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<document id="DA14E42E1FB1367239AB63EF85F3AE9B" ID-CLB-Dataset="53472" ID-DOI="10.11646/phytotaxa.585.2.1" ID-GBIF-Dataset="f648d987-0485-4e81-a650-dec255e19aa8" ID-ISSN="1179-3163" ID-Zenodo-Dep="7681555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677506650544" checkinUser="plazi" docAuthor="Kessous, Igor M. &amp; Costa, Andrea F." docDate="2023" docId="03A287F4FFD295429097F8F0FA3C363E" docLanguage="en" docName="phytotaxa.585.2.1.pdf" docOrigin="Phytotaxa 585 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Tillandsia lutheriana Kessous" docType="treatment" docVersion="7" lastPageNumber="85" masterDocId="FF9BFF8CFFD9954C901FFFCAFFBB347B" masterDocTitle="Novelties in “ incertae sedis ” Vriesea (Bromeliaceae: Tillandsioideae): Redescriptions, new combinations and nomenclatural revision" masterLastPageNumber="101" masterPageNumber="71" pageNumber="82" updateTime="1699475854866" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="606D35FA5824B53C2ED4EDD83649F72A">Novelties in “ incertae sedis ” Vriesea (Bromeliaceae: Tillandsioideae): Redescriptions, new combinations and nomenclatural revision</mods:title>
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<mods:namePart id="DADBEBFF018AF7A5A73AFC24D1236C71">Kessous, Igor M.</mods:namePart>
<mods:affiliation id="774040DB6AC31AFA7C05D17A778B0180">Programa de Pós-Graduação em Ciências Biológicas (Botânica), Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart id="0A52C0116B490D521CA853EA9B0F8F58">Costa, Andrea F.</mods:namePart>
<mods:affiliation id="724749F564885A84E4D3B9572304E3E9">Departamento de Botânica, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:title id="8E5B80714BEADF54D3C7EDF76A316666">Phytotaxa</mods:title>
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<treatment id="03A287F4FFD295429097F8F0FA3C363E" ID-DOI="http://doi.org/10.5281/zenodo.7685863" ID-GBIF-Taxon="209347586" ID-Zenodo-Dep="7685863" LSID="urn:lsid:plazi:treatment:03A287F4FFD295429097F8F0FA3C363E" httpUri="http://treatment.plazi.org/id/03A287F4FFD295429097F8F0FA3C363E" lastPageId="14" lastPageNumber="85" pageId="11" pageNumber="82">
<subSubSection id="C3116569FFD295479097F8F0FBB9332F" box="[136,1026,1850,1877]" pageId="11" pageNumber="82" type="nomenclature">
<paragraph id="8BB436E2FFD295479097F8F0FBB9332F" blockId="11.[136,1452,1850,1983]" box="[136,1026,1850,1877]" pageId="11" pageNumber="82">
<heading id="D0FC818EFFD295479097F8F0FBB9332F" box="[136,1026,1850,1877]" fontSize="11" level="2" pageId="11" pageNumber="82" reason="4">
<emphasis id="B97FEAF0FFD295479097F8F0FF10332F" bold="true" box="[136,171,1850,1876]" pageId="11" pageNumber="82">2.2</emphasis>
<taxonomicName id="4C0B4D61FFD2954790ADF8F0FD34332E" authority="(J. R. Grant) Kessous" authorityName="Kessous" baseAuthorityName="J. R. Grant" box="[178,655,1850,1877]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="11" pageNumber="82" phylum="Tracheophyta" rank="species" species="lutheriana">
<emphasis id="B97FEAF0FFD2954790ADF8F0FE1D332E" bold="true" box="[178,422,1850,1877]" italics="true" pageId="11" pageNumber="82">Tillandsia lutheriana</emphasis>
(J.R.Grant) Kessous
</taxonomicName>
&amp; A.F.Costa,
<emphasis id="B97FEAF0FFD295479330F8F0FC13332F" box="[815,936,1850,1876]" italics="true" pageId="11" pageNumber="82">comb. nov.</emphasis>
(
