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<mods:title id="29F343EBBC370C49DDB8B9D15FA0C6B4">Shedding light into Brazilian subterranean isopods (Isopoda, Oniscidea): expanding distribution data and describing new taxa</mods:title>
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Campos-Filho, Sfenthourakis &amp; Bichuette
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(
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, state of
<collectingRegion id="49C960D1921F633BFE81FB88FE29EB51" box="[358,416,1086,1110]" country="Brazil" name="Bahia" pageId="38" pageNumber="567">Bahia</collectingRegion>
;
<geoCoordinate id="EE39C8F4921F633BFE4CFB88FDB8EB51" box="[427,561,1086,1110]" degrees="13" direction="south" minutes="56" orientation="latitude" pageId="38" pageNumber="567" precision="1" seconds="46.4" value="-13.946222">13°5646.4”S</geoCoordinate>
,
<geoCoordinate id="EE39C8F4921F633BFDDBFB88FD31EB51" box="[572,696,1086,1110]" degrees="44" direction="west" minutes="11" orientation="longitude" pageId="38" pageNumber="567" precision="15" seconds="12" value="-44.18667">44°1112”W</geoCoordinate>
;
<elevation id="00204900921F633BFD21FB88FF38EB76" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" pageId="38" pageNumber="567" unit="m" value="544.0">
<quantity id="4CF503D6921F633BFD21FB88FC8EEB51" box="[710,775,1086,1110]" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" pageId="38" pageNumber="567" unit="m" value="544.0">544 m</quantity>
a.s.l.
</elevation>
;
<date id="FFB388F3921F633BFF59FBEFFEA0EB76" box="[190,297,1113,1137]" pageId="38" pageNumber="567" value="2020-10-15">
<collectingDate id="EFF7711B921F633BFF59FBEFFEA0EB76" box="[190,297,1113,1137]" pageId="38" pageNumber="567" value="2020-10-15">15.X.2020</collectingDate>
</date>
;
<collectorName id="26F8CBE5921F633BFED3FBEFFE5FEB76" box="[308,470,1113,1137]" pageId="38" pageNumber="567">M. E. Bichuette</collectorName>
,
<collectorName id="26F8CBE5921F633BFE06FBEFFDD0EB77" box="[481,601,1113,1136]" pageId="38" pageNumber="567">D. F. Torres</collectorName>
,
<collectorName id="26F8CBE5921F633BFD83FBEFFD44EB76" box="[612,717,1113,1137]" pageId="38" pageNumber="567">J. S. Gallo</collectorName>
,
<collectorName id="26F8CBE5921F633BFD3FFBEFFF36EB8C" pageId="38" pageNumber="567">L. S. Horta</collectorName>
&amp;
<collectorName id="26F8CBE5921F633BFF06FBC2FED8EB8C" box="[225,337,1139,1163]" pageId="38" pageNumber="567">J. E. Gallão</collectorName>
leg.;
<specimenCode id="DBAB0648921F633BFE61FBC5FE70EB8C" box="[390,505,1139,1163]" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" pageId="38" pageNumber="567" type="Museum">LES 28002</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB2AE33921F633BFF63FB38FE2CEBC7" blockId="38.[131,775,1060,1216]" pageId="38" pageNumber="567">
<materialsCitation id="3B65A46E921F633BFF63FB38FE28EBC7" ID-GBIF-Occurrence="4428725353" collectingDate="2020-10-15" collectionCode="LES" collectorName=" M. E. Bichuette &amp; D. F. Torres &amp; J. S. Gallo &amp; L. S. Horta &amp; J. E. Gallao" country="Brazil" elevation="544" latitude="-13.946222" location="Feira da Mata" longLatPrecision="15" longitude="-44.18667" municipality="Gruna Boca da Lapa cave" pageId="38" pageNumber="567" specimenCode="LES 28003" specimenCount="6" specimenCount-female="6" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="B9797221921F633BFF63FB38FEA0EBA0" bold="true" box="[132,297,1166,1191]" pageId="38" pageNumber="567">
<typeStatus id="54B61091921F633BFF63FB38FF6FEBA1" box="[132,230,1166,1190]" pageId="38" pageNumber="567" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F31AEEA3921F633BFF08FB39FEA0EBA0" box="[239,297,1167,1191]" name="Brazil" pageId="38" pageNumber="567">Brazil</collectingCountry>
</emphasis>
<specimenCount id="9D0B65BA921F633BFEDAFB38FED4EBAE" box="[317,349,1166,1193]" pageId="38" pageNumber="567" type="female">6♀</specimenCount>
(one with parts in micropreparations); same data as holotype;
<specimenCode id="DBAB0648921F633BFEC8FB1EFE28EBC7" box="[303,417,1192,1216]" collectionCode="LES" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13487" lsid="urn:lsid:biocol.org:col:13487" name="Leeds Museums and Galleries" pageId="38" pageNumber="567" type="Museum">LES 28003</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C317FDB8921F633BFF63FB57FE5DEA7A" pageId="38" pageNumber="567" type="etymology">
<paragraph id="8BB2AE33921F633BFF63FB57FE5DEA7A" blockId="38.[132,775,1249,1405]" pageId="38" pageNumber="567">
<smallCapsWord id="8D5438EF921F633BFF63FB57FF77EBF0" baselines="1266,1267" box="[132,254,1249,1272]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="38" pageNumber="567">ETYMOLOGY</smallCapsWord>
. — The new species is named after Exu, a divinity of the Candomblé Afro-Brazilian religion. In the Candomblé, Exu is the messenger of the orixás divinities, transiting between the world of humans and the gods. He is considered the owner of the paths, the crossroads, the perspectives, the past, and the future, and he is responsible for order and disorder.
