal article" year="1833">Walker, 1833</bibRefCitation>
: 466
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54BF54F6FFB5FFC469E8FB69FDD5EA8A" box="[492,542,1253,1274]" pageId="1" pageNumber="42">Type</typeStatus>
species:
<taxonomicName id="4C0491D7FFB5FFC46A8FFB68FBCAEA8A" ID-CoL="6YVVC" authority="Walker, 1833" authorityName="Walker" authorityYear="1833" box="[651,1025,1252,1274]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="dissimilis">
<emphasis id="B9703646FFB5FFC46A8FFB68FCA9EA8A" box="[651,866,1252,1274]" italics="true" pageId="1" pageNumber="42">Spaniopus dissimilis</emphasis>
<bibRefCitation id="EF9597A5FFB5FFC46B74FB68FBCAEA8A" author="Walker" box="[880,1025,1252,1274]" pageId="1" pageNumber="42" refString="Walker, F. (1833) Monographia Chalciditum. (Continued.). Entomological Magazine, 1 (5), 455 - 466." type="journal article" year="1833">Walker, 1833</bibRefCitation>
</taxonomicName>
, by monotypy (♂
<typeStatus id="54BF54F6FFB5FFC46CDCFB68FAFCEA8A" box="[1240,1335,1252,1274]" pageId="1" pageNumber="42" type="lectotype">lectotype</typeStatus>
, BMNH, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBBEA54FFB5FFC46893FAADFD10EB04" blockId="1.[151,1436,1252,1427]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="AB14CD7AFFB5FFC46893FAADFD10EB04" pageId="1" pageNumber="42">
<taxonomicName id="4C0491D7FFB5FFC46893FAADFE07EB47" authority="Thomson, 1878: 131" authorityName="Thomson" authorityPageNumber="131" authorityYear="1878" box="[151,460,1313,1335]" class="Rhabditophora" family="Planariidae" genus="Polycelis" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="42" phylum="Platyhelminthes" rank="genus">
<emphasis id="B9703646FFB5FFC46893FAADFF3FEB47" box="[151,244,1313,1335]" italics="true" pageId="1" pageNumber="42">Polycelis</emphasis>
<treatmentCitation id="0AA5CC45FFB5FFC468FDFAADFE07EB47" author="Thomson" box="[249,460,1313,1335]" page="131" pageId="1" pageNumber="42" year="1878">
<bibRefCitation id="EF9597A5FFB5FFC468FDFAADFE50EB47" author="Thomson" box="[249,411,1313,1335]" pageId="1" pageNumber="42" refString="Thomson, C. G. (1878) Hymenoptera Scandinaviae 5. Pteromalus (Svederus) continuatio. Sweden, Lund, 307 pp." type="book" year="1878">Thomson, 1878</bibRefCitation>
: 131
</treatmentCitation>
</taxonomicName>
, 143.
<typeStatus id="54BF54F6FFB5FFC46A0DFAAEFDF0EB47" box="[521,571,1314,1335]" pageId="1" pageNumber="42">Type</typeStatus>
species:
<taxonomicName id="4C0491D7FFB5FFC46A92FAADFBC4EB47" ID-CoL="78MG5" authority="Walker, 1835" authorityName="Walker" authorityYear="1835" box="[662,1039,1313,1335]" class="Insecta" family="Pteromalidae" genus="Pteromalus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="conspersus">
<emphasis id="B9703646FFB5FFC46A92FAADFC4AEB47" box="[662,897,1313,1335]" italics="true" pageId="1" pageNumber="42">Pteromalus conspersus</emphasis>
Walker, 1835
</taxonomicName>
, designated by
<treatmentCitation id="0AA5CC45FFB5FFC46CADFAADFAB1EB47" author="Ashmead" box="[1193,1402,1313,1335]" page="386" pageId="1" pageNumber="42" year="1904">
<bibRefCitation id="EF9597A5FFB5FFC46CADFAADFA80EB47" author="Ashmead" box="[1193,1355,1313,1335]" pageId="1" pageNumber="42" refString="Ashmead, W. H. (1904) Classification of the chalcid flies of the superfamily Chalcidoidea, with descriptions of new species in the Carnegie Museum, collected in South America by Herbert H. Smith. Memoirs of the Carnegie Museum, 1 (4), 225 - 551. http: // dx. doi. org / 10.5962 / bhl. title. 10341" type="journal article" year="1904">Ashmead, 1904</bibRefCitation>
: 386
</treatmentCitation>
(♀
<typeStatus id="54BF54F6FFB5FFC468C3FACCFEEDEB26" box="[199,294,1344,1366]" pageId="1" pageNumber="42" type="lectotype">lectotype</typeStatus>
, BMNH, not examined). Junior primary homonym of
<taxonomicName id="4C0491D7FFB5FFC46B65FACCFBB9EB26" authority="Ehrenberg, 1831" authorityName="Ehrenberg" authorityYear="1831" box="[865,1138,1344,1366]" class="Rhabditophora" family="Planariidae" genus="Polycelis" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="42" phylum="Platyhelminthes" rank="genus">
<emphasis id="B9703646FFB5FFC46B65FACCFC75EB26" box="[865,958,1344,1366]" italics="true" pageId="1" pageNumber="42">Polycelis</emphasis>
Ehrenberg, 1831
</taxonomicName>
(
<taxonomicName id="4C0491D7FFB5FFC46C86FACCFB07EB26" box="[1154,1228,1344,1366]" class="Rhabditophora" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="42" phylum="Platyhelminthes" rank="order">Seriata</taxonomicName>
,
<taxonomicName id="4C0491D7FFB5FFC46CDDFACCFA86EB26" box="[1241,1357,1344,1366]" class="Rhabditophora" family="Planariidae" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="42" phylum="Platyhelminthes" rank="family">Planariidae</taxonomicName>
) (
<treatmentCitation id="0AA5CC45FFB5FFC46D67FACCFE94EB04" author="Dalla" page="87" pageId="1" pageNumber="42" year="1897">