<figureCitation id="13302A67FFD2954793A9F8F0FC41332F" box="[950,1018,1850,1876]" captionStart="FIGURE 6" captionStartId="13.[136,229,1953,1975]" captionTargetBox="[211,1377,198,1922]" captionTargetId="figure-17@13.[203,1385,190,1929]" captionTargetPageId="13" captionText="FIGURE 6. Tillandsia lutheriana (J.R.Grant) Kessous &amp; A.F.Costa. A. Isotype, J.R. Grant 91-01475 (SEL; Courtesy of the Marie Selby Botanical Gardens). B. Habit (Grant 1992: 117;). C. Habit and detail of inflorescence (Grant 1992: 115; Photos: C. Skotak). D. Detail of the flower and the petal (Grant 1992: 116). Images courtesy of the Bromeliad Society International (B, C and D)." figureDoi="http://doi.org/10.5281/zenodo.7681569" httpUri="https://zenodo.org/record/7681569/files/figure.png" pageId="11" pageNumber="82">Fig. 6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFD295479097F8A8FEDA33C4" pageId="11" pageNumber="82" type="reference_group">
<paragraph id="8BB436E2FFD295479097F8A8FEDA33C4" blockId="11.[136,1452,1850,1983]" pageId="11" pageNumber="82">
<emphasis id="B97FEAF0FFD295479097F8A8FF4E3303" bold="true" box="[136,245,1890,1912]" pageId="11" pageNumber="82">Basionym:</emphasis>
<taxonomicName id="4C0B4D61FFD295479112F8A8FDC7330C" ID-CoL="5BRX2" authority="Grant (1992: 114)" authorityName="Grant" authorityPageNumber="114" authorityYear="1992" box="[269,636,1890,1911]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="11" pageNumber="82" phylum="Tracheophyta" rank="species" species="lutheriana">
<emphasis id="B97FEAF0FFD295479112F8A8FE7A330C" box="[269,449,1890,1911]" italics="true" pageId="11" pageNumber="82">Vriesea lutheriana</emphasis>
<materialsCitation id="3B633CBFFFD2954791D6F8A8FDC7330C" ID-GBIF-Occurrence="4039734343" box="[457,636,1890,1911]" country="Costa Rica" location="Puntarenas" pageId="11" pageNumber="82" specimenCount="1" stateProvince="Puntarenas">
<bibRefCitation id="EF9A4B13FFD2954791D6F8A8FDC7330C" author="Grant, J. R." box="[457,636,1890,1911]" pageId="11" pageNumber="82" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant (1992: 114)</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<emphasis id="B97FEAF0FFD295479298F8A8FD7A330C" bold="true" box="[647,705,1890,1911]" pageId="11" pageNumber="82">Type:</emphasis>
<collectingCountry id="F31C7672FFD2954792C6F8A8FCD2330C" box="[729,873,1890,1911]" name="Costa Rica" pageId="11" pageNumber="82">COSTA RICA</collectingCountry>
.
<collectingRegion id="49CFF800FFD29547936BF8A8FC5A330C" box="[884,993,1890,1911]" country="Costa Rica" name="Puntarenas" pageId="11" pageNumber="82">Puntarenas</collectingRegion>
:
<materialsCitation id="3B633CBFFFD2954793F4F8A8FEE633C4" ID-GBIF-Occurrence="4039734374" collectingDate="1991-01-16" collectionCode="CR, SEL, WS" country="United States of America" elevation="1200" location="Lanana" municipality="Angeles" pageId="11" pageNumber="82" specimenCount="1" typeStatus="holotype">
<collectingMunicipality id="6BD0AC98FFD2954793F4F8A8FB873303" box="[1003,1084,1890,1912]" pageId="11" pageNumber="82">Angeles</collectingMunicipality>
,
<locationDeviation id="9A062FC0FFD295479457F8A8FEF233E7" firstLocation="San Vito de Java" pageId="11" pageNumber="82" secondLocation="Agua Buena, Pacific">between San Vito de Java and Agua Buena, Pacific</locationDeviation>
side near
<location id="8ED46039FFD2954791B8F84CFE4B33E0" LSID="urn:lsid:plazi:treatment:03A287F4FFD295429097F8F0FA3C363E:8ED46039FFD2954791B8F84CFE4B33E0" box="[423,496,1926,1947]" country="United States of America" municipality="Angeles" name="Lanana" pageId="11" pageNumber="82">Lanana</location>
, ca.
