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<document id="2787EC991C31A5F5ECA68E10C8853413" ID-DOI="10.5281/zenodo.183126" ID-GBIF-Dataset="5ba6cec4-345d-4f95-8064-dd9dc33dac4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="183126" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459892214864" checkinUser="plazi" docAuthor="Miller, Kelly B. &amp; Spangler, Paul J." docDate="2008" docId="03AD2179AE5DFFC3FF78FF4CFC9FF8C6" docLanguage="en" docName="zt01827p052.pdf" docOrigin="Zootaxa 1827" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Fontidessus ornatus Miller, new species" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="FF945901AE58FFC6FFEFFFDBFFA9FFC9" masterDocTitle="Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species" masterLastPageNumber="52" masterPageNumber="45" pageNumber="50" updateTime="1698228095007" updateUser="plazi">
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<mods:title id="D082749582AD6243D66BE55BCB544333">Fontidessus Miller and Spangler, a new genus of Bidessini from Venezuela (Coleoptera: Dytiscidae: Hydroporinae) with three new species</mods:title>
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<mods:namePart id="CCFD4D36CFD120A50CCFECB73257B01D">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="89135B401DF21AC226F7DE9BEB308C07">Spangler, Paul J.</mods:namePart>
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<mods:typeOfResource id="52159A77E8CAA517459D4E5110C312A8">text</mods:typeOfResource>
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<mods:date id="87C57DEA1FE2827E2E02A9BBF0159A78">2008</mods:date>
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<mods:identifier id="CA1BCC110E9BE1A344BA0DFA5A8262AD" type="DOI">10.5281/zenodo.183126</mods:identifier>
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<treatment id="03AD2179AE5DFFC3FF78FF4CFC9FF8C6" ID-DOI="http://doi.org/10.5281/zenodo.6233626" ID-GBIF-Taxon="119372126" ID-Zenodo-Dep="6233626" LSID="urn:lsid:plazi:treatment:03AD2179AE5DFFC3FF78FF4CFC9FF8C6" httpUri="http://treatment.plazi.org/id/03AD2179AE5DFFC3FF78FF4CFC9FF8C6" lastPageNumber="50" pageId="5" pageNumber="50">
<subSubSection id="C31EC3E4AE5DFFC3FF78FF4CFECEFF1D" pageId="5" pageNumber="50" type="nomenclature">
<paragraph id="8BBB906FAE5DFFC3FF78FF4CFDD4FF78" blockId="5.[151,637,151,212]" box="[151,637,151,177]" pageId="5" pageNumber="50">
<heading id="D0F32703AE5DFFC3FF78FF4CFDD4FF78" bold="true" box="[151,637,151,177]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
<emphasis id="B9704C7DAE5DFFC3FF78FF4CFDD4FF78" bold="true" box="[151,637,151,177]" pageId="5" pageNumber="50">
<taxonomicName id="4C04EBECAE5DFFC3FF78FF4CFE49FF78" ID-CoL="6JDL6" authority="Miller" authorityName="Miller" box="[151,480,151,177]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ornatus" status="sp. nov.">
<emphasis id="B9704C7DAE5DFFC3FF78FF4CFE25FF78" bold="true" box="[151,396,151,177]" italics="true" pageId="5" pageNumber="50">Fontidessus ornatus</emphasis>
Miller
</taxonomicName>
,
<taxonomicNameLabel id="A243F106AE5DFFC3FE02FF4CFDD4FF78" box="[493,637,151,177]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB906FAE5DFFC3FF78FF61FECEFF1D" blockId="5.[151,637,151,212]" box="[151,359,186,212]" pageId="5" pageNumber="50">
(
<figureCitation id="133F8CEAAE5DFFC3FF70FF61FEA0FF1D" box="[159,265,186,212]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Figs 2, 6</figureCitation>
,
<figureCitation id="133F8CEAAE5DFFC3FEFAFF61FEF7FF1D" box="[277,350,186,212]" captionStart="FIGURES 8 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 17. Fontidessus species. 8 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">1113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF78FEDEFA3CFED6" box="[151,1429,261,287]" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="8BBB906FAE5DFFC3FF78FEDEFA3CFED6" blockId="5.[151,1436,261,1807]" box="[151,1429,261,287]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF78FEDEFE94FED6" bold="true" box="[151,317,261,287]" pageId="5" pageNumber="50">
<typeStatus id="54BF2ECDAE5DFFC3FF78FEDEFF7DFED6" box="[151,212,261,287]" pageId="5" pageNumber="50">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F313D0FFAE5DFFC3FEABFEDEFE16FED6" box="[324,447,261,287]" name="Venezuela" pageId="5" pageNumber="50">Venezuela</collectingCountry>
, Territorio Federal Amazonas,
<quantity id="4CFC3D8AAE5DFFC3FCC4FEDEFCC7FED6" box="[811,878,261,287]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="50" unit="km" value="40.0">40km</quantity>
S Puerto Ayacucho, at El Tobogán, Coromoto.
