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<document id="2531E1B55A4F983C1CC8AD1C382A5C0E" ID-CLB-Dataset="53472" ID-DOI="10.11646/phytotaxa.585.2.1" ID-GBIF-Dataset="f648d987-0485-4e81-a650-dec255e19aa8" ID-ISSN="1179-3163" ID-Zenodo-Dep="7681555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677506650544" checkinUser="plazi" docAuthor="Kessous, Igor M. &amp; Costa, Andrea F." docDate="2023" docId="03A287F4FFCE955B9097FF54FDA93143" docLanguage="en" docName="phytotaxa.585.2.1.pdf" docOrigin="Phytotaxa 585 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Werauhia fibrosa Kessous" docType="treatment" docVersion="8" lastPageNumber="94" masterDocId="FF9BFF8CFFD9954C901FFFCAFFBB347B" masterDocTitle="Novelties in “ incertae sedis ” Vriesea (Bromeliaceae: Tillandsioideae): Redescriptions, new combinations and nomenclatural revision" masterLastPageNumber="101" masterPageNumber="71" pageNumber="94" updateTime="1699475854866" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8975CFADA58FBB5699E7D896A6CD84E4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EB20413392EFAE252D7AF8E3C55AEE1A">
<mods:title id="30DF03943AFF824442DCC7DD088DDD32">Novelties in “ incertae sedis ” Vriesea (Bromeliaceae: Tillandsioideae): Redescriptions, new combinations and nomenclatural revision</mods:title>
</mods:titleInfo>
<mods:name id="81F0738A1F01EB9EA065D48B205B699C" type="personal">
<mods:role id="C4562A36200570ACA46C9C505D23F0DE">
<mods:roleTerm id="77B81C1B21A05588B510B4B78AAAC93D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41A68CB0AFC2B70381A3B8243FA790BC">Kessous, Igor M.</mods:namePart>
<mods:affiliation id="965C32669F900628EEE6F4D904C2A56D">Programa de Pós-Graduação em Ciências Biológicas (Botânica), Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
</mods:name>
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<mods:role id="E5ED134E39460750A38F0411D7FD83FA">
<mods:roleTerm id="9BF313A2F7A4DE0F97BA11C2FEA340A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="472DB1B4B3B04E04FF9ED2AA97756C4D">Costa, Andrea F.</mods:namePart>
<mods:affiliation id="F11ABA251FE50F3A96431CAD19697BA9">Departamento de Botânica, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
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<mods:titleInfo id="8D4F075EF8DA4FEA33DCF1C466C5428E">
<mods:title id="4BB2308894646686AF92746BBC150397">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="43C8826455B25DFB8FAECDEF0BD423AA">
<mods:date id="B9D047F722BFEC1447EBA72C2C4F7840">2023</mods:date>
<mods:detail id="9D23CEED806108C370516FC71A72CE5A" type="pubDate">
<mods:number id="180608BCBB23B38BF46768CE4BC6F9A5">2023-02-27</mods:number>
</mods:detail>
<mods:detail id="B6401D5FD2E20C5F3145999FDF63A4C0" type="volume">
<mods:number id="9B3DA65444A2A92B0566F548B08BF479">585</mods:number>
</mods:detail>
<mods:detail id="D6BFC1A3EE834C5C8F6CC3E431DABEF9" type="issue">
<mods:number id="05658E1931E5842B1B2398AA244A6F3B">2</mods:number>
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<mods:end id="FF33638F8A7DC907ACC1E3A0C32E2119">101</mods:end>
</mods:extent>
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<mods:location id="50CCF05822154D444705B519AF332445">
<mods:url id="F94FD35BAA2146616E8079FD16952660">http://dx.doi.org/10.11646/phytotaxa.585.2.1</mods:url>
</mods:location>
<mods:classification id="2051386B4D53AA6C68FF64CB99BB6DD4">journal article</mods:classification>
<mods:identifier id="42CB45AFBF9A284CC9684C83BBA3A01D" type="CLB-Dataset">53472</mods:identifier>
<mods:identifier id="0D6DEEE8121D369C16D64871B84DE7BA" type="DOI">10.11646/phytotaxa.585.2.1</mods:identifier>