<bibRefCitation id="EF9597A5FFB5FFC46D67FACCFEF2EB04" author="Dalla" pageId="1" pageNumber="42" refString="Dalla Torre, K. W. van (1897) Zur Nomenclatur der Chalcididen-Genera. Wiener Entomologische Zeitung, 16, 83 - 88." type="journal article" year="1897">Dalla Torre 1897</bibRefCitation>
: 87
</treatmentCitation>
). Synonymy by
<treatmentCitation id="0AA5CC45FFB5FFC46A01FAD2FD1CEB04" author="Graham" box="[517,727,1374,1396]" page="702" pageId="1" pageNumber="42" year="1969">
<bibRefCitation id="EF9597A5FFB5FFC46A01FAD2FD1CEB04" author="Graham" box="[517,727,1374,1396]" pageId="1" pageNumber="42" refString="Graham, M. W. R. de V. (1969) The Pteromalidae of north-western Europe (Hymenoptera: Chalcidoidea). Bulletin of the British Museum (Natural History) (Entomology), Supplement 16, 1 - 908." type="book chapter" year="1969">Graham (1969: 702)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBBEA54FFB5FFC46893FAF1FAF5EBE3" blockId="1.[151,1436,1252,1427]" box="[151,1342,1405,1427]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="AB14CD7AFFB5FFC46893FAF1FAF5EBE3" box="[151,1342,1405,1427]" pageId="1" pageNumber="42">
<taxonomicName id="4C0491D7FFB5FFC46893FAF1FE1BEBE3" authority="Hincks, 1944: 38" authorityName="Hincks" authorityPageNumber="38" authorityYear="1944" box="[151,464,1405,1427]" class="Insecta" family="Pteromalidae" genus="Neopolycelis" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB5FFC46893FAF1FED1EBE3" box="[151,282,1405,1427]" italics="true" pageId="1" pageNumber="42">Neopolycelis</emphasis>
Hincks, 1944: 38
</taxonomicName>
. Replacement name for
<taxonomicName id="4C0491D7FFB5FFC46AC1FAF1FC00EBE3" authority="Thomson, 1878" authorityName="Thomson" authorityYear="1878" box="[709,971,1405,1427]" class="Rhabditophora" family="Planariidae" genus="Polycelis" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="42" phylum="Platyhelminthes" rank="genus">
<emphasis id="B9703646FFB5FFC46AC1FAF1FCE9EBE3" box="[709,802,1405,1427]" italics="true" pageId="1" pageNumber="42">Polycelis</emphasis>
<bibRefCitation id="EF9597A5FFB5FFC46B2DFAF1FC00EBE3" author="Thomson" box="[809,971,1405,1427]" pageId="1" pageNumber="42" refString="Thomson, C. G. (1878) Hymenoptera Scandinaviae 5. Pteromalus (Svederus) continuatio. Sweden, Lund, 307 pp." type="book" year="1878">Thomson, 1878</bibRefCitation>
</taxonomicName>
. Synonymy by
<treatmentCitation id="0AA5CC45FFB5FFC46C6CFAF1FAF1EBE3" author="Graham" box="[1128,1338,1405,1427]" page="703" pageId="1" pageNumber="42" year="1969">
<bibRefCitation id="EF9597A5FFB5FFC46C6CFAF1FAF1EBE3" author="Graham" box="[1128,1338,1405,1427]" pageId="1" pageNumber="42" refString="Graham, M. W. R. de V. (1969) The Pteromalidae of north-western Europe (Hymenoptera: Chalcidoidea). Bulletin of the British Museum (Natural History) (Entomology), Supplement 16, 1 - 908." type="book chapter" year="1969">Graham (1969: 703)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31EB9DFFFB5FFC46893FA33FB87E9DD" pageId="1" pageNumber="42" type="diagnosis">
<paragraph id="8BBBEA54FFB5FFC46893FA33FB87E9DD" blockId="1.[151,1437,1471,2037]" pageId="1" pageNumber="42">
<emphasis id="B9703646FFB5FFC46893FA33FEDAEBA8" bold="true" box="[151,273,1471,1496]" pageId="1" pageNumber="42">Diagnosis.</emphasis>
Body colour dark metallic green to blush-green or blush-purple and often with diffuse coppery lustre; fore wing often with brownish cloud or patches. Sides of propodeum hairy, pilosity not reaching base of nucha (e.g.
<figureCitation id="133FF6D1FFB5FFC46893F98BFF29E850" box="[151,226,1543,1568]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[200,1384,193,1934]" captionTargetId="figure@4.[200,1385,193,1935]" captionTargetPageId="4" captionText="FIGURES 1 6. Spaniopus amoenus Foerster (non type, Hungary). 1, ♀ habitus, lateral view; 2, ♀ habitus, dorsal view; 3, ♀ head and antenna, lateral view; 4, ♀ scutellum and propodeum, dorsal view; 5, ♀ fore wing; 6, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239871/files/figure.png" pageId="1" pageNumber="42">Figs 4</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC468F5F984FEC4E850" box="[241,271,1544,1568]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">12</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC4691BF984FEF6E850" box="[287,317,1544,1568]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1914,1937]" captionTargetBox="[215,1371,193,1893]" captionTargetId="figure@8.[215,1372,193,1894]" captionTargetPageId="8" captionText="FIGURES 13 18. Spaniopus dissimilis Walker (♀ non type, Great Britain). 13, habitus, lateral view; 14, habitus, dorsal view; 15, head and antenna, lateral view; 16, head and antenna, frontal view; 17, fore wing; 18, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239873/files/figure.png" pageId="1" pageNumber="42">18</figureCitation>
); first metasomal tergite with small patch of hairs (e.g.