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF2AFEF6FEACFDFE" pageId="5" pageNumber="50" type="diagnosis">
<paragraph id="8BBB906FAE5DFFC3FF2AFEF6FEACFDFE" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF2AFEF6FEECFE8E" bold="true" box="[197,325,301,327]" pageId="5" pageNumber="50">Diagnosis.</emphasis>
This species is smaller than
<taxonomicName id="4C04EBECAE5DFFC3FD40FEF6FCC3FE8F" box="[687,874,301,326]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="toboganensis">
<emphasis id="B9704C7DAE5DFFC3FD40FEF6FCC3FE8F" box="[687,874,301,326]" italics="true" pageId="5" pageNumber="50">F. toboganensis</emphasis>
</taxonomicName>
(TL &lt;
<quantity id="4CFC3D8AAE5DFFC3FC25FEF6FB88FE8E" box="[970,1057,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="50" unit="mm" value="1.3">1.3mm</quantity>
), but is similar in size (though slightly smaller) and external appearance to
<taxonomicName id="4C04EBECAE5DFFC3FD4EFE8EFC8AFEA7" box="[673,803,341,366]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="wheeleri">
<emphasis id="B9704C7DAE5DFFC3FD4EFE8EFC8AFEA7" box="[673,803,341,366]" italics="true" pageId="5" pageNumber="50">F. wheeleri</emphasis>
</taxonomicName>
.
<taxonomicName id="4C04EBECAE5DFFC3FCDDFE8EFB89FEA7" box="[818,1056,341,366]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B9704C7DAE5DFFC3FCDDFE8EFB89FEA7" box="[818,1056,341,366]" italics="true" pageId="5" pageNumber="50">Fontidessus ornatus</emphasis>
</taxonomicName>
has the prosternal process moderately elongate with the lateral margins in the apical portion straight and evenly tapered to the pointed apex (
<figureCitation id="133F8CEAAE5DFFC3FF70FE7EFF40FE76" box="[159,233,421,447]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 6</figureCitation>
). The pale regions on the elytron are diffuse and poorly demarcated and are yellow-brown unlike
<taxonomicName id="4C04EBECAE5DFFC3FA69FE7EFF55FE2F" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="wheeleri">
<emphasis id="B9704C7DAE5DFFC3FA69FE7EFF55FE2F" italics="true" pageId="5" pageNumber="50">F. wheeleri</emphasis>
</taxonomicName>
which have the elytral maculae brighter yellow and more clearly demarcated. The male genitalia are diagnostic with the median lobe slender and elongate with a subapical brush of setae on the ventral margin (
<figureCitation id="133F8CEAAE5DFFC3FF70FDC6FF5EFDFE" box="[159,247,541,567]" captionStart="FIGURES 8 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 17. Fontidessus species. 8 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF2AFD9EFB92FAB6" pageId="5" pageNumber="50" type="description">
<paragraph id="8BBB906FAE5DFFC3FF2AFD9EFDE9FD1E" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF2AFD9EFD8CFD96" bold="true" box="[197,549,581,607]" pageId="5" pageNumber="50">Description. Measurements.</emphasis>
TL =
<quantity id="4CFC3D8AAE5DFFC3FD9BFD9EFCACFD96" box="[628,773,581,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="5" pageNumber="50" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
, GW =
<quantity id="4CFC3D8AAE5DFFC3FC8AFD9EFC5EFD96" box="[869,1015,581,607]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="50" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, PW =
<quantity id="4CFC3D8AAE5DFFC3FBBDFD9EFB4DFD96" box="[1106,1252,581,607]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="50" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, HW =
<quantity id="4CFC3D8AAE5DFFC3FAABFD9EFF61FD4E" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="50" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 mm</quantity>