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<mods:identifier id="F4BD091F47887659CC93F3DF21A17A49" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="804F5D048F48CC8E496372DA20A43463" type="Zenodo-Dep">7681555</mods:identifier>
</mods:mods>
<treatment id="03A287F4FFCE955B9097FF54FDA93143" ID-DOI="http://doi.org/10.5281/zenodo.7681919" ID-GBIF-Taxon="209347611" ID-Zenodo-Dep="7681919" LSID="urn:lsid:plazi:treatment:03A287F4FFCE955B9097FF54FDA93143" httpUri="http://treatment.plazi.org/id/03A287F4FFCE955B9097FF54FDA93143" lastPageNumber="94" pageId="23" pageNumber="94">
<subSubSection id="C3116569FFCE955B9097FF54FC7934C3" box="[136,962,158,185]" pageId="23" pageNumber="94" type="nomenclature">
<paragraph id="8BB436E2FFCE955B9097FF54FC7934C3" blockId="23.[136,1452,158,292]" box="[136,962,158,185]" pageId="23" pageNumber="94">
<heading id="D0FC818EFFCE955B9097FF54FC7934C3" box="[136,962,158,185]" fontSize="11" level="2" pageId="23" pageNumber="94" reason="4">
<emphasis id="B97FEAF0FFCE955B9097FF54FF1034C3" bold="true" box="[136,171,158,184]" pageId="23" pageNumber="94">4.1</emphasis>
<taxonomicName id="4C0B4D61FFCE955B90ADFF54FDED34C2" authority="(L. B. Sm.) Kessous" authorityName="Kessous" baseAuthorityName="L. B. Sm." box="[178,598,158,185]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="species" species="fibrosa">
<emphasis id="B97FEAF0FFCE955B90ADFF54FEC034C2" bold="true" box="[178,379,158,185]" italics="true" pageId="23" pageNumber="94">Werauhia fibrosa</emphasis>
(L.B.Sm.) Kessous
</taxonomicName>
&amp; A.F.Costa.
<emphasis id="B97FEAF0FFCE955B92E9FF54FCD334C3" box="[758,872,158,184]" italics="true" pageId="23" pageNumber="94">comb nov.</emphasis>
(
<figureCitation id="13302A67FFCE955B9368FF54FC0234C3" box="[887,953,158,184]" captionStart="FIGURE 8" captionStartId="22.[136,229,1992,2014]" captionTargetBox="[196,1385,198,1960]" captionTargetId="figure-25@22.[189,1398,190,1968]" captionTargetPageId="22" captionText="FIGURE 8. Werauhia fibrosa (L.B.Sm.) Kessous &amp; A.F.Costa. A. Holotype Maguire &amp; Politi 27912 (US; Courtesy of the United States National Herbarium). B. Detail of the inflorescence (adapted from Smith, 1957:317). C. Stigma cupulate (Photo: I.M. Kessous). D. Illustration of the outline of one petal appendage (I.M. Kessous)." figureDoi="http://doi.org/10.5281/zenodo.7681573" httpUri="https://zenodo.org/record/7681573/files/figure.png" pageId="23" pageNumber="94">Fig. 8</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFCE955B9097FF0CFE39355F" pageId="23" pageNumber="94" type="reference_group">
<paragraph id="8BB436E2FFCE955B9097FF0CFE39355F" blockId="23.[136,1452,158,292]" pageId="23" pageNumber="94">
<emphasis id="B97FEAF0FFCE955B9097FF0CFF4E34A7" bold="true" box="[136,245,198,220]" pageId="23" pageNumber="94">Basionym:</emphasis>
—
<taxonomicName id="4C0B4D61FFCE955B9112FF0CFDEC34A7" ID-CoL="7G4TJ" authority="Smith (1957: 315)" authorityName="Smith" authorityPageNumber="315" authorityYear="1957" box="[269,599,198,220]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="species" species="fibrosa">
<emphasis id="B97FEAF0FFCE955B9112FF0CFE2534A0" box="[269,414,198,219]" italics="true" pageId="23" pageNumber="94">Vriesea fibrosa</emphasis>
<bibRefCitation id="EF9A4B13FFCE955B91BBFF0CFDEC34A7" author="Smith, L. B." box="[420,599,198,220]" pageId="23" pageNumber="94" pagination="283 - 318" refId="ref23190" refString="Smith, L. B. (1957) Bromeliaceae. The Botany of the Guiana Highland - Part II. Memoirs of the New York Botanical Garden 9: 283 - 318." type="journal article" year="1957">Smith (1957: 315)</bibRefCitation>
</taxonomicName>
.