<figureCitation id="133FF6D1FFB5FFC46BCBF98BFBD2E850" box="[975,1049,1543,1568]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[200,1384,193,1934]" captionTargetId="figure@4.[200,1385,193,1935]" captionTargetPageId="4" captionText="FIGURES 1 6. Spaniopus amoenus Foerster (non type, Hungary). 1, ♀ habitus, lateral view; 2, ♀ habitus, dorsal view; 3, ♀ head and antenna, lateral view; 4, ♀ scutellum and propodeum, dorsal view; 5, ♀ fore wing; 6, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239871/files/figure.png" pageId="1" pageNumber="42">Figs 2</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C2DF984FBF2E850" box="[1065,1081,1544,1568]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">8</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C4CF984FBACE850" box="[1096,1127,1544,1568]" captionStart="FIGURES 46 52" captionStartId="18.[151,264,1931,1954]" captionTargetBox="[210,1377,193,1909]" captionTargetId="figure@18.[210,1378,193,1910]" captionTargetPageId="18" captionText="FIGURES 46 52. Spaniopus peisonis (Erdös) (♀ non type, Great Britain, ♂ paratype). 46, ♀ habitus, lateral view; 47, ♀ fore and hind wings; 48, ♀ mesosoma and propodeum, dorsal view; 49, ♀ antenna; 50, ♂ habitus, dorsal view; 51, ♂ antenna; 52, ♂ fore and hind wings." httpUri="https://zenodo.org/record/239878/files/figure.png" pageId="1" pageNumber="42">48</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C72F98BFB5DE850" box="[1142,1174,1543,1568]" captionStart="FIGURES 53 58" captionStartId="21.[151,264,1961,1984]" captionTargetBox="[199,1387,286,1940]" captionTargetId="figure@21.[199,1387,193,1940]" captionTargetPageId="21" captionText="FIGURES 53 58. Spaniopus sasacolae Kamijo (♀ holotype, ♂ paratype). 53, ♀ habitus, lateral view; 54, ♀ habitus, dorsal view; 55, ♀ head and antenna, frontal view; 56, ♀ antenna; 57, ♀ fore wing; 58, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239879/files/figure.png" pageId="1" pageNumber="42">54</figureCitation>
). Head and mesosoma distinctly reticulate; clypeus radially striate; nucha of propodeum strongly reticulate; metasoma smooth and shiny (e.g.
<figureCitation id="133FF6D1FFB5FFC468CBF9C3FEEFE818" box="[207,292,1615,1640]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1914,1937]" captionTargetBox="[215,1371,193,1893]" captionTargetId="figure@8.[215,1372,193,1894]" captionTargetPageId="8" captionText="FIGURES 13 18. Spaniopus dissimilis Walker (♀ non type, Great Britain). 13, habitus, lateral view; 14, habitus, dorsal view; 15, head and antenna, lateral view; 16, head and antenna, frontal view; 17, fore wing; 18, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239873/files/figure.png" pageId="1" pageNumber="42">Figs 14</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46934F9DCFE85E818" box="[304,334,1616,1640]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1931,1954]" captionTargetBox="[211,1376,195,1909]" captionTargetId="figure@10.[211,1377,195,1909]" captionTargetPageId="10" captionText="FIGURES 19 24. Spaniopus fulvicornis Bouček (♀ holotype). 19, habitus, lateral view; 20, habitus, dorsal view; 21, head and antenna, frontal view; 22, fore and hind wings; 23, propodeum, dorsal view; 24, head and antenna, lateral view." httpUri="https://zenodo.org/record/239874/files/figure.png" pageId="1" pageNumber="42">19</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC4695EF9C3FEB3E818" box="[346,376,1615,1640]" captionStart="FIGURES 25 31" captionStartId="12.[151,264,1933,1956]" captionTargetBox="[205,1371,193,1906]" captionTargetId="figure@12.[205,1371,193,1907]" captionTargetPageId="12" captionText="FIGURES 25 31. Spaniopus hedqvisti Huggert (♀, ♂ paratypes). 25, ♀ habitus, dorsal view (macropterous); 26, ♀ habitus, dorsal view (slightly brachypterous); 27, ♀ fore and hind wings; 28, ♀ antenna; 29, ♂ head and antenna; 30, ♂ mesosoma, propodeum and part of metasoma, dorsal view, mid tibia, lateral view; 31, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239875/files/figure.png" pageId="1" pageNumber="42">25</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46987F9DCFE68E818" box="[387,419,1616,1640]" captionStart="FIGURES 32 39" captionStartId="13.[151,264,1901,1924]" captionTargetBox="[220,1366,218,1879]" captionTargetId="figure@13.[220,1368,193,1881]" captionTargetPageId="13" captionText="FIGURES 32 39. Spaniopus spp. 32 35. Spaniopus japonicus Kamijo (♀ holotype). 32, habitus, dorsal view; 33, head and antenna, frontal view; 34, fore wing; 35, scutellum and propodeum, dorsal view. 36 39. Spaniopus nigriceps Kamijo (♀ holotype, ♂ paratype). 36, ♀ fore wing; 37, ♀ head and antenna, lateral view; 38, ♀ habitus, dorsal view; 39, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239876/files/figure.png" pageId="1" pageNumber="42">38</figureCitation>
). Head of female (e.g.
