treatments-xml/data/03/A2/87/03A287FCE7072920FF4CF883FAFFFCE1.xml

<document id="00BFBA4B2FA82BCDAF95AFF0437F849E" ID-CLB-Dataset="78963" ID-DOI="10.11646/phytotaxa.552.1.5" ID-GBIF-Dataset="df7b3dba-392e-4bcc-ad10-f676cf336a4f" ID-ISSN="1179-3163" ID-Zenodo-Dep="6673237" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655792421783" checkinUser="plazi" docAuthor="Damián, Alexander, Garzón, Henry X., Baquero, Luis, Jiménez, Marco M., Vélez-Abarca, Leisberth, Iturralde, Gabriel A., Mitidieri, Nicole, Olortegui, Sergio &amp; Cameron, Kenneth M." docDate="2022" docId="03A287FCE7072920FF4CF883FAFFFCE1" docLanguage="en" docName="phytotaxa.552.1.5.pdf" docOrigin="Phytotaxa 552 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Vanilla andina Damian &amp; H. Garzon 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="70" masterDocId="FF9BFF84E7062927FFC4FFD2FFE7FFA1" masterDocTitle="Vanilla andina (Vanilloideae, Orchidaceae), a new species of the membranaceousleaved group from Peru and Ecuador" masterLastPageNumber="72" masterPageNumber="63" pageNumber="64" updateTime="1699349106699" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6D210853F2545C5B9E91D75A46DEC3E0">Vanilla andina (Vanilloideae, Orchidaceae), a new species of the membranaceousleaved group from Peru and Ecuador</mods:title>
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<mods:namePart id="E796BAA2FAE15A20C1FB75704E25B894">Damián, Alexander</mods:namePart>
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<mods:affiliation id="21230EEC13A17B89BCCC33D9B88DA529">Grupo Científico Calaway Dodson: Investigación y Conservación de Orquídeas del Ecuador, Quito, 170510, Pichincha, Ecuador &amp; Herbario Azuay, Facultad de Ciencia y Tecnología, Universidad del Azuay, Av. 24 de Mayo 7 - 77 y Hernán Malo, Cuenca, Ecuador &amp; hg _ palaco 14 @ hotmail. com; https: // orcid. org / 0000 - 0003 - 4049 - 1652</mods:affiliation>
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<taxonomicName id="4C0B4D69E7072926FF4CF883FE0BF8C6" authority="Damian &amp; H. Garzon" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[136,492,1873,1895]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="andina" status="sp. nov.">
<emphasis id="B97FEAF8E7072926FF4CF883FEF2F8C7" box="[136,277,1873,1894]" italics="true" pageId="1" pageNumber="64">Vanilla andina</emphasis>
Damián &amp; H. Garzón
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<figureCitation id="13302A6FE7072926FD8EF883FD27F8C6" box="[586,704,1873,1895]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[136,229,1913,1935]" captionStartId-1="3.[136,229,1989,2011]" captionStartId-2="4.[136,229,1989,2011]" captionStartId-3="5.[136,229,764,786]" captionTargetBox-0="[170,1416,278,1885]" captionTargetBox-1="[151,1436,195,1965]" captionTargetBox-2="[175,1426,194,1958]" captionTargetBox-3="[151,1436,190,740]" captionTargetId-0="figure-75@2.[170,1416,278,1885]" captionTargetId-1="figure-19@3.[151,1436,190,1965]" captionTargetId-2="figure-26@4.[151,1436,190,1965]" captionTargetId-3="figure-19@5.[151,1436,190,740]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Lankester Composite Dissection Plate (LCDP) of Vanilla andina. A. Habit. B. Flower. C. Dissected perianth. D. Lip and column (left) and in longitudinal section (right). E. Column, ventral (left), lateral (center) and dorsal (right) view. F. Anther cap in dorsal. (above left), ventral (above right), lateral (below, left) view, with pollinarium (below right). Photographs by Henry X. Garzón based on M. Jiménez &amp; H. Garzón 1274." captionText-1="FIGURE 2. Line drawing of Vanilla andina based on the Peruvian holotype. A. Habit, B. Inflorescence, C. Flower in lateral view, D. Dissected perianth. E. Lip and column, lateral view. F. Column, ventral (right) and lateral view (left). G. Anther cap in dorsal (left) and ventral (right) view. Drawing by N. Mitidieri." captionText-2="FIGURE 3. Line drawing of Vanilla andina based on the Ecuadorian specimen. A. Habit, B. Flower, C. Dissected perianth. D. Lip and column, lateral view. E. Column, ventral and lateral view. Drawing by Leisberth Velez-Abarca." captionText-3="FIGURE 4. Lip distended comparison of A. Vanilla oroana, B. V. armoriquensis, C. V. andina, D. V. costaricensis. Photographs by (A) F. Tobar, (B) A. Damian, (C) H. Garzón, (D) A. Karremans. Scale bar: 2 cm" figureDoi-0="http://doi.org/10.5281/zenodo.6673239" figureDoi-1="http://doi.org/10.5281/zenodo.6673241" figureDoi-2="http://doi.org/10.5281/zenodo.6673243" figureDoi-3="http://doi.org/10.5281/zenodo.6673245" httpUri-0="https://zenodo.org/record/6673239/files/figure.png" httpUri-1="https://zenodo.org/record/6673241/files/figure.png" httpUri-2="https://zenodo.org/record/6673243/files/figure.png" httpUri-3="https://zenodo.org/record/6673245/files/figure.png" pageId="1" pageNumber="64">Figures 1–4</figureCitation>
).
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</emphasis>
(
<typeStatus id="54B08848E7072926FE1CF84BFDD6F80E" box="[472,561,1945,1967]" pageId="1" pageNumber="64" type="holotype">holotype</typeStatus>
: USM!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3116561E7072925FF4CF805FC61FEA0" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64" type="diagnosis">
<paragraph id="8BB436EAE7072925FF4CF805FC61FEA0" blockId="1.[136,1452,2007,2070]" lastBlockId="2.[136,1452,158,257]" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
The overall flower color and labellum of 
<taxonomicName id="4C0B4D69E7072926FD9EF805FD19F850" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[602,766,2007,2033]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7072926FD9EF805FD19F850" box="[602,766,2007,2033]" italics="true" pageId="1" pageNumber="64">Vanilla andina</emphasis>
</taxonomicName>
are reminiscent of 
<taxonomicName id="4C0B4D69E7072926FC1CF805FB36F850" authorityName="Damian &amp; Mitidieri" authorityYear="2020" box="[984,1233,2007,2033]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="armoriquensis">
<emphasis id="B97FEAF8E7072926FC1CF805FB36F850" box="[984,1233,2007,2033]" italics="true" pageId="1" pageNumber="64">Vanilla armoriquensis</emphasis>
</taxonomicName>
. However, the new species may be easily distinguished by the slightly smaller flower with sepals and petals 4–6.1 × 
<quantity id="4CF39B0FE7072926FB7EF829FAD1F7B4" box="[1210,1334,2043,2069]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="1" pageNumber="64" unit="cm" value="1.3" valueMax="1.7" valueMin="0.9">0.9–1.7 cm</quantity>
(vs. 5–5.5 × 
<quantity id="4CF39B0FE7042925FF64FF4CFEE9FF18" box="[160,270,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="2" pageNumber="65" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 cm</quantity>
), oblong to sub-orbicular lip shape (vs. sub-rectangular), longitudinal narrow keels that start at the base of the labellum and extend fully to the middle of the lateral lobes (vs. 1/4) and a broadly obtuse to emarginate crisped labellum mid lobe (vs. triangular acuminate, with involute margins).
</paragraph>
</subSubSection>
<caption id="DF746662E7042925FF4CF8ABFE51F85B" ID-DOI="http://doi.org/10.5281/zenodo.6673239" ID-Zenodo-Dep="6673239" httpUri="https://zenodo.org/record/6673239/files/figure.png" pageId="2" pageNumber="65" startId="2.[136,229,1913,1935]" targetBox="[170,1416,278,1885]" targetPageId="2">
<paragraph id="8BB436EAE7042925FF4CF8ABFE51F85B" blockId="2.[136,1452,1913,2043]" pageId="2" pageNumber="65">
<emphasis id="B97FEAF8E7042925FF4CF8ABFF1EF82F" bold="true" box="[136,249,1913,1935]" pageId="2" pageNumber="65">FIGURE 1</emphasis>
. Lankester Composite Dissection Plate (LCDP) of 
<emphasis id="B97FEAF8E7042925FD37F8ABFC6EF82F" box="[755,905,1913,1934]" italics="true" pageId="2" pageNumber="65">
<taxonomicName id="4C0B4D69E7042925FD37F8ABFC62F82F" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[755,901,1913,1934]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="65" phylum="Tracheophyta" rank="species" species="andina">Vanilla andina</taxonomicName>
.
</emphasis>
<emphasis id="B97FEAF8E7042925FC4BF8ABFC47F82F" bold="true" box="[911,928,1913,1934]" pageId="2" pageNumber="65">A</emphasis>
. Habit. 
<emphasis id="B97FEAF8E7042925FC36F8ABFBEFF82F" bold="true" box="[1010,1032,1913,1934]" pageId="2" pageNumber="65">B.</emphasis>
Flower. 
<emphasis id="B97FEAF8E7042925FBA4F8ABFB90F82E" bold="true" box="[1120,1143,1913,1935]" pageId="2" pageNumber="65">C.</emphasis>
Dissected perianth. 
<emphasis id="B97FEAF8E7042925FA84F8ABFAB6F82F" bold="true" box="[1344,1361,1913,1934]" pageId="2" pageNumber="65">D</emphasis>
. Lip and column (left) and in longitudinal section (right). 
<emphasis id="B97FEAF8E7042925FD9EF84FFD8DF813" bold="true" box="[602,618,1949,1970]" pageId="2" pageNumber="65">E</emphasis>
. Column, ventral (left), lateral (center) and dorsal (right) view. 
<emphasis id="B97FEAF8E7042925FB0FF84FFB39F813" bold="true" box="[1227,1246,1949,1970]" pageId="2" pageNumber="65">F.</emphasis>
Anther cap in dorsal. (above left), ventral (above right), lateral (below, left) view, with pollinarium (below right). Photographs by Henry X. Garzón based on M. Jiménez &amp; H. Garzón 1274.
</paragraph>
</caption>
<caption id="DF746662E7052924FF4CF817FDD8F782" ID-DOI="http://doi.org/10.5281/zenodo.6673241" ID-Zenodo-Dep="6673241" httpUri="https://zenodo.org/record/6673241/files/figure.png" pageId="3" pageNumber="66" startId="3.[136,229,1989,2011]" targetBox="[151,1436,195,1965]" targetPageId="3">
<paragraph id="8BB436EAE7052924FF4CF817FDD8F782" blockId="3.[136,1452,1989,2083]" pageId="3" pageNumber="66">
<emphasis id="B97FEAF8E7052924FF4CF817FF18F87B" bold="true" box="[136,255,1989,2011]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
Line drawing of 
<taxonomicName id="4C0B4D69E7052924FE68F817FDDDF87B" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[428,570,1989,2010]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7052924FE68F817FDDDF87B" box="[428,570,1989,2010]" italics="true" pageId="3" pageNumber="66">Vanilla andina</emphasis>
</taxonomicName>
based on the Peruvian holotype. 