<emphasis id="B97FEAF0FFCE955B927EFF0CFD2134A0" bold="true" box="[609,666,198,219]" pageId="23" pageNumber="94">Type:</emphasis>
—
<collectingCountry id="F31C7672FFCE955B92ACFF0CFCFF34A0" box="[691,836,198,219]" name="Venezuela" pageId="23" pageNumber="94">VENEZUELA</collectingCountry>
.
<collectingRegion id="49CFF800FFCE955B9353FF0CFC0E34A0" box="[844,949,198,219]" country="Venezuela" name="Amazonas" pageId="23" pageNumber="94">Amazonas</collectingRegion>
: Cerro Sipapo (Paraque), frequent along open banks of lower Cano Negro,
<date id="FFB51022FFCE955B91BFFF20FDE53484" box="[416,606,234,256]" pageId="23" pageNumber="94" value="1948-12-25">25 December 1948</date>
,
<emphasis id="B97FEAF0FFCE955B9274FF20FCE03484" box="[619,859,234,255]" italics="true" pageId="23" pageNumber="94">Maguire &amp; Politi 27912</emphasis>
(
<typeStatus id="54B08840FFCE955B9375FF20FC7F357B" box="[874,964,234,256]" pageId="23" pageNumber="94" type="lectotype">lectotype</typeStatus>
US!, isolectotype NY!—Designated by
<bibRefCitation id="EF9A4B13FFCE955B9548FF20FECC355F" author="Smith, L. B. &amp; Downs, R. J." pageId="23" pageNumber="94" pagination="663 - 1492" refId="ref23503" refString="Smith, L. B. &amp; Downs, R. J. (1977) Bromeliaceae (Tillandsioideae). Flora Neotropica Monograph 14: 663 - 1492." type="journal article" year="1977">Smith &amp; Downs 1977: 1163</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFCE955B9097FE98FCBA30D3" pageId="23" pageNumber="94" type="description">
<paragraph id="8BB436E2FFCE955B9097FE98FC64371F" blockId="23.[136,1452,338,1337]" pageId="23" pageNumber="94">
<emphasis id="B97FEAF0FFCE955B9097FE98FEC33517" bold="true" box="[136,376,338,365]" pageId="23" pageNumber="94">Description:—Plant</emphasis>
terrestrial, with straight fibers at the base of the rosette (the remains of the veins of the decayed leaf sheath), flowering
<quantity id="4CF39B07FFCE955B918DFEBCFDBE35EA" box="[402,517,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="23" pageNumber="94" unit="cm" value="62.5" valueMax="65.0" valueMin="60.0">60−65 cm</quantity>
high.
<emphasis id="B97FEAF0FFCE955B924FFEBCFD1935EB" bold="true" box="[592,674,374,400]" pageId="23" pageNumber="94">Leaves</emphasis>
<quantity id="4CF39B07FFCE955B92B4FEBCFCA935EB" box="[683,786,374,400]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="23" pageNumber="94" unit="in" value="15.0" valueMax="20.0" valueMin="10.0">10−20 in</quantity>
number, forming an open utriculate rosette;
<emphasis id="B97FEAF0FFCE955B950CFEBCFAE735EB" box="[1299,1372,374,400]" italics="true" pageId="23" pageNumber="94">sheath</emphasis>
ovatetriangular, 4.5−6 ×
<quantity id="4CF39B07FFCE955B9179FE50FE7B35CE" box="[358,448,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="94" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3−4 cm</quantity>
, densely lepidote on both faces;
<emphasis id="B97FEAF0FFCE955B9325FE50FCC335CF" box="[826,888,410,436]" italics="true" pageId="23" pageNumber="94">blade</emphasis>
very narrowly subtriangular, 21−37 ×
<quantity id="4CF39B07FFCE955B9529FE50FA1C35CE" box="[1334,1447,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="23" pageNumber="94" unit="mm" value="13.0" valueMax="18.0" valueMin="8.0">8−18 mm</quantity>
, apex long attenuately acute.