<figureCitation id="133FF6D1FFB5FFC46AA5F9C3FD23E818" box="[673,744,1615,1640]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">Figs 8</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46AF0F9DCFCDAE818" box="[756,785,1616,1640]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1931,1954]" captionTargetBox="[211,1376,195,1909]" captionTargetId="figure@10.[211,1377,195,1909]" captionTargetPageId="10" captionText="FIGURES 19 24. Spaniopus fulvicornis Bouček (♀ holotype). 19, habitus, lateral view; 20, habitus, dorsal view; 21, head and antenna, frontal view; 22, fore and hind wings; 23, propodeum, dorsal view; 24, head and antenna, lateral view." httpUri="https://zenodo.org/record/239874/files/figure.png" pageId="1" pageNumber="42">20</figureCitation>
) in dorsal view 2.02.2× as broad as long; scrobes absent; occiput without carina; lower margin of clypeus subemarginate (e.g.
<figureCitation id="133FF6D1FFB5FFC46B9FF9F8FC29E8FC" box="[923,994,1652,1677]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">Figs 9</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46BEAF9F8FBC7E8FC" box="[1006,1036,1652,1676]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1931,1954]" captionTargetBox="[211,1376,195,1909]" captionTargetId="figure@10.[211,1377,195,1909]" captionTargetPageId="10" captionText="FIGURES 19 24. Spaniopus fulvicornis Bouček (♀ holotype). 19, habitus, lateral view; 20, habitus, dorsal view; 21, head and antenna, frontal view; 22, fore and hind wings; 23, propodeum, dorsal view; 24, head and antenna, lateral view." httpUri="https://zenodo.org/record/239874/files/figure.png" pageId="1" pageNumber="42">21</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C1CF9F8FBF3E8FD" box="[1048,1080,1652,1677]" captionStart="FIGURES 53 58" captionStartId="21.[151,264,1961,1984]" captionTargetBox="[199,1387,286,1940]" captionTargetId="figure@21.[199,1387,193,1940]" captionTargetPageId="21" captionText="FIGURES 53 58. Spaniopus sasacolae Kamijo (♀ holotype, ♂ paratype). 53, ♀ habitus, lateral view; 54, ♀ habitus, dorsal view; 55, ♀ head and antenna, frontal view; 56, ♀ antenna; 57, ♀ fore wing; 58, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239879/files/figure.png" pageId="1" pageNumber="42">55</figureCitation>
). Antennae in both sexes 11263 (e.g.
<figureCitation id="133FF6D1FFB5FFC468D6F91BFED0E8C0" box="[210,283,1687,1712]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[200,1384,193,1934]" captionTargetId="figure@4.[200,1385,193,1935]" captionTargetPageId="4" captionText="FIGURES 1 6. Spaniopus amoenus Foerster (non type, Hungary). 1, ♀ habitus, lateral view; 2, ♀ habitus, dorsal view; 3, ♀ head and antenna, lateral view; 4, ♀ scutellum and propodeum, dorsal view; 5, ♀ fore wing; 6, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239871/files/figure.png" pageId="1" pageNumber="42">Figs 3</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC4692DF914FE8CE8C0" box="[297,327,1688,1712]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">10</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46951F914FEB8E8C0" box="[341,371,1688,1712]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1931,1954]" captionTargetBox="[211,1376,195,1909]" captionTargetId="figure@10.[211,1377,195,1909]" captionTargetPageId="10" captionText="FIGURES 19 24. Spaniopus fulvicornis Bouček (♀ holotype). 19, habitus, lateral view; 20, habitus, dorsal view; 21, head and antenna, frontal view; 22, fore and hind wings; 23, propodeum, dorsal view; 24, head and antenna, lateral view." httpUri="https://zenodo.org/record/239874/files/figure.png" pageId="1" pageNumber="42">24</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46985F91BFE54E8C0" box="[385,415,1687,1712]" captionStart="FIGURES 59 66" captionStartId="22.[151,264,1928,1951]" captionTargetBox="[211,1376,193,1906]" captionTargetId="figure@22.[211,1377,193,1907]" captionTargetPageId="22" captionText="FIGURES 59 66. Spaniopus spp. 59 62. Spaniopus polyspilus Graham (♀ holotype). 59, habitus; 60, head, frontal view; 61, antenna; 62, fore wing. 63 66. Spaniopus varicornis Bouček (♂ holotype). 63, habitus, dorsal view; 64, propodeum; 65, mid tibia, lateral view; 66, head and antenna, lateral view." httpUri="https://zenodo.org/record/239880/files/figure.png" pageId="1" pageNumber="42">61</figureCitation>
), inserted a little above level of lower edges of eyes (e.g.