, EW =
<quantity id="4CFC3D8AAE5DFFC3FECBFDB6FE1DFD4E" box="[292,436,621,647]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="50" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.10.2 mm</quantity>
, TL/GW = 1.81.9, HW/EW = 1.51.6. Body (
<figureCitation id="133F8CEAAE5DFFC3FC05FDB6FB98FD4E" box="[1002,1073,621,647]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
) oval, elongate; lateral outline slightly continuous between pronotum and elytron; lateral margins of pronotum gently curved; lateral margins of elytron evenly and gently curved.
</paragraph>
<paragraph id="8BBB906FAE5DFFC3FF29FD3EFB0AFC56" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF29FD3EFEFCFD36" bold="true" box="[198,341,741,767]" pageId="5" pageNumber="50">Coloration.</emphasis>
Head yellow brown, darker in region near medial margins of eyes; pronotum yellow brown, diffusely brown medially along posterior margin and in narrow band along anterior margin; elytron brown with diffuse yellow pattern, yellow basally, laterally along about ¾ length of elytra and in subtriangular subapical area (
<figureCitation id="133F8CEAAE5DFFC3FEC4FC86FEDFFCBE" box="[299,374,861,887]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
). Ventral surfaces of thorax and abdomen brown except prosternum, prosternal process, propleuron and pronotal epipleuron yellow-brown; appendages yellow to yellow-brown.
</paragraph>
<paragraph id="8BBB906FAE5DFFC3FF29FC76FE35FB16" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF29FC76FE73FC0F" box="[198,474,941,966]" italics="true" pageId="5" pageNumber="50">Sculpture and structure</emphasis>
. Head with very fine, inconspicuous, irregular punctation, surface between punctures shiny with indistinct microsculpture in the form of small cells; eyes medium in size (
<figureCitation id="133F8CEAAE5DFFC3FB28FC0EFAB9FC26" box="[1223,1296,981,1007]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
, HW/EW = 1.51.6). Pronotal surface similar to that of head; with posterior angles obtuse; lateral bead slender, of even width throughout; pronotal striae finely incised, extending nearly 1/2 distance across pronotum (
<figureCitation id="133F8CEAAE5DFFC3FAAFFBFEFA24FBF6" box="[1344,1421,1061,1087]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
). Elytron with anterolateral angle obtuse, not extended anteriorly (
<figureCitation id="133F8CEAAE5DFFC3FC60FB96FC7EFBAE" box="[911,983,1101,1127]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
); surface similar to pronotum but with microsculpture more indistinct. Prosternal process relatively narrow, lateral margins tapered to narrowly rounded apex (
<figureCitation id="133F8CEAAE5DFFC3FEA4FB46FE3CFB7E" box="[331,405,1181,1207]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1010,1034]" captionTargetBox="[476,1111,479,980]" captionTargetId="figure@2.[473,1113,477,983]" captionTargetPageId="2" captionText="FIGURES 1 7. Fontidessus species. 1 3) Dorsal habitus, 1) F. toboganensis, 2) F. ornatus, 3) F. wheeleri. 4) F. toboganensis, left metatrochanter and metafemur, anterior aspect. 5 7) prosternal process, 5) F. toboganensis, 6) F. ornatus, 7) F. wheeleri. Scale bar a = 1.0 mm for Figs 1 4. Scale bar b = 0.1 mm for Figs 5 7." httpUri="https://zenodo.org/record/183127/files/figure.png" pageId="5" pageNumber="50">Fig. 6</figureCitation>
); metacoxal process with lateral lobe minute. Pro- and mesotarsi relatively narrow in both male and female.