<emphasis id="B97FEAF8E7052924FC47F817FC7CF87B" bold="true" box="[899,923,1989,2010]" pageId="3" pageNumber="66">A.</emphasis>
Habit, 
<emphasis id="B97FEAF8E7052924FC21F817FC1CF87B" bold="true" box="[997,1019,1989,2010]" pageId="3" pageNumber="66">B.</emphasis>
Inflorescence, 
<emphasis id="B97FEAF8E7052924FB55F817FB45F87A" bold="true" box="[1169,1186,1989,2011]" pageId="3" pageNumber="66">C</emphasis>
. Flower in lateral view, D. Dissected perianth. 
<emphasis id="B97FEAF8E7052924FE83F83BFEB0F85F" bold="true" box="[327,343,2025,2046]" pageId="3" pageNumber="66">E</emphasis>
. Lip and column, lateral view. 
<emphasis id="B97FEAF8E7052924FD43F83BFD71F85F" bold="true" box="[647,662,2025,2046]" pageId="3" pageNumber="66">F</emphasis>
. Column, ventral (right) and lateral view (left). 
<emphasis id="B97FEAF8E7052924FBACF83BFB9CF85E" bold="true" box="[1128,1147,2025,2047]" pageId="3" pageNumber="66">G</emphasis>
. Anther cap in dorsal (left) and ventral (right) view. Drawing by N. Mitidieri.
</paragraph>
</caption>
<caption id="DF746662E7022923FF4CF817FBFCF85E" ID-DOI="http://doi.org/10.5281/zenodo.6673243" ID-Zenodo-Dep="6673243" httpUri="https://zenodo.org/record/6673243/files/figure.png" pageId="4" pageNumber="67" startId="4.[136,229,1989,2011]" targetBox="[175,1426,194,1958]" targetPageId="4">
<paragraph id="8BB436EAE7022923FF4CF817FBFCF85E" blockId="4.[136,1452,1989,2047]" pageId="4" pageNumber="67">
<emphasis id="B97FEAF8E7022923FF4CF817FF19F87B" bold="true" box="[136,254,1989,2011]" pageId="4" pageNumber="67">FIGURE 3.</emphasis>
Line drawing of 
<taxonomicName id="4C0B4D69E7022923FE6CF817FDD1F87B" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[424,566,1989,2010]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="67" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7022923FE6CF817FDD1F87B" box="[424,566,1989,2010]" italics="true" pageId="4" pageNumber="67">Vanilla andina</emphasis>
</taxonomicName>
based on the Ecuadorian specimen. 
<emphasis id="B97FEAF8E7022923FC5CF817FC4EF87B" bold="true" box="[920,937,1989,2010]" pageId="4" pageNumber="67">A</emphasis>
. Habit, 
<emphasis id="B97FEAF8E7022923FC3CF817FBE9F87B" bold="true" box="[1016,1038,1989,2010]" pageId="4" pageNumber="67">B.</emphasis>
Flower, 
<emphasis id="B97FEAF8E7022923FBA1F817FB9BF87A" bold="true" box="[1125,1148,1989,2011]" pageId="4" pageNumber="67">C.</emphasis>
Dissected perianth. 
<emphasis id="B97FEAF8E7022923FA86F817FAB4F87B" bold="true" box="[1346,1363,1989,2010]" pageId="4" pageNumber="67">D</emphasis>
. Lip and column, lateral view. 
<emphasis id="B97FEAF8E7022923FE93F83BFE8AF85F" bold="true" box="[343,365,2025,2046]" pageId="4" pageNumber="67">E.</emphasis>
Column, ventral and lateral view. Drawing by Leisberth Velez-Abarca.
</paragraph>
</caption>
<caption id="DF746662E7032922FF4CFD2EFCAFFC97" ID-DOI="http://doi.org/10.5281/zenodo.6673245" ID-Zenodo-Dep="6673245" httpUri="https://zenodo.org/record/6673245/files/figure.png" pageId="5" pageNumber="68" startId="5.[136,229,764,786]" targetBox="[151,1436,190,740]" targetPageId="5">
<paragraph id="8BB436EAE7032922FF4CFD2EFCAFFC97" blockId="5.[136,1452,764,823]" pageId="5" pageNumber="68">
<emphasis id="B97FEAF8E7032922FF4CFD2EFF1AFCB3" bold="true" box="[136,253,764,786]" pageId="5" pageNumber="68">FIGURE 4.</emphasis>
Lip distended comparison of A. 
<taxonomicName id="4C0B4D69E7032922FDF3FD2FFD22FCB3" box="[567,709,765,787]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="oroana">Vanilla oroana</taxonomicName>
, B. 
<taxonomicName id="4C0B4D69E7032922FD2EFD2FFC68FCB2" authorityName="Damian &amp; Mitidieri" authorityYear="2020" box="[746,911,765,787]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="armoriquensis">V. armoriquensis</taxonomicName>
, C. 
<taxonomicName id="4C0B4D69E7032922FC77FD2FFBF7FCB2" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[947,1040,765,787]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="andina">V. andina</taxonomicName>
, D. 
<taxonomicName id="4C0B4D69E7032922FBF2FD2FFB29FCB2" box="[1078,1230,765,787]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="costaricensis">V. costaricensis</taxonomicName>
. Photographs by (A) F. Tobar, (B) A. Damian, (C) H. Garzón, (D) A. Karremans. Scale bar: 2 cm
</paragraph>
</caption>
<subSubSection id="C3116561E7032922FF79FCB7FEE7F832" pageId="5" pageNumber="68" type="description">
<paragraph id="8BB436EAE7032922FF79FCB7FEE7F832" blockId="5.[136,1452,869,2084]" pageId="5" pageNumber="68">
<emphasis id="B97FEAF8E7032922FF79FCB7FE8AFCDE" bold="true" box="[189,365,869,895]" pageId="5" pageNumber="68">Description:—</emphasis>
Hemiepiphytic 
<emphasis id="B97FEAF8E7032922FDDFFCB7FDA9FCDE" box="[539,590,869,895]" italics="true" pageId="5" pageNumber="68">herb</emphasis>
, leafy, that branches out as it grows up to 
<quantity id="4CF39B0FE7032922FBE8FCB7FB82FC21" box="[1068,1125,869,896]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="68" unit="m" value="12.0">12 m</quantity>
long. Stems flexuose, terete, dark green, usually verruculose, 
<quantity id="4CF39B0FE7032922FE38FC5BFD71FC05" box="[508,662,905,932]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="7.7" metricValueMin="4.0" pageId="5" pageNumber="68" unit="cm" value="0.585" valueMax="0.77" valueMin="0.4">0.40–0.77 cm</quantity>
wide; internodes 
<quantity id="4CF39B0FE7032922FCA7FC5BFC22FC05" box="[867,965,905,932]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="16.0" metricValueMin="3.0" pageId="5" pageNumber="68" unit="cm" value="9.5" valueMax="16.0" valueMin="3.0">3–16 cm</quantity>
long. Aerial 
<emphasis id="B97FEAF8E7032922FB98FC58FB72FC02" box="[1116,1173,906,931]" italics="true" pageId="5" pageNumber="68">roots</emphasis>
, terete, whitish, 1.5–9 × 
<quantity id="4CF39B0FE7032922FF4CFC7FFEF0FC69" box="[136,279,941,968]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.5999999999999996" metricValueMin="2.0" pageId="5" pageNumber="68" unit="cm" value="0.28" valueMax="0.36" valueMin="0.2">0.2–0.36 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FEE5FC7CFE88FC66" box="[289,367,942,967]" italics="true" pageId="5" pageNumber="68">Leaves</emphasis>
green to dark green, sessile, shiny adaxially and abaxially; blade elliptic, lanceolate when young and orbicular close to the flowering buds, base attenuate to rounded, apex acuminate to caudate, membranaceous, turning dark brown when senescent, becoming blackish after herborization, margins wavy when fresh, mid-vein sulcate, veins conspicuously reticulate, 9.5–18 × 
<quantity id="4CF39B0FE7032922FD98FBCBFD2DFB95" box="[604,714,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="5" pageNumber="68" unit="cm" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FD12FBCBFC8BFB92" box="[726,876,1049,1075]" italics="true" pageId="5" pageNumber="68">Inflorescence</emphasis>
axillary, racemose, rarely paniculate, successively 3–5 flowered; peduncle 
<quantity id="4CF39B0FE7032922FE53FBEFFDF3FBF9" box="[407,532,1085,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.8" metricValueMin="0.6" pageId="5" pageNumber="68" unit="cm" value="1.7" valueMax="2.8" valueMin="0.6">0.6–2.8 cm</quantity>
long, rachis 
<quantity id="4CF39B0FE7032922FD61FBEFFCE2FBF9" box="[677,773,1085,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.9" metricValueMin="0.7" pageId="5" pageNumber="68" unit="cm" value="13.0" valueMax="19.0" valueMin="7.0">7–19 cm</quantity>
long; bracts at the base of the inflorescence green, generally small secondary branches, 2–2.5 × 
<quantity id="4CF39B0FE7032922FDCBFBB3FD9DFBDD" box="[527,634,1121,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="68" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FD40FBB3FCFFFBDA" box="[644,792,1121,1147]" italics="true" pageId="5" pageNumber="68">Floral bracts</emphasis>
foliaceous, reticulate, 5.5–8 × 
<quantity id="4CF39B0FE7032922FBABFBB3FB21FBDD" box="[1135,1222,1121,1148]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="68" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FB0BFBB3FACCFBDA" box="[1231,1323,1121,1147]" italics="true" pageId="5" pageNumber="68">Flowers</emphasis>
resupinate, pendular, sepals and petals pale green or dark green, lip white, salmon-colored adaxially at the base, fragrant. 
<emphasis id="B97FEAF8E7032922FAA1FB57FA4CFB3E" box="[1381,1451,1157,1183]" italics="true" pageId="5" pageNumber="68">Ovary</emphasis>
pedicellate, terete, sinuous, curved, something sigmoid, 3–5 × 
<quantity id="4CF39B0FE7032922FC8BFB7BFC7BFB65" box="[847,924,1193,1220]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="68" unit="cm" value="0.4">0.4 cm</quantity>
including the pedicel. 