<quantity id="4CF39B07FFD295479206F84CFDDC33E0" box="[537,615,1926,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="82" unit="m" value="1200.0">
<elevation id="0026D1D1FFD295479206F84CFDDC33E0" box="[537,615,1926,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="82" unit="m" value="1200.0">1,200 m</elevation>
</quantity>
elev.,
<date id="FFB51022FFD2954792BCF84CFCF933E0" box="[675,834,1926,1948]" pageId="11" pageNumber="82" value="1991-01-16">
<collectingDate id="EFF1E9CAFFD2954792BCF84CFCF933E0" box="[675,834,1926,1948]" pageId="11" pageNumber="82" value="1991-01-16">16 January 1991</collectingDate>
</date>
,
<emphasis id="B97FEAF0FFD295479354F84CFC0933E7" box="[843,946,1926,1948]" italics="true" pageId="11" pageNumber="82">Skotak s.n.</emphasis>
preserved as
<emphasis id="B97FEAF0FFD29547942EF84CFB7033E0" box="[1073,1227,1926,1947]" italics="true" pageId="11" pageNumber="82">Grant 91-01475</emphasis>
(
<typeStatus id="54B08840FFD2954794C8F84CFA9633E7" box="[1239,1325,1926,1948]" pageId="11" pageNumber="82" type="holotype">holotype</typeStatus>
<collectionCode id="ED1AAE27FFD29547952DF84CFAEF33E0" box="[1330,1364,1926,1947]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="11" pageNumber="82" type="Herbarium">CR</collectionCode>
, isotypes
<collectionCode id="ED1AAE27FFD2954790A2F860FF5433C4" box="[189,239,1962,1983]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15632" lsid="urn:lsid:biocol.org:col:15632" name="Marie Selby Botanical Gardens" pageId="11" pageNumber="82" type="Herbarium">SEL</collectionCode>
!,
<collectionCode id="ED1AAE27FFD2954790E3F860FE9933C4" box="[252,290,1962,1983]" country="USA" lsid="urn:lsid:biocol.org:col:15672" name="Washington State University" pageId="11" pageNumber="82" type="Herbarium">WS</collectionCode>
, US!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFD295429097F824FB8F365B" lastPageId="14" lastPageNumber="85" pageId="11" pageNumber="82" type="description">
<paragraph id="8BB436E2FFD295479097F824FE403C73" blockId="11.[136,1246,2030,2093]" box="[136,507,2030,2057]" pageId="11" pageNumber="82">
<emphasis id="B97FEAF0FFD295479097F824FEA73C73" bold="true" box="[136,284,2030,2056]" pageId="11" pageNumber="82">Description:</emphasis>
—see
<bibRefCitation id="EF9A4B13FFD29547917CF824FE4C3C73" author="Grant, J. R." box="[355,503,2030,2056]" pageId="11" pageNumber="82" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="8BB436E2FFD2954790A2F7D8FB653C57" blockId="11.[136,1246,2030,2093]" box="[189,1246,2066,2093]" pageId="11" pageNumber="82">
<emphasis id="B97FEAF0FFD2954790A2F7D8FEE23C57" bold="true" box="[189,345,2066,2092]" pageId="11" pageNumber="82">Distribution:</emphasis>
—This species occurs as an epiphyte in
<collectingCountry id="F31C7672FFD295479305F7D8FC2D3C57" box="[794,918,2066,2092]" name="Costa Rica" pageId="11" pageNumber="82">Costa Rica</collectingCountry>
, ca.
