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<document id="15F8CFDC39C519C2A638B7B72B9964DC" ID-DOI="10.1080/00222933.2017.1360960" ID-ISSN="1464-5262" ID-Zenodo-Dep="5182851" ID-ZooBank="2FA403F6-5535-40A4-9FD0-ADB9913EDE83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628708365672" checkinUser="felipe" docAuthor="Wang, Nannan, Zhang, Yuan, Cheng, Yu &amp; Chen, Hongwei" docDate="2017" docId="03A287F6FFC3FFEB050A8723FEA6FB30" docLanguage="en" docName="JNatHist.51.33-34.1943-1970.pdf" docOrigin="Journal of Natural History 51 (33 - 34)" docSource="http://dx.doi.org/10.1080/00222933.2017.1360960" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Stegana (Oxyphortica) spinosa Cheng and Chen 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1967" masterDocId="FF9BFF8EFFDBFFF2043F8408FFC5FF91" masterDocTitle="Nine new species of the subgenus Stegana (Oxyphortica) from the Oriental region, with morphological and molecular evidence based on Chinese species (Diptera: Drosophilidae)" masterLastPageNumber="1970" masterPageNumber="1943" pageNumber="1966" updateTime="1699050360900" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E16AF48D6E23B3EAD27288ED14988D55">Nine new species of the subgenus Stegana (Oxyphortica) from the Oriental region, with morphological and molecular evidence based on Chinese species (Diptera: Drosophilidae)</mods:title>
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<mods:name id="29D7BD207E68DCEF3BE2E38DD030DBBD" type="personal">
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<mods:namePart id="774FEEAAEF288A12391B89B1B26FBB13">Wang, Nannan</mods:namePart>
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<mods:namePart id="E501A8DD177B951E098F72786F04F8FA">Zhang, Yuan</mods:namePart>
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<mods:namePart id="3454506A00DB4CA1BA6726AAACC32D3E">Cheng, Yu</mods:namePart>
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<mods:namePart id="7EECC67BCC0B8B9E2C7A18C41B942C52">Chen, Hongwei</mods:namePart>
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<mods:title id="D042DA66BAEA0997637EA6B81A997120">Journal of Natural History</mods:title>
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<mods:part id="45AE8B51FC44A5C65F2631B8AE676E95">
<mods:date id="6C7B56B63EF7C02E96DA0041AA192197">2017</mods:date>
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<mods:number id="D4B4CCED5F8AC49DDBB74F4E7EE6D636">2017-09-14</mods:number>
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<mods:number id="9F15FF019CBB2440C7635FD6995FD76C">51</mods:number>
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<mods:detail id="E1CFFEF4AE06A714215DC5431C4A2240" type="issue">
<mods:number id="D58B9C83604DDC68FCA140883F3CCFF9">33 - 34</mods:number>
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<mods:start id="9AE1590192F6C86B29D4251AEC77CAC9">1943</mods:start>
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<mods:identifier id="0563C99620E982C4E98AE659A0BC3883" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03A287F6FFC3FFEB050A8723FEA6FB30" LSID="urn:lsid:plazi:treatment:03A287F6FFC3FFEB050A8723FEA6FB30" httpUri="http://treatment.plazi.org/id/03A287F6FFC3FFEB050A8723FEA6FB30" lastPageId="25" lastPageNumber="1967" pageId="24" pageNumber="1966">
<subSubSection id="C311656BFFC3FFEA050A8723FC2AFCD4" box="[309,1007,811,837]" pageId="24" pageNumber="1966" type="nomenclature">
<paragraph id="8BB436E0FFC3FFEA050A8723FC2AFCD4" blockId="24.[309,1007,811,837]" box="[309,1007,811,837]" pageId="24" pageNumber="1966">