</paragraph>
</subSubSection>
<subSubSection id="C317FDB8921F633BFF63FA05FADEEA69" pageId="38" pageNumber="567" type="description">
<paragraph id="8BB2AE33921F633BFF63FA05FE9CEACC" blockId="38.[130,777,1459,2027]" box="[132,277,1459,1485]" pageId="38" pageNumber="567">
<smallCapsWord id="8D5438EF921F633BFF63FA05FE9CEACC" baselines="1478,1478" box="[132,277,1459,1485]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="38" pageNumber="567">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BB2AE33921F633BFF63FA65FADEEA69" blockId="38.[130,777,1459,2027]" lastBlockId="38.[810,1457,215,1390]" pageId="38" pageNumber="567">
Maximum body length:
<quantity id="4CF503D6921F633BFE7FFA65FE7CEAEA" box="[408,501,1491,1518]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="38" pageNumber="567" unit="mm" value="6.5">6.5 mm</quantity>
. Color brown with typical muscle insertion spots; cephalon, pereonite 1, pereonite 2-7 posterior portions, pleon, telson and uropods strongly pigmented. Body (
<figureCitation id="1336B2B6921F633BFE2BF984FDBEE94A" box="[460,567,1586,1613]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27A</figureCitation>
) strongly convex, bearing triangular tubercles, arranged as follows: vertex of cephalon with eight tubercles in two rows; pereonite 1 with 13 tubercles in three rows; pereonite 2 with 11 tubercles in two rows; pereonites 3 and 4 with nine tubercles in two rows; pereonites 5-7 with seven tubercles in one row; pleonites 5 and telson with two tubercles on median portion (
<figureCitation id="1336B2B6921F633BFF02F8A4FECCE82B" box="[229,325,1810,1836]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27B</figureCitation>
). Dorsal surface with semi-circular scale-setae (
<figureCitation id="1336B2B6921F633BFF2DF887FEBAE84B" box="[202,307,1841,1868]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27C</figureCitation>
).
<emphasis id="B9797221921F633BFEAFF887FE7AE84C" box="[328,499,1841,1867]" italics="true" pageId="38" pageNumber="567">Noduli laterales</emphasis>
short, inserted on outer part of second tubercle from lateral margins (
<figureCitation id="1336B2B6921F633BFD76F8E7FD71E86B" box="[657,760,1873,1900]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27A</figureCitation>
). Cephalon (
<figureCitation id="1336B2B6921F633BFEE6F8C7FE2FE88C" box="[257,422,1905,1932]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27A, D, E</figureCitation>
) with frontal shield prominent, distinctly protruding above vertex; eyes with 10 ommatidia; pereonite 1-7 epimera (
<figureCitation id="1336B2B6921F633BFE6BF807FE7AE8CC" box="[396,499,1969,1995]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27A</figureCitation>
, E-H) strongly concave; pereonite 1 epimera with rounded, lateral margin grooved on posterior corner for about one quarter of its length, inner lobe of schisma rounded, not extending beyond posterior margin of outer lobe; pereonite 2-7 epimera (
<figureCitation id="1336B2B6921F633BFAA1FEA1FCDDEE56" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27F, I, J</figureCitation>
) triangular and elongated; pereonite 2 epimera with narrow and rounded lobe directed outwards (
<figureCitation id="1336B2B6921F633BFAAFFEE1FCB5EE96" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27F, J</figureCitation>
); pereonites 3 and 4 without lobe; pereonites 5-7 with ventral sulcus (
<figureCitation id="1336B2B6921F633BFC34FE20FBBFEEB6" box="[979,1078,406,433]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27F</figureCitation>
). Pleonite 3-5 epimera (
<figureCitation id="1336B2B6921F633BFAA5FE20FA2EEEB6" box="[1346,1447,406,433]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27K</figureCitation>
) triangular; telson (
<figureCitation id="1336B2B6921F633BFC1EFE00FBF2EED6" box="[1017,1147,438,465]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27K, L</figureCitation>
) dorsum slightly depressed, distal margin slightly convex. Antennula (
<figureCitation id="1336B2B6921F633BFAE6FE60FAE7EEF7" box="[1281,1390,470,496]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27M</figureCitation>
) with proximal and distal articles similar in length, distal article bearing about six aesthetascs inserted apically and subapically. Antenna (
<figureCitation id="1336B2B6921F633BFC38FD80FBC1ED57" box="[991,1096,566,592]" captionStart="FIG" captionStartId="39.[132,143,1939,1957]" captionTargetBox="[200,1387,215,1888]" captionTargetId="figure-16@39.[199,1388,215,1896]" captionTargetPageId="39" captionText="FIG. 27. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, lateral habitus; B, tubercles schem; C, scale-seta; D, cephalon, back view; E, cephalon and pereonites 1 and 2, frontal view; F, pereonite 1-7 epimera, ventral view; G, pereonite 1 epimeron, dorsal view; H, pereonite 1 epimeron, ventral view;I, pereonite 2 epimeron, dorsal view;J, pereonite 2 epimeron, ventral view; K, pereonite 7, pleon, telson and uropods,dorsal view, L, telson; M, antennula; N, antenna. Scale bar: A, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="38" pageNumber="567">Fig. 27N</figureCitation>
) short, not surpassing posterior margin of pereonite 1 when extended backwards; flagellum distal article about three times as long as first, bearing two lateral aesthetascs. Mandibles (
<figureCitation id="1336B2B6921F633BFB75FD20FA94EDB7" box="[1170,1309,662,688]" captionStart="FIG" captionStartId="40.[133,144,1293,1311]" captionTargetBox="[132,1448,222,1249]" captionTargetId="figure-394@40.[132,1455,214,1250]" captionTargetPageId="40" captionText="FIG. 28. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, left mandible; B, right mandible; C, maxillula; D, maxilla; E, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10027525" httpUri="https://zenodo.org/record/10027525/files/figure.png" pageId="38" pageNumber="567">Fig. 28A, B</figureCitation>
) with molar penicil dichotomized. Maxillula (