<quantity id="4CF39B07FFD2954793D6F7D8FB9E3C56" box="[969,1061,2066,2093]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="82" unit="m" value="1200.0">1,200 m</quantity>
above sea level.
</paragraph>
<caption id="DF74666AFFD595409097F86BFD123384" ID-DOI="http://doi.org/10.5281/zenodo.7681565" ID-Zenodo-Dep="7681565" httpUri="https://zenodo.org/record/7681565/files/figure.png" pageId="12" pageNumber="83" startId="12.[136,229,1953,1975]" targetBox="[182,1399,192,1929]" targetPageId="12">
<paragraph id="8BB436E2FFD595409097F86BFD123384" blockId="12.[136,1452,1953,2047]" pageId="12" pageNumber="83">
<emphasis id="B97FEAF0FFD595409097F86BFF4433CD" bold="true" box="[136,255,1953,1975]" pageId="12" pageNumber="83">FIGURE 5.</emphasis>
<taxonomicName id="4C0B4D61FFD595409119F86BFD9E33CD" authority="L. B. Sm. A." authorityName="L. B. Sm. A." box="[262,549,1953,1974]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="12" pageNumber="83" phylum="Tracheophyta" rank="species" species="duidae">
<emphasis id="B97FEAF0FFD595409119F86BFE1533CD" box="[262,430,1953,1974]" italics="true" pageId="12" pageNumber="83">Tillandsia duidae</emphasis>
L.B.Sm.
<emphasis id="B97FEAF0FFD595409212F86BFD9E33CD" bold="true" box="[525,549,1953,1974]" pageId="12" pageNumber="83">A.</emphasis>
</taxonomicName>
Holotype
<emphasis id="B97FEAF0FFD595409293F868FD6433CD" box="[652,735,1953,1975]" italics="true" pageId="12" pageNumber="83">Tate 437</emphasis>
(NY; Courtesy of the C. V. Starr Virtual Herbarium, New York Botanical Garden, http://sweetgum.nybg.org/science/vh/).
<emphasis id="B97FEAF0FFD595409243F80FFDC933A1" bold="true" box="[604,626,1989,2010]" pageId="12" pageNumber="83">B.</emphasis>
Illustration of the flower, petal, pistil and stamens (Lyman Smiths drawing at GH; Courtesy of the Gray Herbarium of Harvard University).
</paragraph>
</caption>
<caption id="DF74666AFFD495419097F86BFB7D3385" ID-DOI="http://doi.org/10.5281/zenodo.7681569" ID-Zenodo-Dep="7681569" httpUri="https://zenodo.org/record/7681569/files/figure.png" pageId="13" pageNumber="84" startId="13.[136,229,1953,1975]" targetBox="[211,1377,198,1922]" targetPageId="13">
<paragraph id="8BB436E2FFD495419097F86BFB7D3385" blockId="13.[136,1452,1953,2047]" pageId="13" pageNumber="84">
<emphasis id="B97FEAF0FFD495419097F86BFF4533CD" bold="true" box="[136,254,1953,1975]" pageId="13" pageNumber="84">FIGURE 6.</emphasis>
<taxonomicName id="4C0B4D61FFD49541911BF86BFD2D33CD" authority="(J. R. Grant) Kessous" authorityName="Kessous" baseAuthorityName="J. R. Grant" box="[260,662,1953,1974]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="13" pageNumber="84" phylum="Tracheophyta" rank="species" species="lutheriana">
<emphasis id="B97FEAF0FFD49541911BF86BFE7533CD" box="[260,462,1953,1974]" italics="true" pageId="13" pageNumber="84">Tillandsia lutheriana</emphasis>
(J.R.Grant) Kessous
</taxonomicName>
&amp; A.F.Costa.