<figureCitation id="133FF6D1FFB5FFC46C39F91BFB4CE8C0" box="[1085,1159,1687,1712]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">Figs 9</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C91F91BFB78E8C0" box="[1173,1203,1687,1712]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1914,1937]" captionTargetBox="[215,1371,193,1893]" captionTargetId="figure@8.[215,1372,193,1894]" captionTargetPageId="8" captionText="FIGURES 13 18. Spaniopus dissimilis Walker (♀ non type, Great Britain). 13, habitus, lateral view; 14, habitus, dorsal view; 15, head and antenna, lateral view; 16, head and antenna, frontal view; 17, fore wing; 18, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239873/files/figure.png" pageId="1" pageNumber="42">15</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CC5F914FB14E8C0" box="[1217,1247,1688,1712]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1931,1954]" captionTargetBox="[211,1376,195,1909]" captionTargetId="figure@10.[211,1377,195,1909]" captionTargetPageId="10" captionText="FIGURES 19 24. Spaniopus fulvicornis Bouček (♀ holotype). 19, habitus, lateral view; 20, habitus, dorsal view; 21, head and antenna, frontal view; 22, fore and hind wings; 23, propodeum, dorsal view; 24, head and antenna, lateral view." httpUri="https://zenodo.org/record/239874/files/figure.png" pageId="1" pageNumber="42">21</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CE9F91BFAC7E8C0" box="[1261,1292,1687,1712]" captionStart="FIGURES 53 58" captionStartId="21.[151,264,1961,1984]" captionTargetBox="[199,1387,286,1940]" captionTargetId="figure@21.[199,1387,193,1940]" captionTargetPageId="21" captionText="FIGURES 53 58. Spaniopus sasacolae Kamijo (♀ holotype, ♂ paratype). 53, ♀ habitus, lateral view; 54, ♀ habitus, dorsal view; 55, ♀ head and antenna, frontal view; 56, ♀ antenna; 57, ♀ fore wing; 58, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239879/files/figure.png" pageId="1" pageNumber="42">55</figureCitation>
); both anelli transverse or second anellus subquadrate to quadrate; funicular segments in female longer than broad, subquadrate or transverse, with one row of dense sensilla; clava symmetric, usually not wider than funicular segments; funicular segments in male more elongate than in female or of different size and colouration (e.g.
<figureCitation id="133FF6D1FFB5FFC46C86F888FB13E96C" box="[1154,1240,1796,1821]" captionStart="FIGURES 25 31" captionStartId="12.[151,264,1933,1956]" captionTargetBox="[205,1371,193,1906]" captionTargetId="figure@12.[205,1371,193,1907]" captionTargetPageId="12" captionText="FIGURES 25 31. Spaniopus hedqvisti Huggert (♀, ♂ paratypes). 25, ♀ habitus, dorsal view (macropterous); 26, ♀ habitus, dorsal view (slightly brachypterous); 27, ♀ fore and hind wings; 28, ♀ antenna; 29, ♂ head and antenna; 30, ♂ mesosoma, propodeum and part of metasoma, dorsal view, mid tibia, lateral view; 31, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239875/files/figure.png" pageId="1" pageNumber="42">Figs 29</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CE1F888FACEE96C" box="[1253,1285,1796,1820]" captionStart="FIGURES 40 45" captionStartId="15.[151,264,1901,1924]" captionTargetBox="[220,1367,193,1879]" captionTargetId="figure@15.[220,1368,193,1881]" captionTargetPageId="15" captionText="FIGURES 40 45. Spaniopus spp. 40 42. Spaniopus dissimilis Walker (♂ lectotype). 40, habitus, dorsal view; 41, head and antenna, dorsal view; 42, mid tibia, lateral view. 43 45. Spaniopus monospilus (Thomson) (♂ lectotype). 43, antenna; 44, mid tibia, lateral view; 45, head, frontal view." httpUri="https://zenodo.org/record/239877/files/figure.png" pageId="1" pageNumber="42">43</figureCitation>
). Mandibular formula 3:4 or rarely 4:4. Mesosoma usually moderately arched dorsally (e.g.
<figureCitation id="133FF6D1FFB5FFC46C25F8ABFBA2E930" box="[1057,1129,1831,1856]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[200,1384,193,1934]" captionTargetId="figure@4.[200,1385,193,1935]" captionTargetPageId="4" captionText="FIGURES 1 6. Spaniopus amoenus Foerster (non type, Hungary). 1, ♀ habitus, lateral view; 2, ♀ habitus, dorsal view; 3, ♀ head and antenna, lateral view; 4, ♀ scutellum and propodeum, dorsal view; 5, ♀ fore wing; 6, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239871/files/figure.png" pageId="1" pageNumber="42">Figs 1</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C7EF8A4FB41E94F" box="[1146,1162,1832,1855]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">7</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46C9DF8ABFB73E930" box="[1177,1208,1831,1856]" captionStart="FIGURES 53 58" captionStartId="21.[151,264,1961,1984]" captionTargetBox="[199,1387,286,1940]" captionTargetId="figure@21.[199,1387,193,1940]" captionTargetPageId="21" captionText="FIGURES 53 58. Spaniopus sasacolae Kamijo (♀ holotype, ♂ paratype). 53, ♀ habitus, lateral view; 54, ♀ habitus, dorsal view; 55, ♀ head and antenna, frontal view; 56, ♀ antenna; 57, ♀ fore wing; 58, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239879/files/figure.png" pageId="1" pageNumber="42">53</figureCitation>
); pronotum without smooth and shiny carina; notauli incomplete; scutellum with noticeable but shallow frenal line. Propodeum with conspicuous subglobose nucha, plicae sinuate before constriction at nuchal base (e.g.