<heading id="D0FC818CFFC3FFEA050A8723FC2AFCD4" box="[309,1007,811,837]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="1966" reason="5">
<taxonomicName id="4C0B4D63FFC3FFEA050A8723FC43FCD4" authority="Cheng and Chen" authorityName="Cheng and Chen" authorityYear="2017" box="[309,902,811,837]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="1966" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Oxyphortica">
<emphasis id="B97FEAF2FFC3FFEA050A8723FD75FCD4" bold="true" box="[309,688,811,837]" pageId="24" pageNumber="1966">
<emphasis id="B97FEAF2FFC3FFEA050A8723FE5CFCD4" bold="true" box="[309,409,811,837]" italics="true" pageId="24" pageNumber="1966">Stegana</emphasis>
(
<emphasis id="B97FEAF2FFC3FFEA05948723FD84FCD4" bold="true" box="[427,577,811,837]" italics="true" pageId="24" pageNumber="1966">Oxyphortica</emphasis>
)
<emphasis id="B97FEAF2FFC3FFEA066C8723FD75FCD4" bold="true" box="[595,688,811,837]" italics="true" pageId="24" pageNumber="1966">spinosa</emphasis>
</emphasis>
Cheng and Chen
</taxonomicName>
,
<taxonomicNameLabel id="A24C5789FFC3FFEA07AD8724FC2AFCD4" box="[914,1007,812,837]" pageId="24" pageNumber="1966" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<caption id="DF746668FFC3FFEA049F8245FCC1F90F" ID-DOI="http://doi.org/10.5281/zenodo.5182887" ID-Zenodo-Dep="5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="24" pageNumber="1966" startId="24.[160,225,1613,1636]" targetBox="[163,1153,965,1582]" targetPageId="24">
<paragraph id="8BB436E0FFC3FFEA049F8245FCC1F90F" blockId="24.[160,1157,1612,1694]" pageId="24" pageNumber="1966">
<emphasis id="B97FEAF2FFC3FFEA049F8245FECDF9F5" bold="true" box="[160,264,1613,1636]" pageId="24" pageNumber="1966">Figure 15.</emphasis>
<taxonomicName id="4C0B4D63FFC3FFEA052F8244FD23F9F5" authority="Cheng and Chen" authorityName="Cheng and Chen" authorityYear="2017" box="[272,742,1612,1636]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="1966" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Oxyphortica">
<emphasis id="B97FEAF2FFC3FFEA052F8244FDF3F9F2" box="[272,566,1612,1635]" italics="true" pageId="24" pageNumber="1966">Stegana (Oxyphortica) spinosa</emphasis>
Cheng and Chen
</taxonomicName>
,
<taxonomicNameLabel id="A24C5789FFC3FFEA06CE8245FC84F9F5" box="[753,833,1613,1636]" pageId="24" pageNumber="1966" rank="species">sp. nov.</taxonomicNameLabel>
, male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C311656BFFC2FFEB049F8486FC4AFEA3" pageId="25" pageNumber="1967" type="diagnosis">
<paragraph id="8BB436E0FFC2FFEB049F8486FED1FF36" blockId="25.[160,1157,142,306]" box="[160,276,142,167]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F8486FED1FF36" bold="true" box="[160,276,142,167]" italics="true" pageId="25" pageNumber="1967">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F84BAFC4AFEA3" blockId="25.[160,1157,142,306]" pageId="25" pageNumber="1967">
Surstylus subbasally with irregular 1 row of ca. 6 somewhat thickened setae along inner margin, distally with two prensisetae (
<figureCitation id="13302A65FFC2FFEB065884DCFD1DFF7D" box="[615,728,212,236]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(b)); aedeagus erect and broadened, acute apically, its medial part sligtly spinulose and with a pair of thorn-like processes, one sheaf of setulae on submedial portion per side (
<figureCitation id="13302A65FFC2FFEB06C48512FCA3FEA3" box="[763,870,282,306]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(c)).