<emphasis id="B97FEAF0FFD495419300F86BFC8D33CD" bold="true" box="[799,822,1953,1974]" pageId="13" pageNumber="84">A.</emphasis>
Isotype,
<emphasis id="B97FEAF0FFD495419390F868FBEE33CD" box="[911,1109,1953,1975]" italics="true" pageId="13" pageNumber="84">J.R. Grant 91-01475</emphasis>
(SEL; Courtesy of the Marie Selby Botanical Gardens).
<emphasis id="B97FEAF0FFD495419150F80FFEDE33A1" bold="true" box="[335,357,1989,2010]" pageId="13" pageNumber="84">B.</emphasis>
Habit (
<bibRefCitation id="EF9A4B13FFD4954191B2F80FFDEA33A1" author="Grant, J. R." box="[429,593,1989,2010]" pageId="13" pageNumber="84" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant 1992: 117</bibRefCitation>
;).
<emphasis id="B97FEAF0FFD495419278F80FFDC533A0" bold="true" box="[615,638,1989,2011]" pageId="13" pageNumber="84">C.</emphasis>
Habit and detail of inflorescence (
<bibRefCitation id="EF9A4B13FFD4954193D4F80FFBD633A0" author="Grant, J. R." box="[971,1133,1989,2011]" pageId="13" pageNumber="84" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant 1992: 115</bibRefCitation>
; Photos: C. Skotak).
<emphasis id="B97FEAF0FFD495419526F80FFAEB33A1" bold="true" box="[1337,1360,1989,2010]" pageId="13" pageNumber="84">D.</emphasis>
Detail of the flower and the petal (
<bibRefCitation id="EF9A4B13FFD495419168F823FDA23384" author="Grant, J. R." box="[375,537,2025,2047]" pageId="13" pageNumber="84" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant 1992: 116</bibRefCitation>
). Images courtesy of the Bromeliad Society International (B, C and D).
</paragraph>
</caption>
<paragraph id="8BB436E2FFD7954290A2FF54FF6A3517" blockId="14.[136,1452,158,581]" pageId="14" pageNumber="85">
<emphasis id="B97FEAF0FFD7954290A2FF54FD3A34C3" bold="true" box="[189,641,158,184]" pageId="14" pageNumber="85">Taxonomic and nomenclatural notes:</emphasis>
—Despite the ligulate leaf blades (as commonly observed in
<taxonomicName id="4C0B4D61FFD795429557FF55FA2034C3" baseAuthorityName="Smith &amp; Downs" baseAuthorityYear="1977" box="[1352,1435,159,184]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFD795429557FF55FA2034C3" box="[1352,1435,159,184]" italics="true" pageId="14" pageNumber="85">Vriesea</emphasis>
</taxonomicName>
), we observed morphological characters of
<taxonomicName id="4C0B4D61FFD795429272FF08FC6A34A7" box="[621,977,194,220]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="subGenus" subGenus="Pseudovriesea">
<emphasis id="B97FEAF0FFD795429272FF08FD6634A7" box="[621,733,194,220]" italics="true" pageId="14" pageNumber="85">Tillandsia</emphasis>
subg.