<emphasis id="B97FEAF8E7032922FB65FB7BFAD5FB62" box="[1185,1330,1193,1219]" italics="true" pageId="5" pageNumber="68">Dorsal sepal</emphasis>
lanceolate to narrowly elliptic; margins undulate, repand, sinuate and curled; base truncate and cuneate, apex acute recurved, revolute, apically twisted, 8–9–veined, 4.4–6.1 × 
<quantity id="4CF39B0FE7032922FD6BFB23FCD7FAAD" box="[687,816,1265,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="68" unit="cm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FCFEFB23FC3FFAAA" box="[826,984,1265,1291]" italics="true" pageId="5" pageNumber="68">Lateral sepals</emphasis>
similar to the dorsal sepal in shape, 4–5.9 × 
<quantity id="4CF39B0FE7032922FF65FAC7FEF7FA91" box="[161,272,1301,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="5" pageNumber="68" unit="cm" value="1.35" valueMax="1.7" valueMin="1.0">1–1.7 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FEE2FAC7FE8BFA8E" box="[294,364,1301,1327]" italics="true" pageId="5" pageNumber="68">Petals</emphasis>
lanceolate to narrowly lanceolate; margins undulate, repand-crisped, recurved at apex, adaxial surface with a central vein canaliculate along, abaxially sulcate along the midvein, 10–12-veined, 4–5.9 × 
<quantity id="4CF39B0FE7032922FAECFAEBFA4FFAF5" box="[1320,1448,1337,1364]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="5" pageNumber="68" unit="cm" value="1.05" valueMax="1.2" valueMin="0.9">0.9–1.2 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FF4CFA8FFF15FAD6" box="[136,242,1373,1399]" italics="true" pageId="5" pageNumber="68">Labellum</emphasis>
white suffused with salmon markings towards the base and at the midline, free, slightly arcuate, ca. 2.6–3 × 
<quantity id="4CF39B0FE7032922FF65FA53FEC3FA3D" box="[161,292,1409,1436]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="5" pageNumber="68" unit="cm" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 cm</quantity>
, three-lobed when extended and forming a tube in its natural position, the margins gradually change from entire to sinuate and crispate at the apex, narrowly channeled abaxially, truncate at the base where attached to the column, with several longitudinal and tall narrow keels conspicuously antrorse-verrucose that extends to the ½ of the lateral lobes; lateral lobes shoulder-shaped, broad, forming a oblong–suborbicular blade with erose to undulate margins; the disc with a two thick, high, ribbed, sigmoid keels, at both sides of the midline with low internal carinae extending to the apex of the midlobe, which collapse towards a central, thick, sub-rectangular, rugose-striated 8–keeled callus that arise from near the middle of the lip and end into a sharp, sub-obtuse point, 3 × 
<quantity id="4CF39B0FE7032922FB4DF98BFB26F9D5" box="[1161,1217,1625,1652]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="68" unit="cm" value="3.0">3 cm</quantity>
long; midlobe white overall, greenish close to the apex and in the middle portion, broadly obtuse to emarginate, middle portion incurved with margins strongly crispate, 1.2 × 
<quantity id="4CF39B0FE7032922FDE8F973FD9BF91D" box="[556,636,1697,1724]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="68" unit="cm" value="1.5">1.5 cm</quantity>
. 
<emphasis id="B97FEAF8E7032922FD43F973FD07F91A" box="[647,736,1697,1723]" italics="true" pageId="5" pageNumber="68">Column</emphasis>
white to yellowish-greenish, ca. 1.6–2.0 × 
<quantity id="4CF39B0FE7032922FB02F973FA85F91D" box="[1222,1378,1697,1724]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="5" pageNumber="68" unit="cm" value="0.185" valueMax="0.2" valueMin="0.17">0.17–0.20 cm</quantity>
, erect, with 4–8 bullate-rugose, orange longitudinal keels, which extend close to the middle at the base of the ventral surface, glabrous, the apex winged with a two-horned clinandrium. 
<emphasis id="B97FEAF8E7032922FCD8F93BFC8DF8A2" box="[796,874,1769,1795]" italics="true" pageId="5" pageNumber="68">Stigma</emphasis>
trilobed, the lateral lobes subquadrate, ca. 0.1 × 
<quantity id="4CF39B0FE7032922FA4CF93BFF4AF889" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="68" unit="cm" value="0.2">0.2 cm</quantity>
, rostellum ca. 0.3 × 
<quantity id="4CF39B0FE7032922FE53F8DFFE11F889" box="[407,502,1805,1832]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="68" unit="cm" value="0.25">0.25 cm</quantity>
, convex, quadrate, the superior margins denticulate and basal ones revolute. 
<emphasis id="B97FEAF8E7032922FA9BF8DFFA4CF886" box="[1375,1451,1805,1831]" italics="true" pageId="5" pageNumber="68">Anther</emphasis>
ca. 0.35 × 
<quantity id="4CF39B0FE7032922FF3BF8E3FEA9F8ED" box="[255,334,1841,1868]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="68" unit="cm" value="0.2">0.2 cm</quantity>
, saddle-shaped, with two horn-shaped projections and with pollen grains not forming a pollinarium but two granular masses. 
<emphasis id="B97FEAF8E7032922FE67F884FE3AF8CE" box="[419,477,1878,1903]" italics="true" pageId="5" pageNumber="68">Fruit</emphasis>
green to dark green, ca. 18–21 × 
<quantity id="4CF39B0FE7032922FC95F887FC37F8D1" box="[849,976,1877,1904]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="5" pageNumber="68" unit="cm" value="1.05" valueMax="1.3" valueMin="0.8">0.8–1.3 cm</quantity>
, pendular, cylindrical, fleshy, tricarpellate, unilocular.
</paragraph>
</subSubSection>
<subSubSection id="C3116561E7032921FF79F84FFC68FECD" lastPageId="6" lastPageNumber="69" pageId="5" pageNumber="68" type="distribution">
<paragraph id="8BB436EAE7032921FF79F84FFC68FECD" blockId="5.[136,1452,869,2084]" lastBlockId="6.[136,1452,158,365]" lastPageId="6" lastPageNumber="69" pageId="5" pageNumber="68">
<emphasis id="B97FEAF8E7032922FF79F84FFE03F816" bold="true" box="[189,484,1949,1975]" pageId="5" pageNumber="68">Distribution and ecology</emphasis>
:—So far known only from the 
<typeStatus id="54B08848E7032922FC83F84CFC90F819" box="[839,887,1950,1976]" pageId="5" pageNumber="68">type</typeStatus>
locality in the 
<collectingRegion id="49CFF808E7032922FBE4F84FFB7AF816" box="[1056,1181,1949,1975]" country="Peru" name="San Martin" pageId="5" pageNumber="68">San Martin</collectingRegion>
department of 
<collectingCountry id="F31C767AE7032922FA8CF84CFA9CF819" box="[1352,1403,1950,1976]" name="Peru" pageId="5" pageNumber="68">Peru</collectingCountry>
and the 
<collectingRegion id="49CFF808E7032922FF76F813FE07F87A" box="[178,480,1985,2011]" country="Ecuador" name="Morona-Santiago" pageId="5" pageNumber="68">Morona-Santiago province</collectingRegion>
of 
<collectingCountry id="F31C767AE7032922FDCCF813FD82F87A" box="[520,613,1985,2011]" name="Ecuador" pageId="5" pageNumber="68">Ecuador</collectingCountry>
where the species inhabits the pre-montane forest of the eastern slopes of the Andes at ranges of 
<quantity id="4CF39B0FE7032922FE55F837FDCAF7A1" box="[401,557,2021,2048]" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="5" pageNumber="68" unit="m" value="1175.0" valueMax="1250.0" valueMin="1100.0">1100–1250 m</quantity>
in elevation. The Peruvian plant has been seen growing in a coffee plantation surrounded by secondary forest dominated by 
<taxonomicName id="4C0B4D69E7032922FD53F7DBFB4BF782" authority="Cambess. (1828: 320)" authorityName="Cambess." authorityPageNumber="320" authorityYear="1828" box="[663,1196,2057,2083]" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="brasiliense">
<emphasis id="B97FEAF8E7032922FD53F7DBFC4CF782" box="[663,939,2057,2083]" italics="true" pageId="5" pageNumber="68">Calophyllum brasiliense</emphasis>
Cambess. (1828: 320)
</taxonomicName>
and 
<taxonomicName id="4C0B4D69E7032922FB21F7D8FAA5F785" box="[1253,1346,2058,2084]" class="Magnoliopsida" family="Fabaceae" genus="Inga" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="68" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B97FEAF8E7032922FB21F7D8FAFFF782" box="[1253,1304,2058,2083]" italics="true" pageId="5" pageNumber="68">Inga</emphasis>
sp.
</taxonomicName>
, whereas the Ecuadorian specimen has been found climbing trees of 
<taxonomicName id="4C0B4D69E7002921FCF5FF4DFEA6FF7D" authority="(Kunth 1818: 35 - 36) Pruski (1996: 97)" authorityName="Pruski" authorityPageNumber="97" authorityYear="1996" baseAuthorityName="Kunth" baseAuthorityPageNumber="35 - 36" baseAuthorityYear="1818" class="Magnoliopsida" family="Asteraceae" genus="Piptocoma" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="discolor">
<emphasis id="B97FEAF8E7002921FCF5FF4DFBE8FF19" box="[817,1039,158,184]" italics="true" pageId="6" pageNumber="69">Piptocoma discolor</emphasis>
(
<bibRefCitation id="EF9A4B1BE7002921FBDBFF4CFAE7FF18" author="Kunth, K. S." box="[1055,1280,158,185]" pageId="6" pageNumber="69" pagination="301 - 412" refId="ref5030" refString="Kunth, K. S. (1818) Nova Genera et Species Plantarum (folio ed.), vol. 4. N. Maze, Paris, pp. 301 - 412." type="book chapter" year="1818">Kunth 1818: 35–36</bibRefCitation>
) Pruski (1996: 97) (Asteraceae)
</taxonomicName>
in a forest also composed by 
<taxonomicName id="4C0B4D69E7002921FD4AFF10FCC7FF7C" box="[654,800,194,221]" class="Magnoliopsida" family="Fabaceae" genus="Tachigali" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B97FEAF8E7002921FD4AFF10FD10FF7D" box="[654,759,194,220]" italics="true" pageId="6" pageNumber="69">Tachigali</emphasis>
sp.
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7002921FCE8FF11FE9AFEA0" authority="(Willdenow 1806: 1010) Britton &amp; Rose (1928: 40)" authorityName="Britton &amp; Rose" authorityPageNumber="40" authorityYear="1928" baseAuthorityName="Willdenow" baseAuthorityPageNumber="1010" baseAuthorityYear="1806" class="Magnoliopsida" family="Fabaceae" genus="Zygia" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="B97FEAF8E7002921FCE8FF11FC38FF7D" box="[812,991,194,220]" italics="true" pageId="6" pageNumber="69">Zygia longifolia</emphasis>
(
<bibRefCitation id="EF9A4B1BE7002921FC2AFF10FB16FF7D" author="Willdenow, C. L. von" box="[1006,1265,194,220]" pageId="6" pageNumber="69" pagination="633 - 1157" refId="ref5993" refString="Willdenow, C. L. von (1806) Species plantarum, Tomus IV. Pars II. Laurentius Salvius, Stockholm, pp. 633 - 1157." type="book chapter" year="1806">Willdenow 1806: 1010</bibRefCitation>
) 
<bibRefCitation id="EF9A4B1BE7002921FAC4FF10FF1BFEA1" author="Britton, N. L. &amp; Rose, J. N." pageId="6" pageNumber="69" refId="ref4384" refString="Britton, N. L. &amp; Rose, J. N. (1928) North American Flora. Volume 23. Part I. (Rosales). Mimosaceae. The New York Botanical Garden, New York City, 76 pp." type="book" year="1928">Britton &amp; Rose (1928: 40)</bibRefCitation>
(Fabaceae)
</taxonomicName>
and 
<taxonomicName id="4C0B4D69E7002921FE74FF35FC14FEA1" authority="Spruce (1859: 194)" authorityName="Spruce" authorityPageNumber="194" authorityYear="1859" box="[432,1011,230,257]" class="Liliopsida" family="Arecaceae" genus="Wettinia" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="maynensis">
<emphasis id="B97FEAF8E7002921FE74FF35FD60FEA1" box="[432,647,231,256]" italics="true" pageId="6" pageNumber="69">Wettinia maynensis</emphasis>
<bibRefCitation id="EF9A4B1BE7002921FD48FF34FC85FEA1" author="Spruce, R." box="[652,866,230,257]" pageId="6" pageNumber="69" pagination="194 - 196" refId="ref5713" refString="Spruce, R. (1859) On Five New Plants from Eastern Peru. Journal of the Proceedings of the Linnean Society, Botany 3: 194 - 196. https: // doi. org / 10.1111 / j. 1095 - 8339.1859. tb 02054. x" type="journal article" year="1859">Spruce (1859: 194)</bibRefCitation>
(Arecaceae)
</taxonomicName>
. The distance between these two known populations is 
<quantity id="4CF39B0FE7002921FEE9FED8FE61FE85" box="[301,390,266,292]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="69" unit="km" value="400.0">400 km</quantity>
, which is not surprising for the genus; occurrence in nearby areas is highly probable. Flowers and fruits have been seen at the same time (July-September and November-December) in both populations studied. Flowers are slightly aromatic, but no pollinators have been observed.