</paragraph>
</subSubSection>
<subSubSection id="C311656BFFC2FFEB049F8569FBB3FCAE" pageId="25" pageNumber="1967" type="description">
<paragraph id="8BB436E0FFC2FFEB049F8569FEE3FEEB" blockId="25.[160,1156,353,620]" box="[160,294,353,378]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F8569FEE3FEEB" bold="true" box="[160,294,353,378]" italics="true" pageId="25" pageNumber="1967">Description</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F858CFC98FDFD" blockId="25.[160,1156,353,620]" pageId="25" pageNumber="1967">
Body colour had been almost lost, because preservation in 70% ethanol for long time. Male terminalia: Epandrium slightly prominent on posterolateral corners, with pubescence and dense setae (
<figureCitation id="13302A65FFC2FFEB059185C2FDDDFE73" box="[430,536,458,482]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(a)). Cercus lacking pubescence (
<figureCitation id="13302A65FFC2FFEB07B985C2FC35FE73" box="[902,1008,458,482]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(a)). Surstylus basally with a few small setae, submedially curved, distally with ca. 12 long setae except prensisetae (
<figureCitation id="13302A65FFC2FFEB05098607FE6DFDB6" box="[310,424,527,551]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(b)). Hypandrium with one pair of large flaps anteromedially (
<figureCitation id="13302A65FFC2FFEB04988639FEDDFDD8" box="[167,280,561,585]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(c)). Parameres slender, fused basally, with ca. 78 sensilla (
<figureCitation id="13302A65FFC2FFEB07D48639FB9EFDD8" box="[1003,1115,561,585]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(c)). Gonopods low-reaching, fused with each other (
<figureCitation id="13302A65FFC2FFEB06F6865CFCF0FDFD" box="[713,821,596,620]" captionStart="Figure 15" captionStartId="24.[160,225,1613,1636]" captionTargetBox="[163,1153,965,1582]" captionTargetId="figure-151@24.[163,1153,965,1582]" captionTargetPageId="24" captionText="Figure 15. Stegana (Oxyphortica) spinosa Cheng and Chen, sp. nov., male terminalia. (a) epandrium, cercus and surstylus (lateral view); (b) surstylus (ventral view); (c) hypandrium, parameres, aedeagus and aedeagal apodeme (ventral view). Scale lines = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5182887" httpUri="https://zenodo.org/record/5182887/files/figure.png" pageId="25" pageNumber="1967">Figure 15</figureCitation>
(c)).
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F8693FE88FD25" blockId="25.[160,1156,667,831]" box="[160,333,667,692]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F8693FE88FD25" bold="true" box="[160,333,667,692]" italics="true" pageId="25" pageNumber="1967">Measurements</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F86B7FBB3FCAE" blockId="25.[160,1156,667,831]" pageId="25" pageNumber="1967">
BL =
<quantity id="4CF39B05FFC2FFEB04E286B7FE84FD46" box="[221,321,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="2.83" pageId="25" pageNumber="1967" unit="mm" value="2.83">2.83 mm</quantity>
in
<typeStatus id="54B08842FFC2FFEB055786B7FE0AFD46" box="[360,463,703,727]" pageId="25" pageNumber="1967" type="holotype">holotype</typeStatus>
, ThL =
<quantity id="4CF39B05FFC2FFEB061986B7FD49FD46" box="[550,652,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="25" pageNumber="1967" unit="mm" value="1.52">1.52 mm</quantity>
, WL =
<quantity id="4CF39B05FFC2FFEB06E186B7FC81FD46" box="[734,836,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="25" pageNumber="1967" unit="mm" value="2.8">2.80 mm</quantity>
; WW =
<quantity id="4CF39B05FFC2FFEB079F86B7FBC3FD46" box="[928,1030,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="25" pageNumber="1967" unit="mm" value="1.18">1.18 mm</quantity>
, arb = 8/5, avd = 0.60, adf = 2.27, flw = 2.00, FW/HW = 0.30, ch/o = 0.09, prorb = 0.74, rcorb = 0.71, vb = 0.29, dcl = 0.48, presctl = 0.48, sctl = 1.02, sterno = 0.83, orbito = 1.50, dcp = 0.29, sctlp = 0.95, C = 3.12, 4c = 1.00, 4v = 2.36, 5x = 1.67, ac = 5.00, M = 0.70, C3F = 0.82.