<emphasis id="B97FEAF0FFCE955B91CFFE74FDD535A3" bold="true" box="[464,622,446,472]" pageId="23" pageNumber="94">Inflorescence</emphasis>
simple, 5–9 flowered,
<quantity id="4CF39B07FFCE955B9366FE74FC5135A2" box="[889,1002,446,473]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="23" pageNumber="94" unit="cm" value="50.0" valueMax="55.0" valueMin="45.0">45–55 cm</quantity>
long, lax, erect;
<emphasis id="B97FEAF0FFCE955B94B0FE74FAAF35A3" box="[1199,1300,446,472]" italics="true" pageId="23" pageNumber="94">peduncle</emphasis>
erect,
<quantity id="4CF39B07FFCE955B957BFE74FF113586" metricMagnitude="-1" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="23" pageNumber="94" unit="cm" value="37.5" valueMax="40.0" valueMin="35.0">35−40 cm</quantity>
long and
<quantity id="4CF39B07FFCE955B9106FE28FE303586" box="[281,395,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.7" metricValueMin="4.0" pageId="23" pageNumber="94" unit="mm" value="4.35" valueMax="4.7" valueMin="4.0">4–4.7 mm</quantity>
in diameter, glabrous;
<emphasis id="B97FEAF0FFCE955B9293FE28FC853587" box="[652,830,482,508]" italics="true" pageId="23" pageNumber="94">peduncle bracts</emphasis>
erect, imbricate at the base to remote distally, narrowly elliptic, 30−45 ×
<quantity id="4CF39B07FFCE955B9159FDCCFE7F365A" box="[326,452,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="23" pageNumber="94" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13−14 mm</quantity>
, lepidote on adaxial surface;
<emphasis id="B97FEAF0FFCE955B930BFDCCFCE1365B" box="[788,858,518,544]" italics="true" pageId="23" pageNumber="94">rachis</emphasis>
straight to slightly flexuous, ca.
<quantity id="4CF39B07FFCE955B94CBFDCCFAB6365A" box="[1236,1293,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="94" unit="cm" value="8.0">8 cm</quantity>
long,
<quantity id="4CF39B07FFCE955B9548FDCCFF0F363E" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="23" pageNumber="94" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.2–3.9 mm</quantity>
in diameter.
<emphasis id="B97FEAF0FFCE955B9155FDE0FE5D363F" bold="true" box="[330,486,554,580]" pageId="23" pageNumber="94">Floral bracts</emphasis>
orbicular, apex rounded, ca. 20 ×
<quantity id="4CF39B07FFCE955B9370FDE0FC7A363E" box="[879,961,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="94" unit="mm" value="20.0">20 mm</quantity>
, slightly carinate at the apex, much shorter than the sepals, with a yellowish mucilage, lepidote on the adaxial surface.
<emphasis id="B97FEAF0FFCE955B93D1FD84FB963613" bold="true" box="[974,1069,590,616]" pageId="23" pageNumber="94">Flowers</emphasis>
ca.
<quantity id="4CF39B07FFCE955B9445FD84FB133612" box="[1114,1192,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="94" unit="mm" value="35.0">35 mm</quantity>
long,
<quantity id="4CF39B07FFCE955B94F1FD84FADD3612" box="[1262,1382,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="23" pageNumber="94" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12–18 mm</quantity>
apart, distichous, suberect, secund;
<emphasis id="B97FEAF0FFCE955B91CEFDB8FDAC36F7" box="[465,535,626,652]" italics="true" pageId="23" pageNumber="94">sepals</emphasis>
ovate, 23–25 ×
<quantity id="4CF39B07FFCE955B92D1FDB8FCA436F6" box="[718,799,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="94" unit="mm" value="12.0">12 mm</quantity>
, free, slightly carinate at the apex, symmetric, lepidote on the margins of the abaxial surface, the adaxial surface with gland-like indument;
<emphasis id="B97FEAF0FFCE955B9400FD5CFBD936CB" box="[1055,1122,662,688]" italics="true" pageId="23" pageNumber="94">petals</emphasis>
narrowly elliptic, obtuse, ca. 32 ×
<quantity id="4CF39B07FFCE955B90A3FD70FEB436AE" box="[188,271,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="94" unit="mm" value="10.0">10 mm</quantity>
, greenish-white, reddish towards the apex, free;
<emphasis id="B97FEAF0FFCE955B9326FD70FC4436AF" box="[825,1023,698,724]" italics="true" pageId="23" pageNumber="94">petal appendages</emphasis>
ca.