<figureCitation id="1336B2B6921F633BFB4DFD03FA9BEDD7" box="[1194,1298,693,720]" captionStart="FIG" captionStartId="40.[133,144,1293,1311]" captionTargetBox="[132,1448,222,1249]" captionTargetId="figure-394@40.[132,1455,214,1250]" captionTargetPageId="40" captionText="FIG. 28. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, left mandible; B, right mandible; C, maxillula; D, maxilla; E, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10027525" httpUri="https://zenodo.org/record/10027525/files/figure.png" pageId="38" pageNumber="567">Fig. 28C</figureCitation>
) inner endite bearing two penicils, distal margin rounded; outer endite of 4+6 teeth, outer set apically entire. Maxilla (
<figureCitation id="1336B2B6921F633BFAD9FD43FA2FEC08" box="[1342,1446,757,783]" captionStart="FIG" captionStartId="40.[133,144,1293,1311]" captionTargetBox="[132,1448,222,1249]" captionTargetId="figure-394@40.[132,1455,214,1250]" captionTargetPageId="40" captionText="FIG. 28. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, left mandible; B, right mandible; C, maxillula; D, maxilla; E, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10027525" httpUri="https://zenodo.org/record/10027525/files/figure.png" pageId="38" pageNumber="567">Fig. 28D</figureCitation>
) inner lobe covered with thick setae; outer lobe three times as wide as inner lobe covered with thin setae. Maxilliped (
<figureCitation id="1336B2B6921F633BFCD2FCE3FC1EEC68" box="[821,919,853,879]" captionStart="FIG" captionStartId="40.[133,144,1293,1311]" captionTargetBox="[132,1448,222,1249]" captionTargetId="figure-394@40.[132,1455,214,1250]" captionTargetPageId="40" captionText="FIG. 28. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, left mandible; B, right mandible; C, maxillula; D, maxilla; E, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.10027525" httpUri="https://zenodo.org/record/10027525/files/figure.png" pageId="38" pageNumber="567">Fig. 28E</figureCitation>
) basis rectangular bearing sparse setae; proximal article of palp with two distinct setae; endite subrectangular, medial seta surpassing distal margin, ventral sulcus setose ending with short seta. Uropod (
<figureCitation id="1336B2B6921F633BFB4DFC03FABAECC8" box="[1194,1331,949,975]" captionStart="FIG" captionStartId="41.[132,143,1985,2003]" captionTargetBox="[132,1447,216,1813]" captionTargetId="figure-16@41.[132,1455,216,1813]" captionTargetPageId="41" captionText="FIG. 29. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, uropod, frontal view; B, uropod; C, pereopod 1; D, pereopod 7; E, pleopod 2 exopod; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10130514" httpUri="https://zenodo.org/record/10130514/files/figure.png" pageId="38" pageNumber="567">Fig. 29A, B</figureCitation>
) protopod enlarged on basal part, distal part subtriangular, broad, medial margin concave; endopod short bearing setae; exopod short inserted dorsally on distinct lobe near medial margin. Pereopods 1-7 (
<figureCitation id="1336B2B6921F633BFBADFB82FB55EB49" box="[1098,1244,1076,1102]" captionStart="FIG" captionStartId="41.[132,143,1985,2003]" captionTargetBox="[132,1447,216,1813]" captionTargetId="figure-16@41.[132,1455,216,1813]" captionTargetPageId="41" captionText="FIG. 29. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, uropod, frontal view; B, uropod; C, pereopod 1; D, pereopod 7; E, pleopod 2 exopod; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10130514" httpUri="https://zenodo.org/record/10130514/files/figure.png" pageId="38" pageNumber="567">Fig. 29C, D</figureCitation>
) with sparse setae along sternal margin; pereopod 1 carpus with longitudinal antenna-grooming brush; dactylus of two claws, ungual and dactylar setae simple, not surpassing outer claw. Pleopod 1 exopod (not drawn) sub-circular, twice as wide as long, outer portion narrow, median portion rounded bearing five small setae. Pleopod 2 exopod (
<figureCitation id="1336B2B6921F633BFB60FB45FB60EA09" box="[1159,1257,1267,1294]" captionStart="FIG" captionStartId="41.[132,143,1985,2003]" captionTargetBox="[132,1447,216,1813]" captionTargetId="figure-16@41.[132,1455,216,1813]" captionTargetPageId="41" captionText="FIG. 29. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, uropod, frontal view; B, uropod; C, pereopod 1; D, pereopod 7; E, pleopod 2 exopod; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10130514" httpUri="https://zenodo.org/record/10130514/files/figure.png" pageId="38" pageNumber="567">Fig. 29E</figureCitation>
) triangular, outer margin concave. Pleopod 3 and 4 exopods as in
<figureCitation id="1336B2B6921F633BFAD5FAA5FCCAEA4A" captionStart="FIG" captionStartId="41.[132,143,1985,2003]" captionTargetBox="[132,1447,216,1813]" captionTargetId="figure-16@41.[132,1455,216,1813]" captionTargetPageId="41" captionText="FIG. 29. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, uropod, frontal view; B, uropod; C, pereopod 1; D, pereopod 7; E, pleopod 2 exopod; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10130514" httpUri="https://zenodo.org/record/10130514/files/figure.png" pageId="38" pageNumber="567">Figure 29F, G</figureCitation>
, respectively. Pleopod 5 exopod (
<figureCitation id="1336B2B6921F633BFB5DFA85FAADEA4A" box="[1210,1316,1331,1357]" captionStart="FIG" captionStartId="41.[132,143,1985,2003]" captionTargetBox="[132,1447,216,1813]" captionTargetId="figure-16@41.[132,1455,216,1813]" captionTargetPageId="41" captionText="FIG. 29. — Diploexochus exu Campos-Filho, Sfenthourakis &amp; Bichuette, n. sp., female paratype LES 28003: A, uropod, frontal view; B, uropod; C, pereopod 1; D, pereopod 7; E, pleopod 2 exopod; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10130514" httpUri="https://zenodo.org/record/10130514/files/figure.png" pageId="38" pageNumber="567">Fig. 29H</figureCitation>
) rhomboid, distal margin almost straight bearing many setae.