</paragraph>
</subSubSection>
<subSubSection id="C311656BFFC2FFEB049F8766FB41FC38" pageId="25" pageNumber="1967" type="materials_examined">
<paragraph id="8BB436E0FFC2FFEB049F8766FE95FC16" blockId="25.[160,1156,878,937]" box="[160,336,878,903]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F8766FE95FC16" bold="true" box="[160,336,878,903]" italics="true" pageId="25" pageNumber="1967">Type specimen</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F8799FB41FC38" blockId="25.[160,1156,878,937]" box="[160,1156,913,937]" pageId="25" pageNumber="1967">
<materialsCitation id="3B633CBDFFC2FFEB049F8799FB41FC38" box="[160,1156,913,937]" collectingDate="1999-10-04" collectionCode="KPSP" collectorName="MJ Toda." country="Malaysia" location="Mahua" pageId="25" pageNumber="1967" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54B08842FFC2FFEB049F8799FEC2FC38" box="[160,263,913,937]" pageId="25" pageNumber="1967" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED1AAE25FFC2FFEB056B8799FE56FC38" box="[340,403,913,937]" pageId="25" pageNumber="1967">KPSP</collectionCode>
),
<collectingCountry id="F31C7670FFC2FFEB059B8799FDC3FC38" box="[420,518,913,937]" name="Malaysia" pageId="25" pageNumber="1967">Malaysia</collectingCountry>
:
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,
<collectingRegion id="49CFF802FFC2FFEB06538799FD71FC38" box="[620,692,913,937]" country="Malaysia" name="Sabah" pageId="25" pageNumber="1967">Sabah</collectingRegion>
,
<date id="FFB51020FFC2FFEB06808799FCB7FC38" box="[703,882,913,937]" pageId="25" pageNumber="1967" value="1999-10-04">
<collectingDate id="EFF1E9C8FFC2FFEB06808799FCB7FC38" box="[703,882,913,937]" pageId="25" pageNumber="1967" value="1999-10-04">4 October 1999</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26FE5336FFC2FFEB00218799FB41FC38" box="[1054,1156,913,937]" pageId="25" pageNumber="1967">MJ Toda.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311656BFFC2FFEB049F87D0FEC7FBA7" pageId="25" pageNumber="1967" type="etymology">
<paragraph id="8BB436E0FFC2FFEB049F87D0FED8FC60" blockId="25.[160,1155,984,1078]" box="[160,285,984,1009]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F87D0FED8FC60" bold="true" box="[160,285,984,1009]" italics="true" pageId="25" pageNumber="1967">Etymology</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F87F4FEC7FBA7" blockId="25.[160,1155,984,1078]" pageId="25" pageNumber="1967">
From the Latin word:
<emphasis id="B97FEAF2FFC2FFEB058C87F3FDCAFB85" box="[435,527,1019,1044]" italics="true" pageId="25" pageNumber="1967">spinosus</emphasis>
(thorny), referring to the aedeagus with numerous spinules.
</paragraph>
</subSubSection>
<subSubSection id="C311656BFFC2FFEB049F806DFEA6FB30" box="[160,355,1125,1185]" pageId="25" pageNumber="1967" type="distribution">
<paragraph id="8BB436E0FFC2FFEB049F806DFEE9FBEF" blockId="25.[160,355,1125,1185]" box="[160,300,1125,1150]" pageId="25" pageNumber="1967">
<emphasis id="B97FEAF2FFC2FFEB049F806DFEE9FBEF" bold="true" box="[160,300,1125,1150]" italics="true" pageId="25" pageNumber="1967">Distribution</emphasis>
</paragraph>
<paragraph id="8BB436E0FFC2FFEB049F8081FEA6FB30" blockId="25.[160,355,1125,1185]" box="[160,355,1161,1185]" pageId="25" pageNumber="1967">
<collectingCountry id="F31C7670FFC2FFEB049F8081FEC5FB30" box="[160,256,1161,1185]" name="Malaysia" pageId="25" pageNumber="1967">Malaysia</collectingCountry>
(
<collectingRegion id="49CFF802FFC2FFEB05318081FE9CFB30" box="[270,345,1161,1185]" country="Malaysia" name="Sabah" pageId="25" pageNumber="1967">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>