<quantity id="4CF39B07FFCE955B942FFD70FB3236AE" box="[1072,1161,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="23" pageNumber="94" unit="mm" value="5.5">5.5 mm</quantity>
, adnate at the base of the petals for
<quantity id="4CF39B07FFCE955B90E9FD14FE8C3682" box="[246,311,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="94" unit="mm" value="3.0">3 mm</quantity>
, entire, with free lobe spatulate, reduced dactyloid one-fingered;
<emphasis id="B97FEAF0FFCE955B9419FD15FBDB3683" box="[1030,1120,735,760]" italics="true" pageId="23" pageNumber="94">stamens</emphasis>
equaling the petals (
<emphasis id="B97FEAF0FFCE955B955AFD15FA203683" box="[1349,1435,735,760]" italics="true" pageId="23" pageNumber="94">in sicco</emphasis>
);
<emphasis id="B97FEAF0FFCE955B9097FCC8FF5F3767" box="[136,228,770,796]" italics="true" pageId="23" pageNumber="94">filament</emphasis>
ca.
<quantity id="4CF39B07FFCE955B910CFCC8FED93766" box="[275,354,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="94" unit="mm" value="15.0">15 mm</quantity>
long, adnate to the petals for ca.
<quantity id="4CF39B07FFCE955B92C5FCC8FC8F3766" box="[730,820,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="94" unit="mm" value="1.7">1.7 mm</quantity>
;
<emphasis id="B97FEAF0FFCE955B9320FCC8FC333767" box="[831,904,770,796]" italics="true" pageId="23" pageNumber="94">anther</emphasis>
ca. 6 ×
<quantity id="4CF39B07FFCE955B93FBFCC8FB853766" box="[996,1086,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="94" unit="mm" value="1.9">1.9 mm</quantity>
;
<emphasis id="B97FEAF0FFCE955B9455FCC9FB313767" box="[1098,1162,771,796]" italics="true" pageId="23" pageNumber="94">ovary</emphasis>
ellipsoid, ca. 10 ×
<quantity id="4CF39B07FFCE955B957DFCC8FA1C3766" box="[1378,1447,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="94" unit="mm" value="6.0">6 mm</quantity>
;
<emphasis id="B97FEAF0FFCE955B9097FCECFF00373B" box="[136,187,806,832]" italics="true" pageId="23" pageNumber="94">style</emphasis>
ca.
<quantity id="4CF39B07FFCE955B90F4FCECFE80373A" box="[235,315,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="94" unit="mm" value="20.0">20 mm</quantity>
long;
<emphasis id="B97FEAF0FFCE955B919BFCEDFE74373B" box="[388,463,807,832]" italics="true" pageId="23" pageNumber="94">stigma</emphasis>
cupulate.
<emphasis id="B97FEAF0FFCE955B925BFCECFD36373B" bold="true" box="[580,653,806,832]" pageId="23" pageNumber="94">Fruits</emphasis>
unknown.
<emphasis id="B97FEAF0FFCE955B9313FCECFC13373B" bold="true" box="[780,936,806,832]" pageId="23" pageNumber="94">Distribution:</emphasis>
—This species occurs terrestrially on sand, in the region of the state of
<collectingRegion id="49CFF800FFCE955B9181FC80FDAC371F" box="[414,535,842,868]" country="Venezuela" name="Amazonas" pageId="23" pageNumber="94">Amazonas</collectingRegion>
,
<collectingCountry id="F31C7672FFCE955B923DFC80FD23371F" box="[546,664,842,868]" name="Venezuela" pageId="23" pageNumber="94">Venezuela</collectingCountry>
, ca.
<quantity id="4CF39B07FFCE955B92D5FC80FC9D371E" box="[714,806,842,869]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="94" unit="m" value="1400.0">1,400 m</quantity>
above sea level.