</paragraph>
</subSubSection>
<caption id="DF746662E7002921FF4CFE4EFEB3FE5D" ID-Table-UUID="DF746662E7002921FF4CFE4EFEB3FE5D" httpUri="http://table.plazi.org/id/DF746662E7002921FF4CFE4EFEB3FE5D" pageId="6" pageNumber="69" startId="6.[136,224,412,436]" targetBox="[144,1441,529,1548]" targetIsTable="true" targetPageId="6">
<paragraph id="8BB436EAE7002921FF4CFE4EFEB3FE5D" blockId="6.[136,1452,412,508]" pageId="6" pageNumber="69">
<emphasis id="B97FEAF8E7002921FF4CFE4EFF1EFE15" bold="true" box="[136,249,412,436]" pageId="6" pageNumber="69">TABLE 1.</emphasis>
Comparison of morphologically similar species compared to 
<taxonomicName id="4C0B4D69E7002921FC44FE4EFBFCFE15" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[896,1051,412,436]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7002921FC44FE4EFBFCFE15" box="[896,1051,412,436]" italics="true" pageId="6" pageNumber="69">Vanilla andina</emphasis>
</taxonomicName>
. References taken from: (1) presented herein, (2) 
<bibRefCitation id="EF9A4B1BE7002921FEC4FE12FDCDFE79" author="Damian, A. &amp; Mitidieri, N." box="[256,554,448,472]" pageId="6" pageNumber="69" pagination="154 - 160" refId="ref4747" refString="Damian, A. &amp; Mitidieri, N. (2020) Living in the clouds: A new high-elevation species of Vanilla (Orchidaceae, Vanilloideae) from Peru. Phytotaxa 451 (2): 154 - 160. https: // doi. org / 10.11646 / phytotaxa. 451.2.5" type="journal article" year="2020">Damian &amp; Mitidieri (2020)</bibRefCitation>
; (3) 
<bibRefCitation id="EF9A4B1BE7002921FD98FE13FCC6FE79" author="Karremans, A. P." box="[604,801,448,472]" pageId="6" pageNumber="69" pagination="749 - 750" refId="ref4921" refString="Karremans, A. P. (2021) Vanilla costaricensis. Orchids, American Orchid Society 90 (10): 749 - 750." type="journal article" year="2021">Karremans (2021)</bibRefCitation>
and 
<bibRefCitation id="EF9A4B1BE7002921FC92FE12FB4EFE79" author="Soto Arenas, M. A. &amp; Dressler, R. L." box="[854,1193,448,472]" pageId="6" pageNumber="69" pagination="285 - 354" refId="ref5615" refString="Soto Arenas, M. A. &amp; Dressler, R. L. (2010) A revision of the Mexican and Central American species of Vanilla Plumier ex Miller with a characterization of their its region of the nuclear ribosomal DNA. Lankesteriana 9: 285 - 354. https: // doi. org / 10.15517 / lank. v 0 i 0.12065" type="journal article" year="2010">Soto Arenas &amp; Dressler (2010)</bibRefCitation>
; (4) 
<bibRefCitation id="EF9A4B1BE7002921FB1FFE12FA99FE79" author="Dodson, C. H." box="[1243,1406,448,472]" pageId="6" pageNumber="69" pagination="191 - 225" refId="ref4800" refString="Dodson, C. H. (2003) Nuevas especies y combinaciones. Orquideologia 22 (3): 191 - 225." type="journal article" year="2003">Dodson (2003)</bibRefCitation>
and Pérez 
<emphasis id="B97FEAF8E7002921FF0EFE37FEE7FE5D" box="[202,256,484,508]" italics="true" pageId="6" pageNumber="69">et al.</emphasis>
(2010).
</paragraph>
</caption>
<paragraph id="8BB436EAE7002921FE47FDC0FAE2F9AA" pageId="6" pageNumber="69">
<table id="F90BC44AE700D6D8FF54FDC3FA46F9AD" box="[144,1441,529,1548]" gridcols="5" gridrows="15" pageId="6" pageNumber="69">
<tr id="353B34A8E700D6D8FF54FDC3FA46FD86" box="[144,1441,529,551]" gridrow="0" pageId="6" pageNumber="69" rowspan-0="1">
<th id="76EA5DD4E700D6D8FE47FDC3FD9CFD86" box="[387,635,529,551]" gridcol="1" gridrow="0" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FE47FDC0FE3AFD87" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[387,477,529,551]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7002921FE47FDC0FE3AFD87" box="[387,477,529,551]" italics="true" pageId="6" pageNumber="69">V. andina</emphasis>
</taxonomicName>
(1)
</th>
<th id="76EA5DD4E700D6D8FD57FDC3FC6BFD86" box="[659,908,529,551]" gridcol="2" gridrow="0" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FD57FDC0FCD2FD86" authorityName="Damian &amp; Mitidieri" authorityYear="2020" box="[659,821,530,551]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="armoriquensis">
<emphasis id="B97FEAF8E7002921FD57FDC0FCD2FD86" box="[659,821,530,551]" italics="true" pageId="6" pageNumber="69">V. armoriquensis</emphasis>
</taxonomicName>
(2)
</th>
<th id="76EA5DD4E700D6D8FC5FFDC3FB77FD86" box="[923,1168,529,551]" gridcol="3" gridrow="0" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FC5FFDC0FBD7FD86" box="[923,1072,530,551]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="costaricensis">
<emphasis id="B97FEAF8E7002921FC5FFDC0FBD7FD86" box="[923,1072,530,551]" italics="true" pageId="6" pageNumber="69">V. costaricensis</emphasis>
</taxonomicName>
(3)
</th>
<th id="76EA5DD4E700D6D8FB6BFDC3FA46FD86" box="[1199,1441,529,551]" gridcol="4" gridrow="0" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FB6BFDC0FAECFD86" box="[1199,1291,530,551]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="oroana">
<emphasis id="B97FEAF8E7002921FB6BFDC0FAECFD86" box="[1199,1291,530,551]" italics="true" pageId="6" pageNumber="69">V. oroana</emphasis>
</taxonomicName>
(4)
</th>
</tr>
<tr id="353B34A8E700D6D8FF54FD93FA46FDF6" box="[144,1441,577,599]" gridrow="1" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FD93FEAFFDF6" box="[144,328,577,599]" gridcol="0" gridrow="1" pageId="6" pageNumber="69">Leaves (cm)</th>
<td id="76EA5DD4E700D6D8FE47FD93FD9CFDF6" box="[387,635,577,599]" gridcol="1" gridrow="1" pageId="6" pageNumber="69">9.5–18 × 6.5–7</td>
<td id="76EA5DD4E700D6D8FD57FD93FC6BFDF6" box="[659,908,577,599]" gridcol="2" gridrow="1" pageId="6" pageNumber="69">12.5–21.5 × 3.7–5.2</td>
<td id="76EA5DD4E700D6D8FC5FFD93FB77FDF6" box="[923,1168,577,599]" gridcol="3" gridrow="1" pageId="6" pageNumber="69">30–34 × 13–17</td>
<td id="76EA5DD4E700D6D8FB6BFD93FA46FDF6" box="[1199,1441,577,599]" gridcol="4" gridrow="1" pageId="6" pageNumber="69">12–30 × 5–15</td>
</tr>
<tr id="353B34A8E700D6D8FF54FDA1FA46FD0C" box="[144,1441,627,685]" gridrow="2" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FDA1FEAFFD0C" box="[144,328,627,685]" gridcol="0" gridrow="2" pageId="6" pageNumber="69">Inflorescence</th>
<td id="76EA5DD4E700D6D8FE47FDA1FD9CFD0C" box="[387,635,627,685]" gridcol="1" gridrow="2" pageId="6" pageNumber="69">Axillary, racemose rarely paniculate</td>
<td id="76EA5DD4E700D6D8FD57FDA1FC6BFD0C" box="[659,908,627,685]" gridcol="2" gridrow="2" pageId="6" pageNumber="69">Axillary, racemose, sometimes paniculate</td>
<td id="76EA5DD4E700D6D8FC5FFDA1FB77FD0C" box="[923,1168,627,685]" gridcol="3" gridrow="2" pageId="6" pageNumber="69">Lateral, racemose</td>
<td id="76EA5DD4E700D6D8FB6BFDA1FA46FD0C" box="[1199,1441,627,685]" gridcol="4" gridrow="2" pageId="6" pageNumber="69">Axillar, racemose</td>
</tr>
<tr id="353B34A8E700D6D8FF54FD1BFA46FD7E" box="[144,1441,713,735]" gridrow="3" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FD1BFEAFFD7E" box="[144,328,713,735]" gridcol="0" gridrow="3" pageId="6" pageNumber="69">Floral bracts (cm)</th>
<td id="76EA5DD4E700D6D8FE47FD1BFD9CFD7E" box="[387,635,713,735]" gridcol="1" gridrow="3" pageId="6" pageNumber="69">5.5–8 × 3–5</td>
<td id="76EA5DD4E700D6D8FD57FD1BFC6BFD7E" box="[659,908,713,735]" gridcol="2" gridrow="3" pageId="6" pageNumber="69">5–11 × 1.5–4.3</td>
<td id="76EA5DD4E700D6D8FC5FFD1BFB77FD7E" box="[923,1168,713,735]" gridcol="3" gridrow="3" pageId="6" pageNumber="69">13–14 × 6–7</td>
<td id="76EA5DD4E700D6D8FB6BFD1BFA46FD7E" box="[1199,1441,713,735]" gridcol="4" gridrow="3" pageId="6" pageNumber="69">6–15 × 3–7.5</td>
</tr>
<tr id="353B34A8E700D6D8FF54FD29FA46FC94" box="[144,1441,763,821]" gridrow="4" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FD29FEAFFC94" box="[144,328,763,821]" gridcol="0" gridrow="4" pageId="6" pageNumber="69">Sepals shape</th>
<td id="76EA5DD4E700D6D8FE47FD29FD9CFC94" box="[387,635,763,821]" gridcol="1" gridrow="4" pageId="6" pageNumber="69">Lanceolate to narrowly elliptic</td>
<td id="76EA5DD4E700D6D8FD57FD29FC6BFC94" box="[659,908,763,821]" gridcol="2" gridrow="4" pageId="6" pageNumber="69">Lanceolate</td>
<td id="76EA5DD4E700D6D8FC5FFD29FB77FC94" box="[923,1168,763,821]" gridcol="3" gridrow="4" pageId="6" pageNumber="69">Narrowly elliptic</td>
<td id="76EA5DD4E700D6D8FB6BFD29FA46FC94" box="[1199,1441,763,821]" gridcol="4" gridrow="4" pageId="6" pageNumber="69">Elliptic</td>
</tr>
<tr id="353B34A8E700D6D8FF54FC83FA46FCC6" box="[144,1441,849,871]" gridrow="5" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FC83FEAFFCC6" box="[144,328,849,871]" gridcol="0" gridrow="5" pageId="6" pageNumber="69">Dorsal sepal (cm)</th>