<figureCitation id="133FF6D1FFB5FFC46C56F8E3FB52E9F8" box="[1106,1177,1903,1928]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1958,1981]" captionTargetBox="[200,1384,193,1934]" captionTargetId="figure@4.[200,1385,193,1935]" captionTargetPageId="4" captionText="FIGURES 1 6. Spaniopus amoenus Foerster (non type, Hungary). 1, ♀ habitus, lateral view; 2, ♀ habitus, dorsal view; 3, ♀ head and antenna, lateral view; 4, ♀ scutellum and propodeum, dorsal view; 5, ♀ fore wing; 6, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239871/files/figure.png" pageId="1" pageNumber="42">Figs 4</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CA0F8FCFB09E9F8" box="[1188,1218,1904,1928]" captionStart="FIGURES 7 12" captionStartId="6.[151,264,1955,1978]" captionTargetBox="[201,1385,193,1934]" captionTargetId="figure@6.[201,1385,193,1935]" captionTargetPageId="6" captionText="FIGURES 7 12. Spaniopus belokobylskiji sp. nov. (♀ holotype). 7, habitus, lateral view; 8, habitus, dorsal view; 9, head and antenna, frontal view; 10, antenna; 11, fore wing; 12, scutellum and propodeum, dorsal view." httpUri="https://zenodo.org/record/239872/files/figure.png" pageId="1" pageNumber="42">12</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CCAF8E3FB27E9F8" box="[1230,1260,1903,1928]" captionStart="FIGURES 32 39" captionStartId="13.[151,264,1901,1924]" captionTargetBox="[220,1366,218,1879]" captionTargetId="figure@13.[220,1368,193,1881]" captionTargetPageId="13" captionText="FIGURES 32 39. Spaniopus spp. 32 35. Spaniopus japonicus Kamijo (♀ holotype). 32, habitus, dorsal view; 33, head and antenna, frontal view; 34, fore wing; 35, scutellum and propodeum, dorsal view. 36 39. Spaniopus nigriceps Kamijo (♀ holotype, ♂ paratype). 36, ♀ fore wing; 37, ♀ head and antenna, lateral view; 38, ♀ habitus, dorsal view; 39, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239876/files/figure.png" pageId="1" pageNumber="42">35</figureCitation>
,
<figureCitation id="133FF6D1FFB5FFC46CF3F8FCFADCE9F8" box="[1271,1303,1904,1928]" captionStart="FIGURES 46 52" captionStartId="18.[151,264,1931,1954]" captionTargetBox="[210,1377,193,1909]" captionTargetId="figure@18.[210,1378,193,1910]" captionTargetPageId="18" captionText="FIGURES 46 52. Spaniopus peisonis (Erdös) (♀ non type, Great Britain, ♂ paratype). 46, ♀ habitus, lateral view; 47, ♀ fore and hind wings; 48, ♀ mesosoma and propodeum, dorsal view; 49, ♀ antenna; 50, ♂ habitus, dorsal view; 51, ♂ antenna; 52, ♂ fore and hind wings." httpUri="https://zenodo.org/record/239878/files/figure.png" pageId="1" pageNumber="42">48</figureCitation>
). Fore wing with basal cell bare or pilose, veins not thickened. Metasoma ovate or sublanceolate.
</paragraph>
</subSubSection>
<subSubSection id="C31EB9DFFFB5FFC468C3F834FDB4E9A1" box="[199,639,1976,2001]" pageId="1" pageNumber="42" type="distribution">
<paragraph id="8BBBEA54FFB5FFC468C3F834FDB4E9A1" blockId="1.[151,1437,1471,2037]" box="[199,639,1976,2001]" pageId="1" pageNumber="42">
<emphasis id="B9703646FFB5FFC468C3F834FEABE9A1" bold="true" box="[199,352,1976,2001]" pageId="1" pageNumber="42">Distribution.</emphasis>
Palaearctic and Nearctic.
</paragraph>
</subSubSection>
<subSubSection id="C31EB9DFFFB5FFC768C3F850FB61EF85" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42" type="discussion">
<paragraph id="8BBBEA54FFB5FFC768C3F850FDE8EEA4" blockId="1.[151,1437,1471,2037]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42">
<emphasis id="B9703646FFB5FFC468C3F850FE84E985" bold="true" box="[199,335,2012,2037]" pageId="1" pageNumber="42">Comments.</emphasis>
For detailed discussion of the synonymy of
<taxonomicName id="4C0491D7FFB5FFC46B45F851FC7DE984" box="[833,950,2013,2036]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB5FFC46B45F851FC7DE984" box="[833,950,2013,2036]" italics="true" pageId="1" pageNumber="42">Spaniopus</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EF9597A5FFB5FFC46BEBF850FAC4E985" author="Graham" box="[1007,1295,2012,2037]" pageId="1" pageNumber="42" refString="Graham, M. W. R. de V. (1956) A revision of the Walker types of Pteromalidae (Hym., Chalcidoidea). Part 2. Entomologist's Monthly Magazine, 92, 246 - 263." type="journal article" year="1956">Graham (1956: 250251)</bibRefCitation>
. Differences between
<taxonomicName id="4C0491D7FFB6FFC768FFFF14FEBBEEDF" box="[251,368,152,175]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB6FFC768FFFF14FEBBEEDF" box="[251,368,152,175]" italics="true" pageId="2" pageNumber="43">Spaniopus</emphasis>
</taxonomicName>
and the similar genera
<taxonomicName id="4C0491D7FFB6FFC76A70FF1BFC7EEEC0" authority="Thomson, 1878" authorityName="Thomson" authorityYear="1878" box="[628,949,151,176]" class="Insecta" family="Pteromalidae" genus="Trichomalus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB6FFC76A70FF1BFCCBEEC0" box="[628,768,151,176]" italics="true" pageId="2" pageNumber="43">Trichomalus</emphasis>
<bibRefCitation id="EF9597A5FFB6FFC76B02FF1BFC7EEEC0" author="Thomson" box="[774,949,151,176]" pageId="2" pageNumber="43" refString="Thomson, C. G. (1878) Hymenoptera Scandinaviae 5. Pteromalus (Svederus) continuatio. Sweden, Lund, 307 pp." type="book" year="1878">Thomson, 1878</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0491D7FFB6FFC76BEFFF1BFAD9EEC0" authority="Foerster, 1856" authorityName="Foerster" authorityYear="1856" box="[1003,1298,151,176]" class="Insecta" family="Pteromalidae" genus="Peridesmia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB6FFC76BEFFF1BFBA1EEC0" box="[1003,1130,151,176]" italics="true" pageId="2" pageNumber="43">Peridesmia</emphasis>
<bibRefCitation id="EF9597A5FFB6FFC76C76FF14FAD9EEC0" author="Foerster" box="[1138,1298,151,176]" pageId="2" pageNumber="43" refString="Foerster, A. (1856) Hymenopterologische Studien. 2. Chalcidiae und Proctotrupii. Aachen, 152 pp." type="book" year="1856">Foerster, 1856</bibRefCitation>
</taxonomicName>
are given in the key of
<bibRefCitation id="EF9597A5FFB6FFC7690AFF30FDD4EEA4" box="[270,543,188,213]" pageId="2" pageNumber="43" refString="Boucek, Z., Rasplus, J. - Y. (1991) Illustrated key to West-Palaearctic genera of Pteromalidae (Hymenoptera: Chalcidoidea). Institut National de la Recherche Agronomique, Paris, 140 pp." type="book">Bouček, Rasplus (1991)</bibRefCitation>
.