</paragraph>
<paragraph id="8BB436E2FFCE955B90A2FCA4FCBD378E" blockId="23.[136,1452,338,1337]" pageId="23" pageNumber="94">
<emphasis id="B97FEAF0FFCE955B90A2FCA4FD3F37F3" bold="true" box="[189,644,878,904]" pageId="23" pageNumber="94">Taxonomic and nomenclatural notes:</emphasis>
—Despite the lack of detailed information in the protologue and few available specimens, we observed a cupulate stigma in the
<typeStatus id="54B08840FFCE955B9305FC59FCF137D6" box="[794,842,915,941]" pageId="23" pageNumber="94">type</typeStatus>
specimen (
<emphasis id="B97FEAF0FFCE955B93D4FC59FB6237D7" box="[971,1241,914,941]" italics="true" pageId="23" pageNumber="94">Maguire &amp; Politi 27912</emphasis>
) (
<figureCitation id="13302A67FFCE955B94EDFC58FAF737D7" box="[1266,1356,914,940]" captionStart="FIGURE 8" captionStartId="22.[136,229,1992,2014]" captionTargetBox="[196,1385,198,1960]" captionTargetId="figure-25@22.[189,1398,190,1968]" captionTargetPageId="22" captionText="FIGURE 8. Werauhia fibrosa (L.B.Sm.) Kessous &amp; A.F.Costa. A. Holotype Maguire &amp; Politi 27912 (US; Courtesy of the United States National Herbarium). B. Detail of the inflorescence (adapted from Smith, 1957:317). C. Stigma cupulate (Photo: I.M. Kessous). D. Illustration of the outline of one petal appendage (I.M. Kessous)." figureDoi="http://doi.org/10.5281/zenodo.7681573" httpUri="https://zenodo.org/record/7681573/files/figure.png" pageId="23" pageNumber="94">Fig. 8D</figureCitation>
). Within
<taxonomicName id="4C0B4D61FFCE955B9097FC7CFE8C37AB" authorityName="Harms" authorityYear="1930" box="[136,311,950,976]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="subFamily" subFamily="Tillandsioideae">Tillandsioideae</taxonomicName>
, only members of the genus
<taxonomicName id="4C0B4D61FFCE955B9299FC7CFD4937AB" authorityName="Grant" authorityYear="1995" box="[646,754,950,976]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFCE955B9299FC7CFD4937AB" box="[646,754,950,976]" italics="true" pageId="23" pageNumber="94">Werauhia</emphasis>
</taxonomicName>
have this
<typeStatus id="54B08840FFCE955B9374FC7DFC2037AA" box="[875,923,951,977]" pageId="23" pageNumber="94">type</typeStatus>
of stigma (
<bibRefCitation id="EF9A4B13FFCE955B943EFC7CFB0837AA" author="Grant, J. R." box="[1057,1203,950,977]" pageId="23" pageNumber="94" pagination="1 - 57" refId="ref21768" refString="Grant, J. R. (1995 a) Bromelienstudien. The resurrection of Alcantarea and Werauhia, a new genus. Tropische und Subtropische Pflanzenwelt 91: 1 - 57." type="journal article" year="1995">Grant 1995a</bibRefCitation>
,
<bibRefCitation id="EF9A4B13FFCE955B94DFFC7CFA2437AB" author="Barfuss, M. H. J. &amp; Till, W. &amp; Leme, E. M. C. &amp; Pinzon, J. P. &amp; Manzanares, J. M. &amp; Halbritter, H. &amp; Samuel, R. &amp; Brown, G. K." box="[1216,1439,950,976]" pageId="23" pageNumber="94" pagination="1 - 97" refId="ref20630" refString="Barfuss, M. H. J., Till, W., Leme, E. M. C., Pinzon, J. P., Manzanares, J. M., Halbritter, H., Samuel, R. &amp; Brown, G. K. (2016) Taxonomic revision of Bromeliaceae subfam. Tillandsioideae based on a multi-locus DNA sequence phylogeny and morphology. Phytotaxa 279 (1): 1 - 97. https: // doi. org / 10.11646 / phytotaxa. 279.1.1" type="journal article" year="2016">
Barfuss
<emphasis id="B97FEAF0FFCE955B9500FC7DFAE137AB" box="[1311,1370,950,976]" italics="true" pageId="23" pageNumber="94">et al.</emphasis>
2016
</bibRefCitation>
), leading us to propose this new combination in