</paragraph>
</subSubSection>
<subSubSection id="C317FDB8921F633BFCCAFA22FC18EAAB" box="[813,913,1428,1454]" pageId="38" pageNumber="567" type="discussion">
<paragraph id="8BB2AE33921F633BFCCAFA22FC18EAAB" blockId="38.[811,1457,1428,2028]" box="[813,913,1428,1454]" pageId="38" pageNumber="567">
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</paragraph>
</subSubSection>
<subSubSection id="C317FDB8921F6335FCCAFA05FDCAE8EB" lastPageId="40" lastPageNumber="569" pageId="38" pageNumber="567" type="reference_group">
<paragraph id="8BB2AE33921F6335FCCAFA05FD22E86B" blockId="38.[811,1457,1428,2028]" lastBlockId="40.[130,777,1427,2028]" lastPageId="40" lastPageNumber="569" pageId="38" pageNumber="567">
Within
<taxonomicName id="4C0DD5B0921F633BFC60FA05FC74EACA" authority="Morch" box="[903,1021,1459,1485]" class="Gastropoda" family="Cassidae" genus="Oniscidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="38" pageNumber="532" phylum="Mollusca" rank="genus">Oniscidea</taxonomicName>
, most of the morphological characters used in taxonomy to describe and distinguish species are those of males (e.g.,
<bibRefCitation id="EF9CD3C2921F633BFBEDFA45FB66E90A" author="TAITI S. &amp; FERRARA F." box="[1034,1263,1523,1549]" pageId="38" pageNumber="567" pagination="249 - 300" refId="ref39384" refString="TAITI S. &amp; FERRARA F. 1980. - ‰ uovi studi sugli isopodi terrestri dell'Arcipelago Toscano. Redia 63: 249 - 300." type="journal article" year="1980">Taiti &amp; Ferrara 1980</bibRefCitation>
; Leistikow 1999;
<bibRefCitation id="EF9CD3C2921F633BFCCAF9A4FBE5E92A" author="CAMPOS-FILHO I. S. &amp; TAITI S. &amp; ARAUJO P. B." box="[813,1132,1554,1581]" pageId="38" pageNumber="567" pagination="1 - 73" refId="ref35072" refString="CAMPOS-FILHO I. S., TAITI S. &amp; ARAUJO P. B. 2015 a. - Taxonomic revision of the genus Benthana Budde-Lund, 1908 (Isopoda: Oniscidea: Philosciidae). Zootaxa 4022 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4022.1.1" type="journal article" year="2015">
Campos-Filho
<emphasis id="B9797221921F633BFC38F9A5FB90E92B" box="[991,1049,1554,1580]" italics="true" pageId="38" pageNumber="567">et al.</emphasis>
2015a
</bibRefCitation>
;
<bibRefCitation id="EF9CD3C2921F633BFB9AF9A4FAF0E92A" author="REBOLEIRA A. S. P. S. &amp; GONCALVES P. O. &amp; TAITI S." box="[1149,1401,1554,1581]" pageId="38" pageNumber="567" pagination="1 - 61" refId="ref38027" refString="REBOLEIRA A. S. P. S., GONCALVES P. O. &amp; TAITI S. 2015. - The cavernicolous Oniscidea (Crustacea: Isopoda) of Portugal. European Journal of Taxonomy 161: 1 - 61. https: // doi. org / 10.5852 / ejt. 2015.161" type="journal article" year="2015">
Reboleira
<emphasis id="B9797221921F633BFB10F9A5FAB8E92B" box="[1271,1329,1554,1580]" italics="true" pageId="38" pageNumber="567">et al.</emphasis>
2015
</bibRefCitation>
). As mentioned previously, though, conglobating representatives, especially within the
<taxonomicName id="4C0DD5B0921F633BFBB9F9E4FB7CE96B" ID-CoL="7NFZG" authorityName="Brandt" authorityYear="1831" box="[1118,1269,1618,1644]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="38" pageNumber="567" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
, male characters are not always useful, e.g., in
<taxonomicName id="4C0DD5B0921F633BFB97F9C4FB54E98B" ID-CoL="7NTYG" authorityName="Verhoeff" authorityYear="1942" box="[1136,1245,1650,1676]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="38" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="B9797221921F633BFB97F9C4FB54E98B" box="[1136,1245,1650,1676]" italics="true" pageId="38" pageNumber="567">Ctenorillo</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0DD5B0921F633BFB0CF9C4FCE0E9AB" ID-CoL="7Q5GF" authority="Schultz, 1982" authorityName="Schultz" authorityYear="1982" class="Malacostraca" family="Armadillidae" genus="Tuberillo" kingdom="Animalia" order="Isopoda" pageId="38" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="B9797221921F633BFB0CF9C4FAC7E98B" box="[1259,1358,1650,1676]" italics="true" pageId="38" pageNumber="567">Tuberillo</emphasis>
<bibRefCitation id="EF9CD3C2921F633BFAB3F9C4FCE0E9AB" author="SCHULTZ G. A." pageId="38" pageNumber="567" pagination="101 - 117" refId="ref38728" refString="SCHULTZ G. A. 1982. - Terrestrial isopod crustaceans (Oniscoidea) from Mulu Caves, Sarawak, Borneo. Journal of natural History 16: 101 - 117. https: // doi. org / 10.1080 / 00222938200770081" type="journal article" year="1982">Schultz, 1982</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0DD5B0921F633BFC43F924FB8FE9AB" ID-CoL="7Q68J" authorityName="Verhoeff" authorityYear="1928" box="[932,1030,1682,1708]" class="Malacostraca" family="Armadillidae" genus="Venezillo" kingdom="Animalia" order="Isopoda" pageId="38" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="B9797221921F633BFC43F924FB8FE9AB" box="[932,1030,1682,1708]" italics="true" pageId="38" pageNumber="567">Venezillo</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9CD3C2921F633BFBFFF925FB59E9AB" author="VAN NAME W. G." box="[1048,1232,1682,1709]" pageId="38" pageNumber="567" pagination="1 - 535" refId="ref40015" refString="VAN NAME W. G. 1936. - The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535. http: // hdl. handle. net / 2246 / 1185" type="journal article" year="1936">
<collectingRegion id="49C960D1921F633BFBFFF925FBCCE9AA" box="[1048,1093,1683,1709]" country="Turkey" name="Van" pageId="38" pageNumber="567">Van</collectingRegion>
Name 1936
</bibRefCitation>
;
<bibRefCitation id="EF9CD3C2921F633BFB39F924FC13E9CB" author="SCHMALFUSS H. &amp; FERRARA F." pageId="38" pageNumber="567" pagination="111 - 157" refId="ref38417" refString="SCHMALFUSS H. &amp; FERRARA F. 1983. - Terrestrial isopods from West Africa, Part 3: Family Armadillidae Verhoeff, 1917. Monitore zoologico italiano, Nuova Serie, Supplemento 18: 111 - 157. https: // doi. org / 10.1080 / 00269786.1983.11758568" type="journal article" year="1983">Schmalfuss &amp; Ferrara 1983</bibRefCitation>
;
<bibRefCitation id="EF9CD3C2921F633BFC40F904FB1EE9CB" author="TAITI S. &amp; GRUBER G. A." box="[935,1175,1714,1740]" pageId="38" pageNumber="567" pagination="205 - 230" refId="ref39449" refString="TAITI S. &amp; GRUBER G. A. 2010. - The genus Tuberillo Schultz 1982 (Crustacea Oniscidea Armadillidae) with descriptions of four new species. Tropical Zoology 23: 205 - 230." type="journal article" year="2010">Taiti &amp; Gruber 2010</bibRefCitation>
). Thus, other characters, such as the conglobating mechanism, the presence and arrangement of dorsal tubercles (in tuberculated animals), and the shape of the lateral schisma of pereonite 1 (if present), are useful to distinguish species, even in the absence of males.