<emphasis id="B97FEAF0FFD795429331FF08FC6A34A7" box="[814,977,194,220]" italics="true" pageId="14" pageNumber="85">Pseudovriesea</emphasis>
</taxonomicName>
, such as the presence of a conduplicatespiral stigma and bluish-gray petals (
<bibRefCitation id="EF9A4B13FFD795429223FF2CFD7E357B" author="Grant, J. R." box="[572,709,230,256]" pageId="14" pageNumber="85" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant 1992</bibRefCitation>
). Thus, the set of these traits and the geographical distribution (Central America) lead us to propose this combination in
<taxonomicName id="4C0B4D61FFD795429317FEC0FCC3355F" authorityName="Linnaeus" authorityYear="1753" box="[776,888,266,292]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFD795429317FEC0FCC3355F" box="[776,888,266,292]" italics="true" pageId="14" pageNumber="85">Tillandsia</emphasis>
</taxonomicName>
. This taxon is closely related to
<taxonomicName id="4C0B4D61FFD7954294FEFEC0FA1F355F" box="[1249,1444,266,292]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="species" species="duidae">
<emphasis id="B97FEAF0FFD7954294FEFEC0FA1F355F" box="[1249,1444,266,292]" italics="true" pageId="14" pageNumber="85">Tillandsia duidae</emphasis>
</taxonomicName>
, distinguished mainly by the robustness of its inflorescence and the bluish-gray petals (
<emphasis id="B97FEAF0FFD795429478FEE5FB3D3533" box="[1127,1158,303,328]" italics="true" pageId="14" pageNumber="85">vs.</emphasis>
blue in
<taxonomicName id="4C0B4D61FFD7954294F9FEE5FAEB3533" box="[1254,1360,302,328]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="species" species="duidae">
<emphasis id="B97FEAF0FFD7954294F9FEE5FAEB3533" box="[1254,1360,302,328]" italics="true" pageId="14" pageNumber="85">T. duidae</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EF9A4B13FFD795429575FEE4FF7F3517" author="Grant, J. R." pageId="14" pageNumber="85" pagination="114 - 117" refId="ref21735" refString="Grant, J. R. (1992) Vriesea lutheriana, a new species with tripinnate inflorescence from Costa Rica. Journal of the Bromeliad Society 42: 114 - 117." type="journal article" year="1992">Grant 1992</bibRefCitation>
).
</paragraph>
<paragraph id="8BB436E2FFD7954290A2FEBDFB8F365B" blockId="14.[136,1452,158,581]" pageId="14" pageNumber="85">
In the protologue, the specimen at US is not mentioned; however, we found this specimen in the collection of
<typeStatus id="54B08840FFD795429097FE51FF7835CE" box="[136,195,411,437]" pageId="14" pageNumber="85">types</typeStatus>
in this herbarium.
<bibRefCitation id="EF9A4B13FFD795429189FE50FD3635CF" author="Machado, T. M. &amp; Loiseau, O. &amp; Paris, M. &amp; Weigand, A. &amp; Versieux, L. M. &amp; Stehmann, J. R. &amp; Lexer, C. &amp; Salamin, N." box="[406,653,410,436]" pageId="14" pageNumber="85" refId="ref22298" refString="Machado, T. M., Loiseau, O., Paris, M., Weigand, A., Versieux, L. M., Stehmann, J. R., Lexer, C. &amp; Salamin, N. (2020) Systematics of Vriesea (Bromeliaceae): phylogenetic relationships based on nuclear gene and partial plastome sequences. Botanical Journal of the" type="book" year="2020">
Machado
<emphasis id="B97FEAF0FFD79542921BFE51FD8735CF" box="[516,572,410,436]" italics="true" pageId="14" pageNumber="85">et al.</emphasis>
(2020)
</bibRefCitation>
discussed the morphological similarity between
<taxonomicName id="4C0B4D61FFD7954294B1FE51FAC635CF" authorityName="Grant" authorityYear="1992" box="[1198,1405,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="species" species="lutheriana">