<taxonomicName id="4C0B4D61FFCE955B928CFC10FD44378F" authorityName="Grant" authorityYear="1995" box="[659,767,986,1012]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFCE955B928CFC10FD44378F" box="[659,767,986,1012]" italics="true" pageId="23" pageNumber="94">Werauhia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB436E2FFCE955B90A2FC35FCBA30D3" blockId="23.[136,1452,338,1337]" pageId="23" pageNumber="94">
<taxonomicName id="4C0B4D61FFCE955B90A2FC35FEAB3063" authorityName="Lindley" authorityYear="1843" box="[189,272,1023,1048]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFCE955B90A2FC35FEAB3063" box="[189,272,1023,1048]" italics="true" pageId="23" pageNumber="94">Vriesea</emphasis>
</taxonomicName>
has a convolute-bladed stigma, which is the most important distinguishing feature between these two genera. In addition, we observed the presence of dactyloid petal appendages (one-fingered) (
<figureCitation id="13302A67FFCE955B94A0FBE8FAA03047" box="[1215,1307,1058,1084]" captionStart="FIGURE 8" captionStartId="22.[136,229,1992,2014]" captionTargetBox="[196,1385,198,1960]" captionTargetId="figure-25@22.[189,1398,190,1968]" captionTargetPageId="22" captionText="FIGURE 8. Werauhia fibrosa (L.B.Sm.) Kessous &amp; A.F.Costa. A. Holotype Maguire &amp; Politi 27912 (US; Courtesy of the United States National Herbarium). B. Detail of the inflorescence (adapted from Smith, 1957:317). C. Stigma cupulate (Photo: I.M. Kessous). D. Illustration of the outline of one petal appendage (I.M. Kessous)." figureDoi="http://doi.org/10.5281/zenodo.7681573" httpUri="https://zenodo.org/record/7681573/files/figure.png" pageId="23" pageNumber="94">Fig. 8D</figureCitation>
), typical for
<taxonomicName id="4C0B4D61FFCE955B9097FB8CFF4F301B" authorityName="Grant" authorityYear="1995" box="[136,244,1094,1120]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FEAF0FFCE955B9097FB8CFF4F301B" box="[136,244,1094,1120]" italics="true" pageId="23" pageNumber="94">Werauhia</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="4C0B4D61FFCE955B91BEFB8DFDAE301B" authorityName="Smith" authorityYear="1957" box="[417,533,1094,1120]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="species" species="fibrosa">
<emphasis id="B97FEAF0FFCE955B91BEFB8DFDAE301B" box="[417,533,1094,1120]" italics="true" pageId="23" pageNumber="94">W. fibrosa</emphasis>
</taxonomicName>
fits the geographical distribution of other species in this genus (northern South America). This species is closely related to
<taxonomicName id="4C0B4D61FFCE955B9271FBA0FAB830FF" authority="(Mez &amp; Werckl.) Grant (1995 a: 32)" authorityName="Grant" authorityPageNumber="32" authorityYear="1995" baseAuthorityName="Mez &amp; Werckl." box="[622,1283,1130,1157]" class="Liliopsida" family="Bromeliaceae" genus="Werauhia" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="species" species="graminifolia">
<emphasis id="B97FEAF0FFCE955B9271FBA0FCD730FF" box="[622,876,1130,1156]" italics="true" pageId="23" pageNumber="94">Werauhia graminifolia</emphasis>
(Mez &amp; Werckl.)
<bibRefCitation id="EF9A4B13FFCE955B9427FBA0FAB830FF" author="Grant, J. R." box="[1080,1283,1130,1157]" pageId="23" pageNumber="94" pagination="1 - 57" refId="ref21768" refString="Grant, J. R. (1995 a) Bromelienstudien. The resurrection of Alcantarea and Werauhia, a new genus. Tropische und Subtropische Pflanzenwelt 91: 1 - 57." type="journal article" year="1995">Grant (1995a: 32)</bibRefCitation>
</taxonomicName>
, particularly in the graminoid rosette and the shape of the inflorescence.