</paragraph>
<paragraph id="8BBB906FAE5DFFC3FF29FB36FB92FAB6" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF29FB36FED1FACF" box="[198,376,1261,1286]" italics="true" pageId="5" pageNumber="50">Male genitalia</emphasis>
. Median lobe in ventral aspect slender, elongate and with lateral margins parallel to pointed apex (
<figureCitation id="133F8CEAAE5DFFC3FED0FACEFE3FFAE6" box="[319,406,1301,1327]" captionStart="FIGURES 8 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 17. Fontidessus species. 8 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">Fig. 11</figureCitation>
); in lateral aspect moderately broad at base, curved, with apical portion evenly tapered to pointed apex, subapically with small brush of setae (
<figureCitation id="133F8CEAAE5DFFC3FCD4FAE6FC3FFA9E" box="[827,918,1341,1367]" captionStart="FIGURES 8 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 17. Fontidessus species. 8 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">Fig. 12</figureCitation>
). Lateral lobe elongate, expanded apically, broadly rounded at apex, with series of long setae along dorsal margin (
<figureCitation id="133F8CEAAE5DFFC3FC32FABEFB84FAB6" box="[989,1069,1381,1407]" captionStart="FIGURES 8 17" captionStartId="4.[151,269,1053,1077]" captionTargetBox="[469,1097,360,1021]" captionTargetId="figure@4.[463,1103,354,1031]" captionTargetPageId="4" captionText="FIGURES 8 17. Fontidessus species. 8 10) F. toboganensis, male genitalia, 8) median lobe, dorsal aspect, 9) median lobe, right lateral aspect, 10) right lateral lobe, right lateral aspect. 11 13) F. ornatus, male genitalia, 11) median lobe and left lateral lobe, dorsal aspect, 12) median lobe, right lateral aspect, 13) right lateral lobe, right lateral aspect. 14 16) F. wheeleri, male genitalia, 14) median lobe and left lateral lobe, dorsal aspect, 15) median lobe, left lateral aspect, 16) right lateral lobe, right lateral aspect. 17) F. toboganensis, female genitalia, ventral aspect. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/183128/files/figure.png" pageId="5" pageNumber="50">Fig 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF2AFA56FDF9FA06" pageId="5" pageNumber="50" type="etymology">
<paragraph id="8BBB906FAE5DFFC3FF2AFA56FDF9FA06" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF2AFA56FEFBFA6E" bold="true" box="[197,338,1421,1447]" pageId="5" pageNumber="50">Etymology.</emphasis>
The species name is derived from the Greek root
<emphasis id="B9704C7DAE5DFFC3FC70FA56FC7FFA6F" box="[927,982,1421,1446]" italics="true" pageId="5" pageNumber="50">orno</emphasis>
, meaning “adorned,” for the attractive coloration characterizing this species.
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF2AFA06FD5EF9D6" pageId="5" pageNumber="50" type="distribution">
<paragraph id="8BBB906FAE5DFFC3FF2AFA06FD5EF9D6" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF2AFA06FE57FA3E" bold="true" box="[197,510,1501,1527]" pageId="5" pageNumber="50">Distribution and habitat.</emphasis>
The species is known only from the
<typeStatus id="54BF2ECDAE5DFFC3FC5EFA06FC4DFA3E" box="[945,996,1501,1527]" pageId="5" pageNumber="50">type</typeStatus>
locality. Label data indicated the following habitats; “seep,” and “bare rock streambed.”