<td id="76EA5DD4E700D6D8FE47FC83FD9CFCC6" box="[387,635,849,871]" gridcol="1" gridrow="5" pageId="6" pageNumber="69">4.4–6.1 × 1.3–1.5</td>
<td id="76EA5DD4E700D6D8FD57FC83FC6BFCC6" box="[659,908,849,871]" gridcol="2" gridrow="5" pageId="6" pageNumber="69">5.2–5.5 × 1–1.3</td>
<td id="76EA5DD4E700D6D8FC5FFC83FB77FCC6" box="[923,1168,849,871]" gridcol="3" gridrow="5" pageId="6" pageNumber="69">4.3–1.4</td>
<td id="76EA5DD4E700D6D8FB6BFC83FA46FCC6" box="[1199,1441,849,871]" gridcol="4" gridrow="5" pageId="6" pageNumber="69">5 × 1.2</td>
</tr>
<tr id="353B34A8E700D6D8FF54FC51FA46FC38" box="[144,1441,899,921]" gridrow="6" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FC51FEAFFC38" box="[144,328,899,921]" gridcol="0" gridrow="6" pageId="6" pageNumber="69">Lateral sepals (cm)</th>
<td id="76EA5DD4E700D6D8FE47FC51FD9CFC38" box="[387,635,899,921]" gridcol="1" gridrow="6" pageId="6" pageNumber="69">4–5.9 × 1–1.7</td>
<td id="76EA5DD4E700D6D8FD57FC51FC6BFC38" box="[659,908,899,921]" gridcol="2" gridrow="6" pageId="6" pageNumber="69">5.2–5.5 × 1–1.3</td>
<td id="76EA5DD4E700D6D8FC5FFC51FB77FC38" box="[923,1168,899,921]" gridcol="3" gridrow="6" pageId="6" pageNumber="69">3.9–4.1 × 1.4</td>
<td id="76EA5DD4E700D6D8FB6BFC51FA46FC38" box="[1199,1441,899,921]" gridcol="4" gridrow="6" pageId="6" pageNumber="69">5 × 1.2</td>
</tr>
<tr id="353B34A8E700D6D8FF54FC67FA46FC4E" box="[144,1441,949,1007]" gridrow="7" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FC67FEAFFC4E" box="[144,328,949,1007]" gridcol="0" gridrow="7" pageId="6" pageNumber="69">Petals shape</th>
<td id="76EA5DD4E700D6D8FE47FC67FD9CFC4E" box="[387,635,949,1007]" gridcol="1" gridrow="7" pageId="6" pageNumber="69">Lanceolate to narrowly lanceolate</td>
<td id="76EA5DD4E700D6D8FD57FC67FC6BFC4E" box="[659,908,949,1007]" gridcol="2" gridrow="7" pageId="6" pageNumber="69">Lanceolate</td>
<td id="76EA5DD4E700D6D8FC5FFC67FB77FC4E" box="[923,1168,949,1007]" gridcol="3" gridrow="7" pageId="6" pageNumber="69">Narrowly elliptic</td>
<td id="76EA5DD4E700D6D8FB6BFC67FA46FC4E" box="[1199,1441,949,1007]" gridcol="4" gridrow="7" pageId="6" pageNumber="69">Elliptic</td>
</tr>
<tr id="353B34A8E700D6D8FF54FBD8FA46FB80" box="[144,1441,1034,1057]" gridrow="8" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FBD8FEAFFB80" box="[144,328,1034,1057]" gridcol="0" gridrow="8" pageId="6" pageNumber="69">Petals (cm)</th>
<td id="76EA5DD4E700D6D8FE47FBD8FD9CFB80" box="[387,635,1034,1057]" gridcol="1" gridrow="8" pageId="6" pageNumber="69">4–5.9 × 0.9–1.2</td>
<td id="76EA5DD4E700D6D8FD57FBD8FC6BFB80" box="[659,908,1034,1057]" gridcol="2" gridrow="8" pageId="6" pageNumber="69">5 × 1</td>
<td id="76EA5DD4E700D6D8FC5FFBD8FB77FB80" box="[923,1168,1034,1057]" gridcol="3" gridrow="8" pageId="6" pageNumber="69">3.6–3.8 × 1.2</td>
<td id="76EA5DD4E700D6D8FB6BFBD8FA46FB80" box="[1199,1441,1034,1057]" gridcol="4" gridrow="8" pageId="6" pageNumber="69">4.8 × 0.8</td>
</tr>
<tr id="353B34A8E700D6D8FF54FBEFFA46FBD6" box="[144,1441,1085,1143]" gridrow="9" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FBEFFEAFFBD6" box="[144,328,1085,1143]" gridcol="0" gridrow="9" pageId="6" pageNumber="69">Lip overall shape</th>
<td id="76EA5DD4E700D6D8FE47FBEFFD9CFBD6" box="[387,635,1085,1143]" gridcol="1" gridrow="9" pageId="6" pageNumber="69">Oblong-suborbicular, trilobed</td>
<td id="76EA5DD4E700D6D8FD57FBEFFC6BFBD6" box="[659,908,1085,1143]" gridcol="2" gridrow="9" pageId="6" pageNumber="69">Sub-rectangular, trilobed</td>
<td id="76EA5DD4E700D6D8FC5FFBEFFB77FBD6" box="[923,1168,1085,1143]" gridcol="3" gridrow="9" pageId="6" pageNumber="69">Quadrate-suborbicular, incospicously trilobed</td>
<td id="76EA5DD4E700D6D8FB6BFBEFFA46FBD6" box="[1199,1441,1085,1143]" gridcol="4" gridrow="9" pageId="6" pageNumber="69">Trilobed</td>
</tr>
<tr id="353B34A8E700D6D8FF54FB40FA46FB08" box="[144,1441,1170,1193]" gridrow="10" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FB40FEAFFB08" box="[144,328,1170,1193]" gridcol="0" gridrow="10" pageId="6" pageNumber="69">Lip size (cm)</th>
<td id="76EA5DD4E700D6D8FE47FB40FD9CFB08" box="[387,635,1170,1193]" gridcol="1" gridrow="10" pageId="6" pageNumber="69">2.6–3 × 3.7–4.1</td>
<td id="76EA5DD4E700D6D8FD57FB40FC6BFB08" box="[659,908,1170,1193]" gridcol="2" gridrow="10" pageId="6" pageNumber="69">4.5 × 2.3</td>
<td id="76EA5DD4E700D6D8FC5FFB40FB77FB08" box="[923,1168,1170,1193]" gridcol="3" gridrow="10" pageId="6" pageNumber="69">3.1–3.2 × 2.8</td>
<td id="76EA5DD4E700D6D8FB6BFB40FA46FB08" box="[1199,1441,1170,1193]" gridcol="4" gridrow="10" pageId="6" pageNumber="69">5.2 × 3.3</td>
</tr>
<tr id="353B34A8E700D6D8FF54FB16FA46FB5F" box="[144,1441,1220,1278]" gridrow="11" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FB16FEAFFB5F" box="[144,328,1220,1278]" gridcol="0" gridrow="11" pageId="6" pageNumber="69">Lateral lobes</th>
<td id="76EA5DD4E700D6D8FE47FB16FD9CFB5F" box="[387,635,1220,1278]" gridcol="1" gridrow="11" pageId="6" pageNumber="69">Shoulder-shaped, erose to undulate</td>
<td id="76EA5DD4E700D6D8FD57FB16FC6BFB5F" box="[659,908,1220,1278]" gridcol="2" gridrow="11" pageId="6" pageNumber="69">Shoulder-shaped,undulate margins</td>
<td id="76EA5DD4E700D6D8FC5FFB16FB77FB5F" box="[923,1168,1220,1278]" gridcol="3" gridrow="11" pageId="6" pageNumber="69">Truncate</td>
<td id="76EA5DD4E700D6D8FB6BFB16FA46FB5F" box="[1199,1441,1220,1278]" gridcol="4" gridrow="11" pageId="6" pageNumber="69">Cuneate, triangular</td>
</tr>
<tr id="353B34A8E700D6D8FF54FAC8FA46FAF5" box="[144,1441,1306,1364]" gridrow="12" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FAC8FEAFFAF5" box="[144,328,1306,1364]" gridcol="0" gridrow="12" pageId="6" pageNumber="69">Disc</th>
<td id="76EA5DD4E700D6D8FE47FAC8FD9CFAF5" box="[387,635,1306,1364]" gridcol="1" gridrow="12" pageId="6" pageNumber="69">Two thick high, ribbed, sigmoid keels</td>
<td id="76EA5DD4E700D6D8FD57FAC8FC6BFAF5" box="[659,908,1306,1364]" gridcol="2" gridrow="12" pageId="6" pageNumber="69">Two parallel longitudinal keels</td>
<td id="76EA5DD4E700D6D8FC5FFAC8FB77FAF5" box="[923,1168,1306,1364]" gridcol="3" gridrow="12" pageId="6" pageNumber="69">A pair of flat, broad keels along the axial line</td>
<td id="76EA5DD4E700D6D8FB6BFAC8FA46FAF5" box="[1199,1441,1306,1364]" gridcol="4" gridrow="12" pageId="6" pageNumber="69">Two parallel longitudinal keels</td>
</tr>
<tr id="353B34A8E700D6D8FF54FAA4FA46FA75" box="[144,1441,1398,1492]" gridrow="13" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FAA4FEAFFA75" box="[144,328,1398,1492]" gridcol="0" gridrow="13" pageId="6" pageNumber="69">Midlobe</th>
<td id="76EA5DD4E700D6D8FE47FAA4FD9CFA75" box="[387,635,1398,1492]" gridcol="1" gridrow="13" pageId="6" pageNumber="69">Broadly obtuse to emarginate, incurved, margins crispate</td>
<td id="76EA5DD4E700D6D8FD57FAA4FC6BFA75" box="[659,908,1398,1492]" gridcol="2" gridrow="13" pageId="6" pageNumber="69">Triangular-acuminate, incurved with margins involute</td>
<td id="76EA5DD4E700D6D8FC5FFAA4FB77FA75" box="[923,1168,1398,1492]" gridcol="3" gridrow="13" pageId="6" pageNumber="69">Truncate, incurved</td>
<td id="76EA5DD4E700D6D8FB6BFAA4FA46FA75" box="[1199,1441,1398,1492]" gridcol="4" gridrow="13" pageId="6" pageNumber="69">Narrowly obtuse, decurved</td>
</tr>
<tr id="353B34A8E700D6D8FF54FA24FA46F9AD" box="[144,1441,1526,1548]" gridrow="14" pageId="6" pageNumber="69">
<th id="76EA5DD4E700D6D8FF54FA24FEAFF9AD" box="[144,328,1526,1548]" gridcol="0" gridrow="14" pageId="6" pageNumber="69">Column (cm)</th>
<td id="76EA5DD4E700D6D8FE47FA24FD9CF9AD" box="[387,635,1526,1548]" gridcol="1" gridrow="14" pageId="6" pageNumber="69">1.6–2 × 0.17–0.2</td>
<td id="76EA5DD4E700D6D8FD57FA24FC6BF9AD" box="[659,908,1526,1548]" gridcol="2" gridrow="14" pageId="6" pageNumber="69">2 × 0.2</td>
<td id="76EA5DD4E700D6D8FC5FFA24FB77F9AD" box="[923,1168,1526,1548]" gridcol="3" gridrow="14" pageId="6" pageNumber="69">1.6–1.9</td>
<td id="76EA5DD4E700D6D8FB6BFA24FA46F9AD" box="[1199,1441,1526,1548]" gridcol="4" gridrow="14" pageId="6" pageNumber="69">5.4 × 0.8</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3116561E7002921FF79F993FEA4F9DE" pageId="6" pageNumber="69" type="etymology">
<paragraph id="8BB436EAE7002921FF79F993FEA4F9DE" blockId="6.[136,1452,1601,2060]" pageId="6" pageNumber="69">
<emphasis id="B97FEAF8E7002921FF79F993FED9F9FA" bold="true" box="[189,318,1601,1627]" pageId="6" pageNumber="69">Etymology</emphasis>
:—The specific epithet of the new taxon is a reference to the Andean mountain range, where the species is found.