</paragraph>
<paragraph id="8BBBEA54FFB6FFC768C3FF6CFC18EFDD" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="43">
Most species of
<taxonomicName id="4C0491D7FFB6FFC76980FF6CFE32EE87" box="[388,505,224,247]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB6FFC76980FF6CFE32EE87" box="[388,505,224,247]" italics="true" pageId="2" pageNumber="43">Spaniopus</emphasis>
</taxonomicName>
have long fore wings that distinctly exceed the apex of the metasoma. However, some species, such as
<emphasis id="B9703646FFB6FFC7698AFE89FDE3EF6D" box="[398,552,260,285]" italics="true" pageId="2" pageNumber="43">
<taxonomicName id="4C0491D7FFB6FFC7698AFE89FDEEEF6D" box="[398,549,260,285]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="fulvicornis">S. fulvicornis</taxonomicName>
,
</emphasis>
are characterized by shortened fore wings. The length of the wings was used in some keys as the main feature for separating species (
<bibRefCitation id="EF9597A5FFB6FFC76AF6FEABFC47EF30" author="Boucek" box="[754,908,295,320]" pageId="2" pageNumber="43" refString="Boucek, Z. (1972) On European Pteromalidae (Hymenoptera): a revision of Cleonymus, Eunotus and Spaniopus, with descriptions of new genera and species. Bulletin of the British Museum (Natural History) (Entomology), 27 (5), 265 - 315." type="journal article" year="1972">Bouček 1972</bibRefCitation>
;
<bibRefCitation id="EF9597A5FFB6FFC76B93FEABFBE0EF30" author="Kamijo" box="[919,1067,295,320]" pageId="2" pageNumber="43" refString="Kamijo, K. (1981) Three new species of Spaniopus (Hymenoptera: Pteromalidae) from Japan. Akitu, 36, 1 - 9." type="journal article" year="1981">Kamijo 1981</bibRefCitation>
) because wing-length variability within a single species was not realized. However,
<bibRefCitation id="EF9597A5FFB6FFC76B0CFEC1FC77EF15" author="Huggert" box="[776,956,332,357]" pageId="2" pageNumber="43" refString="Huggert, L. (1976) Descriptions of a previously unknown male of a new genus and three new species of Pteromalidae (Hym., Chalcidoidea) from northern Sweden. Entomologisk Tidskrift, 97 (1 - 2), 55 - 64." type="journal article" year="1976">Huggert (1976)</bibRefCitation>
described wing-length variability for
<taxonomicName id="4C0491D7FFB6FFC76D83FEC1FF37EFF8" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="hedqvisti">
<emphasis id="B9703646FFB6FFC76D83FEC1FF37EFF8" italics="true" pageId="2" pageNumber="43">S. hedqvisti</emphasis>
</taxonomicName>
(
<figureCitation id="133FF6D1FFB6FFC76908FEE3FE5BEFF8" box="[268,400,367,392]" captionStart="FIGURES 25 31" captionStartId="12.[151,264,1933,1956]" captionTargetBox="[205,1371,193,1906]" captionTargetId="figure@12.[205,1371,193,1907]" captionTargetPageId="12" captionText="FIGURES 25 31. Spaniopus hedqvisti Huggert (♀, ♂ paratypes). 25, ♀ habitus, dorsal view (macropterous); 26, ♀ habitus, dorsal view (slightly brachypterous); 27, ♀ fore and hind wings; 28, ♀ antenna; 29, ♂ head and antenna; 30, ♂ mesosoma, propodeum and part of metasoma, dorsal view, mid tibia, lateral view; 31, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239875/files/figure.png" pageId="2" pageNumber="43">Figs 25, 26</figureCitation>
), which suggests that wing-length may be variable for more species. We have not used this feature in our new identification key to species because of this possibility.