<taxonomicName id="4C0DD5B0921F633BFBD4F8E7FBFAE88C" authority="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityName="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityYear="2023" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="38" pageNumber="567" phylum="Arthropoda" rank="species" species="exu" status="sp. nov.">
<emphasis id="B9797221921F633BFBD4F8E7FB7FE86C" box="[1075,1270,1873,1899]" italics="true" pageId="38" pageNumber="567">Diploexochus exu</emphasis>
Campos-Filho, Sfenthourakis &amp; Bichuette
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A921F633BFB66F8C4FB4DE88B" box="[1153,1220,1906,1932]" pageId="38" pageNumber="567" rank="species">n. sp.</taxonomicNameLabel>
easily differs in the shape and arrangement of dorsal tubercles. Moreover, it can be distinguished in having the pereonite 1 epimera with lateral groove of about one quarter of the length of the epimeron (vs all epimeron grooved in
<taxonomicName id="4C0DD5B092116335FD94FA22FD76EAAA" authority="Brandt, 1833" box="[627,767,1427,1453]" class="Malacostraca" family="Armadillidae" genus="Armadillo" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B979722192116335FD94FA22FD76EAAA" box="[627,767,1427,1453]" italics="true" pageId="40" pageNumber="569">D. echinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0DD5B092116335FF63FA02FD35EACA" authority="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityName="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityYear="2023" box="[132,700,1459,1485]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="carrapicho" status="sp. nov.">
<emphasis id="B979722192116335FF63FA02FE9DEACA" box="[132,276,1459,1485]" italics="true" pageId="40" pageNumber="569">D. carrapicho</emphasis>
Campos-Filho, López-Orozco &amp; Taiti
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A92116335FD22FA05FC8BEACA" box="[709,770,1459,1485]" pageId="40" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
, and
<taxonomicName id="4C0DD5B092116335FF50FA62FEBCEAEA" ID-CoL="117af273-11c6-441f-ae7e-288891d52caf" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[183,309,1492,1517]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="B979722192116335FF50FA62FEBCEAEA" box="[183,309,1492,1517]" italics="true" pageId="40" pageNumber="569">D. spinatus</emphasis>
</taxonomicName>
; about two thirds of the epimeron lenth in
<taxonomicName id="4C0DD5B092116335FF44FA45FEAAE90A" ID-CoL="1a487283-85d7-4168-b944-d30c317b5ab3" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[163,291,1523,1549]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B979722192116335FF44FA45FEAAE90A" box="[163,291,1523,1549]" italics="true" pageId="40" pageNumber="569">D. obscurus</emphasis>
</taxonomicName>
), telson with distal margin slightly convex (vs concave in
<taxonomicName id="4C0DD5B092116335FED4F9A5FE48E92B" authority="Brandt, 1833" box="[307,449,1554,1580]" class="Malacostraca" family="Armadillidae" genus="Armadillo" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B979722192116335FED4F9A5FE48E92B" box="[307,449,1554,1580]" italics="true" pageId="40" pageNumber="569">D. echinatus</emphasis>
</taxonomicName>
; slightly concave in
<taxonomicName id="4C0DD5B092116335FD52F9A5FD24E94B" authority="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityName="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityYear="2023" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="carrapicho" status="sp. nov.">
<emphasis id="B979722192116335FD52F9A5FF5EE94B" italics="true" pageId="40" pageNumber="569">D. carrapicho</emphasis>
Campos-Filho, López-Orozco &amp; Taiti
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A92116335FD5BF985FC89E94A" box="[700,768,1587,1613]" pageId="40" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
; slightly convex in
<taxonomicName id="4C0DD5B092116335FEB8F9E5FE6AE96B" ID-CoL="1a487283-85d7-4168-b944-d30c317b5ab3" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[351,483,1618,1644]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B979722192116335FEB8F9E5FE6AE96B" box="[351,483,1618,1644]" italics="true" pageId="40" pageNumber="569">D. obscurus</emphasis>
</taxonomicName>
; straight in