</paragraph>
</subSubSection>
<subSubSection id="C3116561E7002921FF79F95BFADAF966" pageId="6" pageNumber="69" type="materials_examined">
<paragraph id="8BB436EAE7002921FF79F95BFADAF966" blockId="6.[136,1452,1601,2060]" pageId="6" pageNumber="69">
<materialsCitation id="3B633CB7E7002921FF79F95BFADEF966" ID-GBIF-Occurrence="3817021302" collectingDate="2021-08-23" collectorName="M. Jimenez &amp; H. Garzon" country="Ecuador" elevation="1125" latitude="-3.1276944" location="Near San Juan Bosco" longLatPrecision="1" longitude="-78.54253" pageId="6" pageNumber="69" specimenCount="2" stateProvince="Morona-Santiago" typeStatus="paratype">
<emphasis id="B97FEAF8E7002921FF79F95BFD2FF902" bold="true" box="[189,712,1673,1699]" pageId="6" pageNumber="69">
Additional specimens examined (
<typeStatus id="54B08848E7002921FD95F95BFD27F902" box="[593,704,1673,1699]" pageId="6" pageNumber="69" type="paratype">paratype</typeStatus>
)
</emphasis>
:— 
<collectingCountry id="F31C767AE7002921FD2CF95BFC91F902" box="[744,886,1673,1699]" name="Ecuador" pageId="6" pageNumber="69">ECUADOR</collectingCountry>
. 
<collectingRegion id="49CFF808E7002921FC40F958FBAAF902" box="[900,1101,1673,1700]" country="Ecuador" name="Morona-Santiago" pageId="6" pageNumber="69">Morona Santiago</collectingRegion>
: 
<location id="8ED46031E7002921FB98F958FABFF905" LSID="urn:lsid:plazi:treatment:03A287FCE7072920FF4CF883FAFFFCE1:8ED46031E7002921FB98F958FABFF905" box="[1116,1368,1673,1700]" country="Ecuador" latitude="-3.1276944" longLatPrecision="1" longitude="-78.54253" name="Near San Juan Bosco" pageId="6" pageNumber="69" stateProvince="Morona-Santiago">Near San Juan Bosco</location>
, 
<geoCoordinate id="EE3F502DE7002921FAA3F95BFF3BF966" degrees="3" direction="south" minutes="07" orientation="latitude" pageId="6" pageNumber="69" precision="1" seconds="39.7" value="-3.1276944">3º 07’ 39.7” S</geoCoordinate>
<geoCoordinate id="EE3F502DE7002921FF27F97FFE68F969" box="[227,399,1709,1736]" degrees="78" direction="west" minutes="32" orientation="longitude" pageId="6" pageNumber="69" precision="1" seconds="33.1" value="-78.54253">78 32’ 33.1” W</geoCoordinate>
, 
<quantity id="4CF39B0FE7002921FE5EF97FFE17F969" box="[410,496,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="6" pageNumber="69" unit="m" value="1125.0">
<elevation id="0026D1D9E7002921FE5EF97FFE17F969" box="[410,496,1709,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="6" pageNumber="69" unit="m" value="1125.0">1125 m</elevation>
</quantity>
, 
<date id="FFB5102AE7002921FE38F97FFD57F966" box="[508,688,1709,1735]" pageId="6" pageNumber="69" value="2021-08-23">
<collectingDate id="EFF1E9C2E7002921FE38F97FFD57F966" box="[508,688,1709,1735]" pageId="6" pageNumber="69" value="2021-08-23">23 August 2021</collectingDate>
</date>
, 
<emphasis id="B97FEAF8E7002921FD7FF97CFBF3F966" box="[699,1044,1709,1736]" italics="true" pageId="6" pageNumber="69">
<collectorName id="26FE533CE7002921FD7FF97CFCDCF969" box="[699,827,1710,1736]" pageId="6" pageNumber="69">M. Jiménez</collectorName>
&amp; 
<collectorName id="26FE533CE7002921FC9AF97CFC32F969" box="[862,981,1710,1736]" pageId="6" pageNumber="69">H. Garzón</collectorName>
1274
</emphasis>
(QCNE!; HA!; HUTPL!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3116561E7002920FF79F903FAFFFCE1" lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69" type="discussion">
<paragraph id="8BB436EAE7002920FF79F903FE73FEE9" blockId="6.[136,1452,1601,2060]" lastBlockId="7.[136,1452,158,833]" lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69">
<emphasis id="B97FEAF8E7002921FF79F903FE21F94A" bold="true" box="[189,454,1745,1771]" pageId="6" pageNumber="69">Taxonomic Discussion</emphasis>
:— 
<taxonomicName id="4C0B4D69E7002921FE2EF903FD68F94A" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[490,655,1745,1771]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7002921FE2EF903FD68F94A" box="[490,655,1745,1771]" italics="true" pageId="6" pageNumber="69">Vanilla andina</emphasis>
</taxonomicName>
is almost certainly closely related to a small group of morphologically similar species that includes 
<emphasis id="B97FEAF8E7002921FE00F924FD9CF8AE" box="[452,635,1782,1807]" italics="true" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FE00F924FD9FF8AE" box="[452,632,1782,1807]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="amoriquensis">V. amoriquensis</taxonomicName>
,
</emphasis>
<taxonomicName id="4C0B4D69E7002921FD45F927FB9CF8AE" authority="Soto Arenas (2010: 297)" authorityName="Soto Arenas" authorityPageNumber="297" authorityYear="2010" box="[641,1147,1781,1807]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="costaricensis">
<emphasis id="B97FEAF8E7002921FD45F927FC8FF8AE" box="[641,872,1781,1807]" italics="true" pageId="6" pageNumber="69">Vanilla costaricensis</emphasis>
Soto Arenas (2010: 297)
</taxonomicName>
and 
<emphasis id="B97FEAF8E7002921FB6BF924FAF8F8AE" box="[1199,1311,1782,1807]" italics="true" pageId="6" pageNumber="69">
<taxonomicName id="4C0B4D69E7002921FB6BF924FAFBF8AE" box="[1199,1308,1782,1807]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="oroana">V. oroana</taxonomicName>
,
</emphasis>
all featuring foliaceous bracts, a white colored lip with salmon-pink markings, conspicuously tall parallel callus close to or below the middle, and a horned anther. Species members of this group can be easily distinguished mainly by its different lip shape, and midlobe morphology (see 
<tableCitation id="C6890351E7002921FD95F8B3FD43F8DA" box="[593,676,1889,1915]" captionStart="TABLE 1" captionStartId="6.[136,224,412,436]" captionTargetPageId="6" captionText="TABLE 1. Comparison of morphologically similar species compared to Vanilla andina. References taken from: (1) presented herein, (2) Damian &amp; Mitidieri (2020); (3) Karremans (2021) and Soto Arenas &amp; Dressler (2010); (4) Dodson (2003) and Pérez et al. (2010)." httpUri="http://table.plazi.org/id/DF746662E7002921FF4CFE4EFEB3FE5D" pageId="6" pageNumber="69" tableUuid="DF746662E7002921FF4CFE4EFEB3FE5D">Table 1</tableCitation>
, 
<figureCitation id="13302A6FE7002921FD77F8B3FCF3F8DA" box="[691,788,1889,1915]" captionStart="FIGURE 4" captionStartId="5.[136,229,764,786]" captionTargetBox="[151,1436,190,740]" captionTargetId="figure-19@5.[151,1436,190,740]" captionTargetPageId="5" captionText="FIGURE 4. Lip distended comparison of A. Vanilla oroana, B. V. armoriquensis, C. V. andina, D. V. costaricensis. Photographs by (A) F. Tobar, (B) A. Damian, (C) H. Garzón, (D) A. Karremans. Scale bar: 2 cm" figureDoi="http://doi.org/10.5281/zenodo.6673245" httpUri="https://zenodo.org/record/6673245/files/figure.png" pageId="6" pageNumber="69">Figure 4</figureCitation>
). Other membranaceous species differs from 
<taxonomicName id="4C0B4D69E7002921FAD6F8B0FA9BF8DA" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[1298,1404,1889,1915]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7002921FAD6F8B0FA9BF8DA" box="[1298,1404,1889,1915]" italics="true" pageId="6" pageNumber="69">V. andina</emphasis>
</taxonomicName>
and allies by having: a pair of tall broad confluent keels in 
<taxonomicName id="4C0B4D69E7002921FD25F854FB57F83E" authority="Soto Arenas (2010: 342)" authorityName="Soto Arenas" authorityPageNumber="342" authorityYear="2010" box="[737,1200,1925,1951]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="sarapiquensis">
<emphasis id="B97FEAF8E7002921FD25F854FC7FF83E" box="[737,920,1926,1951]" italics="true" pageId="6" pageNumber="69">V. sarapiquensis</emphasis>
Soto Arenas (2010: 342)
</taxonomicName>
; flat disc with truncate mid lobe in 
<taxonomicName id="4C0B4D69E7002921FECBF878FC87F862" authority="Reichenbach &amp; Warszewicz (1854: 97)" authorityName="Reichenbach &amp; Warszewicz" authorityPageNumber="97" authorityYear="1854" box="[271,864,1961,1988]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="methonica">
<emphasis id="B97FEAF8E7002921FECBF878FE78F862" box="[271,415,1961,1987]" italics="true" pageId="6" pageNumber="69">V. methonica</emphasis>
<bibRefCitation id="EF9A4B1BE7002921FE62F87BFC87F862" author="Reichenbach, H. G. &amp; Warszewicz, J." box="[422,864,1961,1988]" pageId="6" pageNumber="69" pagination="97" refId="ref5323" refString="Reichenbach, H. G. &amp; Warszewicz, J. (1854) Orchideae Warscewiczianae recentiores. Bonplandia 2: 97." type="journal article" year="1854">Reichenbach &amp; Warszewicz (1854: 97)</bibRefCitation>
</taxonomicName>
; fleshy disc with 3 thickened keels in 
<taxonomicName id="4C0B4D69E7002921FACDF878FA77F862" box="[1289,1424,1962,1987]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis id="B97FEAF8E7002921FACDF878FA77F862" box="[1289,1424,1962,1987]" italics="true" pageId="6" pageNumber="69">V. mexicana</emphasis>
</taxonomicName>
; a massive cushion callus in 
<taxonomicName id="4C0B4D69E7002921FE69F81CFCE8F849" authority="Schiede (1829: 574)" authorityName="Schiede" authorityPageNumber="574" authorityYear="1829" box="[429,783,1997,2024]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="inodora">
<emphasis id="B97FEAF8E7002921FE69F81CFDC5F846" box="[429,546,1997,2023]" italics="true" pageId="6" pageNumber="69">V. inodora</emphasis>