</paragraph>
<paragraph id="8BBBEA54FFB6FFC768C3FE3BFB61EF85" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="43">
Sexual dimorphism in
<taxonomicName id="4C0491D7FFB6FFC769CDFE34FDF5EFBF" box="[457,574,440,463]" class="Insecta" family="Pteromalidae" genus="Spaniopus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9703646FFB6FFC769CDFE34FDF5EFBF" box="[457,574,440,463]" italics="true" pageId="2" pageNumber="43">Spaniopus</emphasis>
</taxonomicName>
is exhibited in the different shape and colouration of the antennae (
<figureCitation id="133FF6D1FFB6FFC76D3BFE3BFA5CEFA0" box="[1343,1431,439,464]" captionStart="FIGURES 25 31" captionStartId="12.[151,264,1933,1956]" captionTargetBox="[205,1371,193,1906]" captionTargetId="figure@12.[205,1371,193,1907]" captionTargetPageId="12" captionText="FIGURES 25 31. Spaniopus hedqvisti Huggert (♀, ♂ paratypes). 25, ♀ habitus, dorsal view (macropterous); 26, ♀ habitus, dorsal view (slightly brachypterous); 27, ♀ fore and hind wings; 28, ♀ antenna; 29, ♂ head and antenna; 30, ♂ mesosoma, propodeum and part of metasoma, dorsal view, mid tibia, lateral view; 31, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239875/files/figure.png" pageId="2" pageNumber="43">Figs 29</figureCitation>
,
<figureCitation id="133FF6D1FFB6FFC76893FE50FF7DEF84" box="[151,182,476,500]" captionStart="FIGURES 40 45" captionStartId="15.[151,264,1901,1924]" captionTargetBox="[220,1367,193,1879]" captionTargetId="figure@15.[220,1368,193,1881]" captionTargetPageId="15" captionText="FIGURES 40 45. Spaniopus spp. 40 42. Spaniopus dissimilis Walker (♂ lectotype). 40, habitus, dorsal view; 41, head and antenna, dorsal view; 42, mid tibia, lateral view. 43 45. Spaniopus monospilus (Thomson) (♂ lectotype). 43, antenna; 44, mid tibia, lateral view; 45, head, frontal view." httpUri="https://zenodo.org/record/239877/files/figure.png" pageId="2" pageNumber="43">43</figureCitation>
,
<figureCitation id="133FF6D1FFB6FFC768C5FE50FF2BEF85" box="[193,224,476,501]" captionStart="FIGURES 59 66" captionStartId="22.[151,264,1928,1951]" captionTargetBox="[211,1376,193,1906]" captionTargetId="figure@22.[211,1377,193,1907]" captionTargetPageId="22" captionText="FIGURES 59 66. Spaniopus spp. 59 62. Spaniopus polyspilus Graham (♀ holotype). 59, habitus; 60, head, frontal view; 61, antenna; 62, fore wing. 63 66. Spaniopus varicornis Bouček (♂ holotype). 63, habitus, dorsal view; 64, propodeum; 65, mid tibia, lateral view; 66, head and antenna, lateral view." httpUri="https://zenodo.org/record/239880/files/figure.png" pageId="2" pageNumber="43">66</figureCitation>
), mid tibiae (
<figureCitation id="133FF6D1FFB6FFC76970FE50FE00EF84" box="[372,459,476,501]" captionStart="FIGURES 25 31" captionStartId="12.[151,264,1933,1956]" captionTargetBox="[205,1371,193,1906]" captionTargetId="figure@12.[205,1371,193,1907]" captionTargetPageId="12" captionText="FIGURES 25 31. Spaniopus hedqvisti Huggert (♀, ♂ paratypes). 25, ♀ habitus, dorsal view (macropterous); 26, ♀ habitus, dorsal view (slightly brachypterous); 27, ♀ fore and hind wings; 28, ♀ antenna; 29, ♂ head and antenna; 30, ♂ mesosoma, propodeum and part of metasoma, dorsal view, mid tibia, lateral view; 31, ♂ habitus, lateral view." httpUri="https://zenodo.org/record/239875/files/figure.png" pageId="2" pageNumber="43">Figs 30</figureCitation>
,
<figureCitation id="133FF6D1FFB6FFC769D2FE50FDD4EF84" box="[470,543,476,500]" captionStart="FIGURES 40 45" captionStartId="15.[151,264,1901,1924]" captionTargetBox="[220,1367,193,1879]" captionTargetId="figure@15.[220,1368,193,1881]" captionTargetPageId="15" captionText="FIGURES 40 45. Spaniopus spp. 40 42. Spaniopus dissimilis Walker (♂ lectotype). 40, habitus, dorsal view; 41, head and antenna, dorsal view; 42, mid tibia, lateral view. 43 45. Spaniopus monospilus (Thomson) (♂ lectotype). 43, antenna; 44, mid tibia, lateral view; 45, head, frontal view." httpUri="https://zenodo.org/record/239877/files/figure.png" pageId="2" pageNumber="43">42, 44</figureCitation>
,
<figureCitation id="133FF6D1FFB6FFC76A2FFE50FD83EF85" box="[555,584,476,501]" captionStart="FIGURES 59 66" captionStartId="22.[151,264,1928,1951]" captionTargetBox="[211,1376,193,1906]" captionTargetId="figure@22.[211,1377,193,1907]" captionTargetPageId="22" captionText="FIGURES 59 66. Spaniopus spp. 59 62. Spaniopus polyspilus Graham (♀ holotype). 59, habitus; 60, head, frontal view; 61, antenna; 62, fore wing. 63 66. Spaniopus varicornis Bouček (♂ holotype). 63, habitus, dorsal view; 64, propodeum; 65, mid tibia, lateral view; 66, head and antenna, lateral view." httpUri="https://zenodo.org/record/239880/files/figure.png" pageId="2" pageNumber="43">65</figureCitation>
) and metasoma in males and females of some species.
</paragraph>
</subSubSection>
</treatment>
</document>