<taxonomicName id="4C0DD5B092116335FD90F9E5FD71E96B" ID-CoL="117af273-11c6-441f-ae7e-288891d52caf" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[631,760,1619,1644]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="B979722192116335FD90F9E5FD71E96B" box="[631,760,1619,1644]" italics="true" pageId="40" pageNumber="569">D. spinatus</emphasis>
</taxonomicName>
), eyes composed of about 10 ommatidia (vs about
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<taxonomicName id="4C0DD5B092116335FF61F925FE98E9AB" authority="Brandt, 1833" box="[134,273,1682,1708]" class="Malacostraca" family="Armadillidae" genus="Armadillo" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B979722192116335FF61F925FE98E9AB" box="[134,273,1682,1708]" italics="true" pageId="40" pageNumber="569">D. echinatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0DD5B092116335FEACF925FE40E9AB" ID-CoL="117af273-11c6-441f-ae7e-288891d52caf" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[331,457,1683,1708]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="B979722192116335FEACF925FE40E9AB" box="[331,457,1683,1708]" italics="true" pageId="40" pageNumber="569">D. spinatus</emphasis>
</taxonomicName>
;
<quantity id="4CF503D692116335FE3FF924FD9CE9AB" box="[472,533,1682,1708]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="40" pageNumber="569" unit="in" value="16.0">16 in</quantity>
<taxonomicName id="4C0DD5B092116335FDFBF925FD14E9AB" ID-CoL="1a487283-85d7-4168-b944-d30c317b5ab3" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[540,669,1682,1708]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B979722192116335FDFBF925FD14E9AB" box="[540,669,1682,1708]" italics="true" pageId="40" pageNumber="569">D. obscurus</emphasis>
</taxonomicName>
), mandibles with molar penicil semi-dichotomized (vs simple in all other species), and uropod protopod with distal portion subquadrangular (vs subrectangular in
<taxonomicName id="4C0DD5B092116335FD92F945FD76E80B" authority="Brandt, 1833" box="[629,767,1778,1804]" class="Malacostraca" family="Armadillidae" genus="Armadillo" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B979722192116335FD92F945FD76E80B" box="[629,767,1778,1804]" italics="true" pageId="40" pageNumber="569">D. echinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0DD5B092116335FF63F8A4FE8FE82B" ID-CoL="1a487283-85d7-4168-b944-d30c317b5ab3" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[132,262,1810,1836]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="B979722192116335FF63F8A4FE8FE82B" box="[132,262,1810,1836]" italics="true" pageId="40" pageNumber="569">D. obscurus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0DD5B092116335FEAEF8A4FE4EE82C" ID-CoL="117af273-11c6-441f-ae7e-288891d52caf" authorityName="Cardoso, Bastos-Pereira &amp; Ferreira" authorityYear="2023" box="[329,455,1810,1835]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="B979722192116335FEAEF8A4FE4EE82C" box="[329,455,1810,1835]" italics="true" pageId="40" pageNumber="569">D. spinatus</emphasis>
</taxonomicName>
; triangular in
<taxonomicName id="4C0DD5B092116335FD88F8A4FDCDE84C" authority="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityName="Campos-Filho, Lopez-Orozco &amp; Taiti" authorityYear="2023" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="carrapicho" status="sp. nov.">
<emphasis id="B979722192116335FD88F8A4FC8EE82B" box="[623,775,1810,1836]" italics="true" pageId="40" pageNumber="569">D. carrapicho</emphasis>
Campos-Filho, López-Orozco &amp; Taiti
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A92116335FDB5F884FD1AE84B" box="[594,659,1842,1868]" pageId="40" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
) (see also
<bibRefCitation id="EF9CD3C292116335FF61F8E7FE23E86B" author="CAMPOS-FILHO I. S. &amp; MONTESANTO G. &amp; ARAUJO P. B. &amp; TAITI S." box="[134,426,1873,1900]" pageId="40" pageNumber="569" pagination="2017034" refId="ref34955" refString="CAMPOS-FILHO I. S., MONTESANTO G., ARAUJO P. B. &amp; TAITI S. 2017. - ‰ ew species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Brazil. Iheringia, Serie Zoologia 107: e 2017034. https: // doi. org / 10.1590 / 1678 - 4766 e 2017034" type="journal article" year="2017">
Campos-Filho