</paragraph>
</subSubSection>
<subSubSection id="C31EC3E4AE5DFFC3FF2AF9F6FC9FF8C6" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="8BBB906FAE5DFFC3FF2AF9F6FC9FF8C6" blockId="5.[151,1436,261,1807]" pageId="5" pageNumber="50">
<emphasis id="B9704C7DAE5DFFC3FF2AF9F6FE69F98E" bold="true" box="[197,448,1581,1607]" pageId="5" pageNumber="50">Material examined.</emphasis>
<typeStatus id="54BF2ECDAE5DFFC3FE20F9F6FDC3F98E" box="[463,618,1581,1607]" pageId="5" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
in
<collectionCode id="ED1508AAAE5DFFC3FD72F9F6FD5FF98E" LSID="urn:lsid:biocol.org:col:34871" box="[669,758,1581,1607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="50">USNM</collectionCode>
: male labeled, “
<collectingCountry id="F313D0FFAE5DFFC3FC2CF9F6FBD0F98E" box="[963,1145,1581,1607]" name="Venezuela" pageId="5" pageNumber="50">VENEZUELA</collectingCountry>
: T.F. Amazonas Puerto Ayacucho (
<quantity id="4CFC3D8AAE5DFFC3FEF5F98EFECCF9A6" box="[282,357,1621,1647]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="50" unit="km" value="40.0">40km</quantity>
.S) at Tobogán,
<date id="FFBAB6AFAE5DFFC3FDF7F98EFD35F9A6" box="[536,668,1621,1647]" pageId="5" pageNumber="50" value="1986-02-22">22Feb1986</date>
P.J. Spangler, Colln.#6/ Bare rock streambed in sunlight; beneath wet leaves at margin/
<typeStatus id="54BF2ECDAE5DFFC3FE71F9A6FD97F95E" box="[414,574,1661,1687]" pageId="5" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
:
<taxonomicName id="4C04EBECAE5DFFC3FDA5F9A6FC7CF95E" authority="Miller, 2008" authorityName="Miller" authorityYear="2008" box="[586,981,1661,1687]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B9704C7DAE5DFFC3FDA5F9A6FC90F95F" box="[586,825,1661,1686]" italics="true" pageId="5" pageNumber="50">Fontidessus ornatus</emphasis>
Miller, 2008
</taxonomicName>
[red label with double black line bor- der].”
<typeStatus id="54BF2ECDAE5DFFC3FF09F97EFE38F976" box="[230,401,1701,1727]" pageId="5" pageNumber="50" type="paratype">PARATYPES</typeStatus>
, 14 total; 3 with same label data as
<typeStatus id="54BF2ECDAE5DFFC3FCB1F97EFC61F976" box="[862,968,1701,1727]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
; 9,
<collectingCountry id="F313D0FFAE5DFFC3FC13F97EFBDEF976" box="[1020,1143,1701,1727]" name="Venezuela" pageId="5" pageNumber="50">Venezuela</collectingCountry>
, T.F. Amazonas, Puerto Ayacucho,
<quantity id="4CFC3D8AAE5DFFC3FEF4F916FEF7F92E" box="[283,350,1741,1767]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="50" unit="km" value="40.0">40km</quantity>
S El Tobogán, Caño Coromoto,
<date id="FFBAB6AFAE5DFFC3FD09F916FC04F92E" box="[742,941,1741,1767]" pageId="5" pageNumber="50" value="1989-01-23">23 January 1989</date>
, seep at upper shelter, P.J. Spangler, R.A. Faitoute, and C.B. Barr; 2, same except
<date id="FFBAB6AFAE5DFFC3FD83F92EFC98F8C6" box="[620,817,1781,1807]" pageId="5" pageNumber="50" value="1989-01-26">26 January 1989</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>