<emphasis id="B97FEAF0FFD7954294B1FE51FAC635CF" box="[1198,1405,410,436]" italics="true" pageId="14" pageNumber="85">Vriesea lutheriana</emphasis>
</taxonomicName>
and species with compound inflorescences of the
<emphasis id="B97FEAF0FFD79542928EFE74FC0E35A3" box="[657,949,446,472]" italics="true" pageId="14" pageNumber="85">Cipuropsis-Mezobromelia</emphasis>
clade (from
<bibRefCitation id="EF9A4B13FFD795429454FE74FA9135A3" author="Barfuss, M. H. J. &amp; Till, W. &amp; Leme, E. M. C. &amp; Pinzon, J. P. &amp; Manzanares, J. M. &amp; Halbritter, H. &amp; Samuel, R. &amp; Brown, G. K." box="[1099,1322,446,473]" pageId="14" pageNumber="85" pagination="1 - 97" refId="ref20630" refString="Barfuss, M. H. J., Till, W., Leme, E. M. C., Pinzon, J. P., Manzanares, J. M., Halbritter, H., Samuel, R. &amp; Brown, G. K. (2016) Taxonomic revision of Bromeliaceae subfam. Tillandsioideae based on a multi-locus DNA sequence phylogeny and morphology. Phytotaxa 279 (1): 1 - 97. https: // doi. org / 10.11646 / phytotaxa. 279.1.1" type="journal article" year="2016">
Barfuss
<emphasis id="B97FEAF0FFD7954294B5FE75FB6635A3" box="[1194,1245,446,472]" italics="true" pageId="14" pageNumber="85">et al</emphasis>
. 2016
</bibRefCitation>
). However, this species was nested in a well-supported
<taxonomicName id="4C0B4D61FFD79542929BFE28FD4F3587" authorityName="Linnaeus" authorityYear="1753" box="[644,756,482,508]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFD79542929BFE28FD4F3587" box="[644,756,482,508]" italics="true" pageId="14" pageNumber="85">Tillandsia</emphasis>
</taxonomicName>
clade by its nuclear gene and partial plastome phylogenetic analysis (
<bibRefCitation id="EF9A4B13FFD7954290EEFDCCFE65365B" author="Machado, T. M. &amp; Loiseau, O. &amp; Paris, M. &amp; Weigand, A. &amp; Versieux, L. M. &amp; Stehmann, J. R. &amp; Lexer, C. &amp; Salamin, N." box="[241,478,518,544]" pageId="14" pageNumber="85" refId="ref22298" refString="Machado, T. M., Loiseau, O., Paris, M., Weigand, A., Versieux, L. M., Stehmann, J. R., Lexer, C. &amp; Salamin, N. (2020) Systematics of Vriesea (Bromeliaceae): phylogenetic relationships based on nuclear gene and partial plastome sequences. Botanical Journal of the" type="book" year="2020">
Machado
<emphasis id="B97FEAF0FFD79542917DFDCDFE21365B" box="[354,410,518,544]" italics="true" pageId="14" pageNumber="85">et al.</emphasis>
2020
</bibRefCitation>
). The species is known only from the
<typeStatus id="54B08840FFD795429399FDCDFC0D365A" box="[902,950,519,545]" pageId="14" pageNumber="85">type</typeStatus>
collection.
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFD7954290A2FDE0FA3C363E" box="[189,1415,554,581]" pageId="14" pageNumber="85" type="etymology">
<paragraph id="8BB436E2FFD7954290A2FDE0FA3C363E" blockId="14.[136,1452,158,581]" box="[189,1415,554,581]" pageId="14" pageNumber="85">
<emphasis id="B97FEAF0FFD7954290A2FDE0FEF3363F" bold="true" box="[189,328,554,580]" pageId="14" pageNumber="85">Etymology:</emphasis>
—The specific epithet “
<taxonomicName id="4C0B4D61FFD79542924BFDE0FD72363F" authorityName="Grant" authorityYear="1992" box="[596,713,554,580]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="14" pageNumber="85" phylum="Tracheophyta" rank="species" species="lutheriana">
<emphasis id="B97FEAF0FFD79542924BFDE0FD72363F" box="[596,713,554,580]" italics="true" pageId="14" pageNumber="85">lutheriana</emphasis>
</taxonomicName>
” refers to the bromeliad expert, Harry E. Luther (19522012).
</paragraph>
</subSubSection>
</treatment>
</document>