<emphasis id="B979722192116335FED7F8E4FEEEE86C" box="[304,359,1873,1899]" italics="true" pageId="40" pageNumber="569">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EF9CD3C292116335FE5EF8E7FD15E86B" author="CARDOSO G. M. &amp; BASTOS-PEREIRA R. &amp; FERREIRA R. L." box="[441,668,1873,1900]" pageId="40" pageNumber="569" pagination="2023008" refId="ref35453" refString="CARDOSO G. M., BASTOS-PEREIRA R. &amp; FERREIRA R. L. 2023. - Cave-dwellers Diploexochus (Isopoda, Armadillidae): new species and new records of the genus from Brazil. Nauplius 31: e 2023008. https: // doi. org / 10.1590 / 2358 - 2936 e 2023008" type="journal article" year="2023">
Cardoso
<emphasis id="B979722192116335FDC6F8E4FDD1E86C" box="[545,600,1873,1899]" italics="true" pageId="40" pageNumber="569">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
<caption id="DF72FEBB921E633AFF63F825FDD0E8EE" ID-DOI="http://doi.org/10.5281/zenodo.10027523" ID-Zenodo-Dep="10027523" httpUri="https://zenodo.org/record/10027523/files/figure.png" pageId="39" pageNumber="568" startId="39.[132,143,1939,1957]" targetBox="[200,1387,215,1888]" targetPageId="39" targetType="figure">
<paragraph id="8BB2AE33921E633AFF63F825FDD0E8EE" blockId="39.[132,1457,1939,2025]" pageId="39" pageNumber="568">
<smallCapsWord id="8D5438EF921E633AFF63F825FF14E8A3" baselines="1952,1953" box="[132,157,1939,1957]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="39" pageNumber="568">FIG</smallCapsWord>
. 27. —
<taxonomicName id="4C0DD5B0921E633AFF39F825FD59E8A2" authority="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityName="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityYear="2023" box="[222,720,1939,1957]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="39" pageNumber="568" phylum="Arthropoda" rank="species" species="exu" status="sp. nov.">
<emphasis id="B9797221921E633AFF39F825FEF9E8A2" box="[222,368,1939,1957]" italics="true" pageId="39" pageNumber="568">Diploexochus exu</emphasis>
Campos-Filho, Sfenthourakis &amp; Bichuette
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A921E633AFD31F825FC81E8A2" box="[726,776,1939,1957]" pageId="39" pageNumber="568" rank="species">n. sp.</taxonomicNameLabel>
, female paratype LES 28003:
<emphasis id="B9797221921E633AFC1AF825FB83E8A2" bold="true" box="[1021,1034,1939,1957]" pageId="39" pageNumber="568">A</emphasis>
, lateral habitus;
<emphasis id="B9797221921E633AFB75F825FB16E8A2" bold="true" box="[1170,1183,1939,1957]" pageId="39" pageNumber="568">B</emphasis>
, tubercles schem;
<emphasis id="B9797221921E633AFADBF825FAC3E8A2" bold="true" box="[1340,1354,1939,1957]" pageId="39" pageNumber="568">C</emphasis>
, scale-seta;
<emphasis id="B9797221921E633AFF63F81CFF1BE8BB" bold="true" box="[132,146,1962,1980]" pageId="39" pageNumber="568">D</emphasis>
, cephalon, back view;
<emphasis id="B9797221921E633AFEB6F81CFED4E8BB" bold="true" box="[337,349,1962,1980]" pageId="39" pageNumber="568">E</emphasis>
, cephalon and pereonites 1 and 2, frontal view;
<emphasis id="B9797221921E633AFD0BF81CFD7EE8BB" bold="true" box="[748,759,1962,1980]" pageId="39" pageNumber="568">F</emphasis>
, pereonite 1-7 epimera, ventral view;
<emphasis id="B9797221921E633AFBC8F81CFBB4E8BB" bold="true" box="[1071,1085,1962,1980]" pageId="39" pageNumber="568">G</emphasis>
, pereonite 1 epimeron, dorsal view;
<emphasis id="B9797221921E633AFA8CF81CFAF0E8BB" bold="true" box="[1387,1401,1962,1980]" pageId="39" pageNumber="568">H</emphasis>
, pereonite 1 epimeron, ventral view;
<emphasis id="B9797221921E633AFE63F876FE03E8D5" bold="true" box="[388,394,1984,2002]" pageId="39" pageNumber="568">I</emphasis>
, pereonite 2 epimeron, dorsal view;
<emphasis id="B9797221921E633AFD53F876FD37E8D5" bold="true" box="[692,702,1984,2002]" pageId="39" pageNumber="568">J</emphasis>
, pereonite 2 epimeron, ventral view;
<emphasis id="B9797221921E633AFC09F876FC72E8D5" bold="true" box="[1006,1019,1984,2002]" pageId="39" pageNumber="568">K</emphasis>
, pereonite 7, pleon, telson and uropods,dorsal view,
<emphasis id="B9797221921E633AFF63F861FF06E8EE" bold="true" box="[132,143,2007,2025]" pageId="39" pageNumber="568">L</emphasis>
, telson;
<emphasis id="B9797221921E633AFF30F861FF61E8EE" bold="true" box="[215,232,2007,2025]" pageId="39" pageNumber="568">M</emphasis>
, antennula;
<emphasis id="B9797221921E633AFEA9F861FED5E8EE" bold="true" box="[334,348,2007,2025]" pageId="39" pageNumber="568">N</emphasis>
, antenna. Scale bar: A, 1 mm.