<bibRefCitation id="EF9A4B1BE7002921FDEDF81FFCE8F849" author="Schiede, C. J." box="[553,783,1997,2024]" pageId="6" pageNumber="69" pagination="554 - 575" refId="ref5396" refString="Schiede, C. J. (1829) Botanische Berichte aus Mexico, mitgetheilt vom Dr. Schiede. Linnaea 4 (4): 554 - 575." type="journal article" year="1829">Schiede (1829: 574)</bibRefCitation>
</taxonomicName>
; broadly hexagonal lip in 
<taxonomicName id="4C0B4D69E7002921FBF0F81CFA40F846" authority="Splitg. (1841: 279)" authorityName="Splitg." authorityPageNumber="279" authorityYear="1841" box="[1076,1447,1997,2023]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="guianensis">
<emphasis id="B97FEAF8E7002921FBF0F81CFB2FF846" box="[1076,1224,1998,2023]" italics="true" pageId="6" pageNumber="69">V. guianensis</emphasis>
Splitg. (1841: 279)
</taxonomicName>
; subtrilobed truncate lip in 
<taxonomicName id="4C0B4D69E7002921FE69F820FC23F7AD" authority="Pansarin &amp; M. R. Miranda (2016: 85)" authorityName="Pansarin &amp; M. R. Miranda" authorityPageNumber="85" authorityYear="2016" box="[429,964,2033,2060]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="arcuata">
<emphasis id="B97FEAF8E7002921FE69F820FDF9F7AA" box="[429,542,2034,2059]" italics="true" pageId="6" pageNumber="69">V. arcuata</emphasis>
Pansarin &amp; M.R. Miranda (2016: 85)
</taxonomicName>
; laminar apical crests of 
<taxonomicName id="4C0B4D69E7002920FB1DF820FF19FF18" authority="Hoehne (1950: 61)" authorityName="Hoehne" authorityPageNumber="61" authorityYear="1950" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" lastPageId="7" lastPageNumber="70" order="Asparagales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="edwallii">
<emphasis id="B97FEAF8E7002921FB1DF820FAA9F7AA" box="[1241,1358,2033,2059]" italics="true" pageId="6" pageNumber="69">V. edwallii</emphasis>
<bibRefCitation id="EF9A4B1BE7002920FA90F823FF19FF18" author="Hoehne, F. C." lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69" pagination="105 - 110" refId="ref4885" refString="Hoehne, F. C. (1950) Algumas novidades da flora do Brasil Austro-Oriental de entre Orchidaceas e Convolvulaceas. Arquivos de Botanica do Estado de Sao Paulo 2: 105 - 110." type="journal article" year="1950">Hoehne (1950: 61)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C0B4D69E7012920FEF7FF4DFD29FF19" authority="Barb. Rodr (1881: 271)" authorityName="Barb. Rodr" authorityPageNumber="271" authorityYear="1881" box="[307,718,158,184]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="B97FEAF8E7012920FEF7FF4DFE5AFF19" box="[307,445,158,184]" italics="true" pageId="7" pageNumber="70">V. parvifolia</emphasis>
Barb. Rodr (1881: 271)
</taxonomicName>
; strongly papillate disc in 
<taxonomicName id="4C0B4D69E7012920FC30FF4DFA90FF19" authority="Schltr. (1922: 19)" authorityName="Schltr." authorityPageNumber="19" authorityYear="1922" box="[1012,1399,158,184]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="angustipetala">
<emphasis id="B97FEAF8E7012920FC30FF4DFB40FF19" box="[1012,1191,158,184]" italics="true" pageId="7" pageNumber="70">V. angustipetala</emphasis>
Schltr. (1922: 19)
</taxonomicName>
; and terrestrial habit in 
<taxonomicName id="4C0B4D69E7012920FE91FF11FD33FF7D" authority="Edwall (1903: 192)" authorityName="Edwall" authorityPageNumber="192" authorityYear="1903" box="[341,724,194,220]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="dietschiana">
<emphasis id="B97FEAF8E7012920FE91FF11FE16FF7D" box="[341,497,194,220]" italics="true" pageId="7" pageNumber="70">V. dietschiana</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FE33FF10FD33FF7D" author="Edwall, G." box="[503,724,194,220]" pageId="7" pageNumber="70" pagination="191 - 195" refId="ref4824" refString="Edwall, G. (1903) Plantas paulistas novas ou menos conhecidas. Revista do Centro de Ciencias, Letras e Artes de Campinas 2: 191 - 195." type="journal article" year="1903">Edwall (1903: 192)</bibRefCitation>
</taxonomicName>
. The rest of membranaceus names [
<taxonomicName id="4C0B4D69E7012920FBA6FF11FA6EFF7C" authority="Rolfe (1896: 453)" authorityName="Rolfe" authorityPageNumber="453" authorityYear="1896" box="[1122,1417,194,221]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B97FEAF8E7012920FBA6FF11FB5CFF7D" box="[1122,1211,195,220]" italics="true" pageId="7" pageNumber="70">V. acuta</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FB06FF10FA6EFF7C" author="Rolfe, R. A." box="[1218,1417,194,221]" pageId="7" pageNumber="70" pagination="439 - 478" refId="ref5346" refString="Rolfe, R. A. (1896) A Revision of the Genus Vanilla. Journal of the Linnean Society of London, Botany 32: 439 - 478. https: // doi. org / 10.1111 / j. 1095 - 8339.1896. tb 00702. x" type="journal article" year="1896">Rolfe (1896: 453)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7012920FA53FF11FE00FEA1" authority="Bertoni (1910: 10)" authorityName="Bertoni" authorityPageNumber="10" authorityYear="1910" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="bertoniensis">
<emphasis id="B97FEAF8E7012920FA53FF11FEF6FEA1" italics="true" pageId="7" pageNumber="70">V. bertoniensis</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FED3FF34FE00FEA1" author="Bertoni, M." box="[279,487,230,256]" pageId="7" pageNumber="70" pagination="10" refId="ref4204" refString="Bertoni, M. (1910) Contribuciones preliminares al estudio sistematico, biologico y economico de las plantas del Paraguay. Anales cientificos paraguayos serie I, 8: 10." type="journal article" year="1910">Bertoni (1910: 10)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7012920FE37FF35FC41FEA1" authority="Schltr. ex Mansf. (1928: 243)" authorityName="Mansf." authorityPageNumber="243" authorityYear="1928" box="[499,934,230,257]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="bradei">
<emphasis id="B97FEAF8E7012920FE37FF35FDB0FEA1" box="[499,599,230,256]" italics="true" pageId="7" pageNumber="70">V. bradei</emphasis>
Schltr. ex Mansf. (1928: 243)
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7012920FC75FF35FAF3FEA0" authority="Rolfe (1896: 452)" authorityName="Rolfe" authorityPageNumber="452" authorityYear="1896" box="[945,1300,230,257]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="organensis">
<emphasis id="B97FEAF8E7012920FC75FF35FBA1FEA1" box="[945,1094,231,256]" italics="true" pageId="7" pageNumber="70">V. organensis</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FB88FF34FAF3FEA0" author="Rolfe, R. A." box="[1100,1300,230,257]" pageId="7" pageNumber="70" pagination="439 - 478" refId="ref5346" refString="Rolfe, R. A. (1896) A Revision of the Genus Vanilla. Journal of the Linnean Society of London, Botany 32: 439 - 478. https: // doi. org / 10.1111 / j. 1095 - 8339.1896. tb 00702. x" type="journal article" year="1896">Rolfe (1896: 452)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7012920FAE5FF35FE91FE84" authority="Hauman (1917: 365)" authorityName="Hauman" authorityPageNumber="365" authorityYear="1917" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="verrucosa">
<emphasis id="B97FEAF8E7012920FAE5FF35FA4CFEA1" box="[1313,1451,231,256]" italics="true" pageId="7" pageNumber="70">V. verrucosa</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FF4CFED9FE91FE84" author="Hauman, L." box="[136,374,266,293]" pageId="7" pageNumber="70" pagination="353 - 389" refId="ref4855" refString="Hauman, L. (1917) Orchidees de largenine. Anales del Museo Nacional de Historia Natural de Buenos Aires 3 (29): 353 - 389." type="journal article" year="1917">Hauman (1917: 365)</bibRefCitation>
</taxonomicName>
] are of dubious definition and might represent synonyms of previous names according to 
<bibRefCitation id="EF9A4B1BE7012920FABCFED8FE77FEE9" author="Soto Arenas, M. A. &amp; Cribb, P." pageId="7" pageNumber="70" pagination="355 - 398" refId="ref5557" refString="Soto Arenas, M. A. &amp; Cribb, P. (2010) A new infrageneric classification and synopsis of the genus Vanilla Plum. ex Mill. (Orchidaceae: Vanillinae). Lankesteriana 3: 355 - 398. https: // doi. org / 10.15517 / lank. v 0 i 0.12071" type="journal article" year="2010">Soto Arenas &amp; Cribb (2010)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB436EAE7012920FF79FE81FD2EFE5D" blockId="7.[136,1452,158,833]" pageId="7" pageNumber="70">
Known populations of 
<taxonomicName id="4C0B4D69E7012920FE7FFE80FDB8FECD" authorityName="Damian &amp; H. Garzon" authorityYear="2022" box="[443,607,338,364]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="andina">
<emphasis id="B97FEAF8E7012920FE7FFE80FDB8FECD" box="[443,607,338,364]" italics="true" pageId="7" pageNumber="70">Vanilla andina</emphasis>
</taxonomicName>
shows some morphological variation, the most conspicuous being the shape of the lip midlobe (
<figureCitation id="13302A6FE7012920FEA2FEA4FE33FE31" box="[358,468,374,400]" captionStart="FIGURE 2" captionStartId="3.[136,229,1989,2011]" captionTargetBox="[151,1436,195,1965]" captionTargetId="figure-19@3.[151,1436,190,1965]" captionTargetPageId="3" captionText="FIGURE 2. Line drawing of Vanilla andina based on the Peruvian holotype. A. Habit, B. Inflorescence, C. Flower in lateral view, D. Dissected perianth. E. Lip and column, lateral view. F. Column, ventral (right) and lateral view (left). G. Anther cap in dorsal (left) and ventral (right) view. Drawing by N. Mitidieri." figureDoi="http://doi.org/10.5281/zenodo.6673241" httpUri="https://zenodo.org/record/6673241/files/figure.png" pageId="7" pageNumber="70">Figures 2</figureCitation>
, 