</paragraph>
</subSubSection>
<subSubSection id="C3116569FFCE955B90A2FB78FDA93143" pageId="23" pageNumber="94" type="etymology">
<paragraph id="8BB436E2FFCE955B90A2FB78FE07308B" blockId="23.[136,1452,338,1337]" pageId="23" pageNumber="94">
<emphasis id="B97FEAF0FFCE955B90A2FB78FEF330B7" bold="true" box="[189,328,1202,1228]" pageId="23" pageNumber="94">Etymology:</emphasis>
—The specific epithet “
<taxonomicName id="4C0B4D61FFCE955B924CFB78FD1A30B7" authorityName="Smith" authorityYear="1957" box="[595,673,1202,1228]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="23" pageNumber="94" phylum="Tracheophyta" rank="species" species="fibrosa">
<emphasis id="B97FEAF0FFCE955B924CFB78FD1A30B7" box="[595,673,1202,1228]" italics="true" pageId="23" pageNumber="94">fibrosa</emphasis>
</taxonomicName>
” refers to the fibers at the base of the rosette (the remains of the veins of the decayed leaf sheath).
</paragraph>
<paragraph id="8BB436E2FFCE955B90A2FB30FDA93143" blockId="23.[136,1452,338,1337]" pageId="23" pageNumber="94">
<emphasis id="B97FEAF0FFCE955B90A2FB30FDC1316F" bold="true" box="[189,634,1274,1300]" pageId="23" pageNumber="94">Representative Specimen Examined:</emphasis>
—
<materialsCitation id="3B633CBFFFCE955B9288FB30FE2D3143" ID-GBIF-Occurrence="4039734322" collectingDate="1949-01-01" collectorName="Maguire &amp; Politi" country="Venezuela" location="Paraque" municipality="Cerro Sipapo" pageId="23" pageNumber="94" specimenCount="1" stateProvince="Amazonas">
<collectingCountry id="F31C7672FFCE955B9288FB30FCFB316F" box="[663,832,1274,1300]" name="Venezuela" pageId="23" pageNumber="94">VENEZUELA</collectingCountry>
.
<collectingRegion id="49CFF800FFCE955B9353FB30FC7E316F" box="[844,965,1274,1300]" country="Venezuela" name="Amazonas" pageId="23" pageNumber="94">Amazonas</collectingRegion>
,
<collectingMunicipality id="6BD0AC98FFCE955B93CCFB30FBD0316F" box="[979,1131,1274,1300]" pageId="23" pageNumber="94">Cerro Sipapo</collectingMunicipality>
(
<location id="8ED46039FFCE955B9462FB30FB66316F" LSID="urn:lsid:plazi:treatment:03A287F4FFCE955B9097FF54FDA93143:8ED46039FFCE955B9462FB30FB66316F" box="[1149,1245,1274,1300]" country="Venezuela" municipality="Cerro Sipapo" name="Paraque" pageId="23" pageNumber="94" stateProvince="Amazonas">Paráque</location>
),
<date id="FFB51022FFCE955B94ECFB30FA1C316F" box="[1267,1447,1274,1301]" pageId="23" pageNumber="94" value="1949-01-01">
<collectingDate id="EFF1E9CAFFCE955B94ECFB30FA1C316F" box="[1267,1447,1274,1301]" pageId="23" pageNumber="94" value="1949-01-01">1 January 1949</collectingDate>
</date>
,
<emphasis id="B97FEAF0FFCE955B9097FAD5FE2D3143" box="[136,406,1310,1337]" italics="true" pageId="23" pageNumber="94">
<collectorName id="26FE5334FFCE955B9097FAD5FF5C3143" box="[136,231,1311,1336]" pageId="23" pageNumber="94">Maguire</collectorName>
&amp;
<collectorName id="26FE5334FFCE955B9114FAD4FEF23143" box="[267,329,1310,1336]" pageId="23" pageNumber="94">Politi</collectorName>
28108
</emphasis>
</materialsCitation>
<materialsCitation id="3B633CBFFFCE955B9182FAD4FDB53143" ID-GBIF-Occurrence="4039734384" box="[413,526,1310,1336]" collectionCode="NY" country="United States of America" location="US" pageId="23" pageNumber="94" specimenCount="1">
(US,
<collectionCode id="ED1AAE27FFCE955B91C6FAD4FDBD3143" box="[473,518,1310,1336]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="23" pageNumber="94" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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