</paragraph>
</caption>
<caption id="DF72FEBB92116335FF62FABBFEDAEA32" ID-DOI="http://doi.org/10.5281/zenodo.10027525" ID-Zenodo-Dep="10027525" httpUri="https://zenodo.org/record/10027525/files/figure.png" pageId="40" pageNumber="569" startId="40.[133,144,1293,1311]" targetBox="[132,1448,222,1249]" targetPageId="40" targetType="figure">
<paragraph id="8BB2AE3392116335FF62FABBFEDAEA32" blockId="40.[133,1457,1293,1333]" pageId="40" pageNumber="569">
<smallCapsWord id="8D5438EF92116335FF62FABBFF14EA19" baselines="1306,1307" box="[133,157,1293,1311]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="40" pageNumber="569">FIG</smallCapsWord>
. 28. —
<taxonomicName id="4C0DD5B092116335FF05FABBFD57EA18" authority="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityName="Campos-Filho, Sfenthourakis &amp; Bichuette" authorityYear="2023" box="[226,734,1293,1311]" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" pageId="40" pageNumber="569" phylum="Arthropoda" rank="species" species="exu" status="sp. nov.">
<emphasis id="B979722192116335FF05FABBFEFFEA18" box="[226,374,1293,1311]" italics="true" pageId="40" pageNumber="569">Diploexochus exu</emphasis>
Campos-Filho, Sfenthourakis &amp; Bichuette
</taxonomicName>
,
<taxonomicNameLabel id="A24ACF5A92116335FD01FABBFC93EA18" box="[742,794,1293,1311]" pageId="40" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
, female paratype LES 28003:
<emphasis id="B979722192116335FBFEFABBFBAFEA18" bold="true" box="[1049,1062,1293,1311]" pageId="40" pageNumber="569">A</emphasis>
, left mandible;
<emphasis id="B979722192116335FB4DFABBFB3EEA18" bold="true" box="[1194,1207,1293,1311]" pageId="40" pageNumber="569">B</emphasis>
, right mandible;
<emphasis id="B979722192116335FAAFFABBFADFEA18" bold="true" box="[1352,1366,1293,1311]" pageId="40" pageNumber="569">C</emphasis>
, maxillula;
<emphasis id="B979722192116335FF62FA95FF1AEA32" bold="true" box="[133,147,1315,1333]" pageId="40" pageNumber="569">D</emphasis>
, maxilla;
<emphasis id="B979722192116335FF06FA95FF64EA32" bold="true" box="[225,237,1315,1333]" pageId="40" pageNumber="569">E</emphasis>
, maxilliped.
</paragraph>
</caption>
<paragraph id="8BB2AE3392116335FF7CF8C7FDCAE8EB" blockId="40.[130,777,1427,2028]" pageId="40" pageNumber="569">
This species is considered to be a troglophile due to the absence of troglomorphic characteristics. It probably inhabits caves to exploit their resources and favourable micro-habitat conditions (
<bibRefCitation id="EF9CD3C292116335FEE1F867FE64E8EC" author="FERNANDES C. S. &amp; BATALHA M. A. &amp; BICHUETTE M. E." box="[262,493,2001,2027]" pageId="40" pageNumber="569" pagination="0151958" refId="ref36014" refString="FERNANDES C. S., BATALHA M. A. &amp; BICHUETTE M. E. 2016. - Does the cave environment reduce functional diversity? PLoS ONE 11 (3): e 0151958. https: // doi. org / 10.1371 / journal. pone. 0151958" type="journal article" year="2016">
Fernandes
<emphasis id="B979722192116335FE9CF864FE27E8EC" box="[379,430,2001,2027]" italics="true" pageId="40" pageNumber="569">et al.</emphasis>
2016
</bibRefCitation>
, 2019).
</paragraph>
</subSubSection>
</treatment>
</document>