<figureCitation id="13302A6FE7012920FE26FEA4FE17FE31" box="[482,496,374,400]" captionStart="FIGURE 3" captionStartId="4.[136,229,1989,2011]" captionTargetBox="[175,1426,194,1958]" captionTargetId="figure-26@4.[151,1436,190,1965]" captionTargetPageId="4" captionText="FIGURE 3. Line drawing of Vanilla andina based on the Ecuadorian specimen. A. Habit, B. Flower, C. Dissected perianth. D. Lip and column, lateral view. E. Column, ventral and lateral view. Drawing by Leisberth Velez-Abarca." figureDoi="http://doi.org/10.5281/zenodo.6673243" httpUri="https://zenodo.org/record/6673243/files/figure.png" pageId="7" pageNumber="70">3</figureCitation>
) which in the Peruvian material is broadly obtuse vs. emarginate in the Ecuadorian population. This sort of variation is expected in such a wide range of distribution (&gt; 
<quantity id="4CF39B0FE7012920FB80FE48FB7DFE15" box="[1092,1178,410,436]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="70" unit="km" value="400.0">400 km</quantity>
distance between them) and is coherent with other members of the genus such as 
<taxonomicName id="4C0B4D69E7012920FCC1FE6DFC6CFE79" box="[773,907,447,472]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis id="B97FEAF8E7012920FCC1FE6DFC6CFE79" box="[773,907,447,472]" italics="true" pageId="7" pageNumber="70">V. mexicana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0B4D69E7012920FC5DFE6DFAE1FE78" authority="Schiede (1829: 573)" authorityName="Schiede" authorityPageNumber="573" authorityYear="1829" box="[921,1286,446,473]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="pompona">
<emphasis id="B97FEAF8E7012920FC5DFE6DFBFBFE79" box="[921,1052,447,472]" italics="true" pageId="7" pageNumber="70">V. pompona</emphasis>
<bibRefCitation id="EF9A4B1BE7012920FBE7FE6CFAE1FE78" author="Schiede, C. J." box="[1059,1286,446,473]" pageId="7" pageNumber="70" pagination="554 - 575" refId="ref5396" refString="Schiede, C. J. (1829) Botanische Berichte aus Mexico, mitgetheilt vom Dr. Schiede. Linnaea 4 (4): 554 - 575." type="journal article" year="1829">Schiede (1829: 573)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF9A4B1BE7012920FAD0FE6CFF01FE5D" author="Karremans, A. P. &amp; Chinchilla, I. F. &amp; Rojas-Alvarado, G. &amp; Cedeno-Fonseca, M. &amp; Damian, A. &amp; Leotard, G." pageId="7" pageNumber="70" pagination="395 - 497" refId="ref4947" refString="Karremans, A. P., Chinchilla, I. F., Rojas-Alvarado, G., Cedeno-Fonseca, M., Damian, A. &amp; Leotard, G. (2020) A reappraisal of Neotropical Vanilla. With a note on taxonomic inflation and the importance of alpha taxonomy in biological studies. Lankesteriana 20 (3): 395 - 497. https: // doi. org / 10.15517 / lank. v 20 i 3.45203" type="journal article" year="2020">
Karremans 
<emphasis id="B97FEAF8E7012920FA53FE6DFF42FE5D" italics="true" pageId="7" pageNumber="70">et al.</emphasis>
2020
</bibRefCitation>
) and 
<taxonomicName id="4C0B4D69E7012920FEE0FE31FE68FE5D" box="[292,399,483,508]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="oroana">
<emphasis id="B97FEAF8E7012920FEE0FE31FE68FE5D" box="[292,399,483,508]" italics="true" pageId="7" pageNumber="70">V. oroana</emphasis>
</taxonomicName>
(protologue vs. 
<figureCitation id="13302A6FE7012920FD82FE30FD5AFE5D" box="[582,701,482,508]" captionStart="FIGURE 4" captionStartId="5.[136,229,764,786]" captionTargetBox="[151,1436,190,740]" captionTargetId="figure-19@5.[151,1436,190,740]" captionTargetPageId="5" captionText="FIGURE 4. Lip distended comparison of A. Vanilla oroana, B. V. armoriquensis, C. V. andina, D. V. costaricensis. Photographs by (A) F. Tobar, (B) A. Damian, (C) H. Garzón, (D) A. Karremans. Scale bar: 2 cm" figureDoi="http://doi.org/10.5281/zenodo.6673245" httpUri="https://zenodo.org/record/6673245/files/figure.png" pageId="7" pageNumber="70">Figure 4A</figureCitation>
).
</paragraph>
<paragraph id="8BB436EAE7012920FF79FDD4FAFFFCE1" blockId="7.[136,1452,158,833]" pageId="7" pageNumber="70">
Members of the subgenus 
<taxonomicName id="4C0B4D69E7012920FE1DFDD4FDCFFD81" authorityName="Plumier ex Miller" authorityYear="1754" box="[473,552,518,544]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF8E7012920FE1DFDD4FDCFFD81" box="[473,552,518,544]" italics="true" pageId="7" pageNumber="70">Vanilla</emphasis>
</taxonomicName>
are poorly understood, and details about their pollination biology and evolutionary relationships have not been fully explored. Morphologically they were merged by 
<bibRefCitation id="EF9A4B1BE7012920FBF7FDF8FA90FDE5" author="Soto Arenas, M. A. &amp; Cribb, P." box="[1075,1399,554,580]" pageId="7" pageNumber="70" pagination="355 - 398" refId="ref5557" refString="Soto Arenas, M. A. &amp; Cribb, P. (2010) A new infrageneric classification and synopsis of the genus Vanilla Plum. ex Mill. (Orchidaceae: Vanillinae). Lankesteriana 3: 355 - 398. https: // doi. org / 10.15517 / lank. v 0 i 0.12071" type="journal article" year="2010">Soto Arenas &amp; Cribb (2010)</bibRefCitation>
into to informal groups, the 
<taxonomicName id="4C0B4D69E7012920FE55FD9DFDFFFDC9" box="[401,536,591,616]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis id="B97FEAF8E7012920FE55FD9DFDFFFDC9" box="[401,536,591,616]" italics="true" pageId="7" pageNumber="70">V. mexicana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0B4D69E7012920FD8BFD9DFD3EFDC9" authorityName="Barb. Rodr" authorityYear="1881" box="[591,729,590,616]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="B97FEAF8E7012920FD8BFD9DFD3EFDC9" box="[591,729,590,616]" italics="true" pageId="7" pageNumber="70">V. parvifolia</emphasis>
</taxonomicName>
groups, but the limit of their artificial grouping is controversial. For example, 
<taxonomicName id="4C0B4D69E7012920FEE0FDA1FE34FD2D" authorityName="Soto Arenas" authorityYear="2010" box="[292,467,627,652]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="costaricensis">
<emphasis id="B97FEAF8E7012920FEE0FDA1FE34FD2D" box="[292,467,627,652]" italics="true" pageId="7" pageNumber="70">V. costaricensis</emphasis>
</taxonomicName>
was included in the 
<taxonomicName id="4C0B4D69E7012920FD7AFDA1FCA1FD2D" box="[702,838,627,652]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis id="B97FEAF8E7012920FD7AFDA1FCA1FD2D" box="[702,838,627,652]" italics="true" pageId="7" pageNumber="70">V. mexicana</emphasis>
</taxonomicName>
group due to the lack of a basal triangular keel on the abaxial surface of the column, but at least 
<taxonomicName id="4C0B4D69E7012920FDA0FD45FCC5FD11" authorityName="Damian &amp; Mitidieri" authorityYear="2020" box="[612,802,663,688]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="armoriquensis">
<emphasis id="B97FEAF8E7012920FDA0FD45FCC5FD11" box="[612,802,663,688]" italics="true" pageId="7" pageNumber="70">V. armoriquensis</emphasis>
</taxonomicName>
which is alike to 
<taxonomicName id="4C0B4D69E7012920FC2EFD45FB7FFD11" authorityName="Soto Arenas" authorityYear="2010" box="[1002,1176,663,688]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="costaricensis">
<emphasis id="B97FEAF8E7012920FC2EFD45FB7FFD11" box="[1002,1176,663,688]" italics="true" pageId="7" pageNumber="70">V. costaricensis</emphasis>
</taxonomicName>
, has this keel. The same inconsistency is seen with the non-undulate or involute petals used to distinguish the 
<taxonomicName id="4C0B4D69E7012920FB92FD69FB06FD75" authorityName="Barb. Rodr" authorityYear="1881" box="[1110,1249,698,724]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="B97FEAF8E7012920FB92FD69FB06FD75" box="[1110,1249,698,724]" italics="true" pageId="7" pageNumber="70">V. parvifolia</emphasis>
</taxonomicName>
group, but which are present in 
<taxonomicName id="4C0B4D69E7012920FEECFD0DFE3BFD59" box="[296,476,734,760]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="angustipetala">
<emphasis id="B97FEAF8E7012920FEECFD0DFE3BFD59" box="[296,476,734,760]" italics="true" pageId="7" pageNumber="70">V. angustipetala</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0B4D69E7012920FDD1FD0DFD6CFD59" authorityName="Hoehne" authorityYear="1950" box="[533,651,734,760]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="edwallii">
<emphasis id="B97FEAF8E7012920FDD1FD0DFD6CFD59" box="[533,651,734,760]" italics="true" pageId="7" pageNumber="70">V. edwallii</emphasis>
</taxonomicName>
that are proposed to belong to that group. A more thorough systematic and evolutionary assessment of subgenus 
<taxonomicName id="4C0B4D69E7012920FD98FCD0FD4CFCBD" authorityName="Plumier ex Miller" authorityYear="1754" box="[604,683,770,796]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF8E7012920FD98FCD0FD4CFCBD" box="[604,683,770,796]" italics="true" pageId="7" pageNumber="70">Vanilla</emphasis>
</taxonomicName>
is needed to clarify their taxonomy, but also to bring to light a better understanding of how this ancient 
<taxonomicName id="4C0B4D69E7012920FDCFFCF4FDBDFCE1" authorityName="Plumier ex Miller" authorityYear="1754" box="[523,602,806,832]" class="Liliopsida" family="Orchidaceae" genus="Vanilla" kingdom="Plantae" order="Asparagales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF8E7012920FDCFFCF4FDBDFCE1" box="[523,602,806,832]" italics="true" pageId="7" pageNumber="70">Vanilla</emphasis>
</taxonomicName>
lineage evolved and maintains its place across the Neotropics.
</paragraph>
</subSubSection>
</treatment>
</document>