treatments-xml/data/03/A2/87/03A287E3FFFE1D1BFF0DF9D44E68F6B2.xml

<document id="BE4BE1B06BC5F8FD745FCCAC52F4DC2E" ID-DOI="10.11646/zootaxa.3609.2.5" ID-GBIF-Dataset="fad27fbd-389c-4c60-a927-77ac88afa267" ID-ISSN="1175-5326" ID-Zenodo-Dep="220831" ID-ZooBank="0B91082F-CCA0-4CE0-B221-AE62151CB985" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460571845318" checkinUser="plazi" docAuthor="Ban, Teruaki &amp; Ishikawa, Tadashi" docDate="2013" docId="03A287E3FFFE1D1BFF0DF9D44E68F6B2" docLanguage="en" docName="zt03609p203.pdf" docOrigin="Zootaxa 3609 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sadoletus izzardi Hidaka" docType="treatment" docVersion="6" lastPageNumber="201" masterDocId="FF9BFF9BFFFF1D1DFF9AFFD04C1CF00E" masterDocTitle="The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan" masterLastPageNumber="203" masterPageNumber="195" pageNumber="196" updateTime="1698322288420" updateUser="plazi">
<mods:mods id="C91F7EE0BF1B02C2BD8CF69C6AC07DF3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F57E513DA6DE0C6A7493AEF8CA440B15">
<mods:title id="CDED29432C8B1C4E9473D82C6802D90A">The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="63DEFC7E19EDFE3CDEF99EFEDFB6BFDE" type="personal">
<mods:role id="3DF2183DA5E5DCA2160A0D44399B3FEB">
<mods:roleTerm id="8B6A0B4F5AA91D6C1BBB64E6A9DE88EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="355D9C55D0C04F7041851FD92C10DB83">Ban, Teruaki</mods:namePart>
</mods:name>
<mods:name id="1F98827D5515EF9C1BF095FECDAB1C2F" type="personal">
<mods:role id="8C33BEB833CECF36A591B62011DB55A1">
<mods:roleTerm id="1E4863BE2CD9EED75936F384182D55CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="695053718A4EB40F309385B616E687EA">Ishikawa, Tadashi</mods:namePart>
</mods:name>
<mods:typeOfResource id="5413F86F6C1479B521D85C834903ABA4">text</mods:typeOfResource>
<mods:relatedItem id="D73B4AE91F7A9D6DCBF6B65FA73C0FED" type="host">
<mods:titleInfo id="C8E6800EC49D56E37E871BA0A2868645">
<mods:title id="77D19BA016E0D7B3AF949D3A9DE85966">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D1F8BB7EE8BB6D153EB099FD14A530A8">
<mods:date id="B3C15D3AC6B69EB39A2EF97E310DEF97">2013</mods:date>
<mods:detail id="C60F0E0F40509050F859C01617754F78" type="volume">
<mods:number id="02D9EDC2891F3E125BFB1BBF503026CF">3609</mods:number>
</mods:detail>
<mods:detail id="911A2D477F7A85A6089E8DA58A781110" type="issue">
<mods:number id="2A333A37A964508F604BA0B469B66F16">2</mods:number>
</mods:detail>
<mods:extent id="A423987063A39E4AE1BBB1892D8F8710" unit="page">
<mods:start id="8D24836252209F2D2339A87BCC97E0F1">195</mods:start>
<mods:end id="FE8E50EE1C0C8B1133AA2AB188ADDA9C">203</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EE1237E1A1636144C078B36D1B45D558">journal article</mods:classification>
<mods:identifier id="4DB38C5122C6494EBC593B0D57354C42" type="DOI">10.11646/zootaxa.3609.2.5</mods:identifier>
<mods:identifier id="3C0DBC8847C7DE73082F8D6A84C499CA" type="GBIF-Dataset">fad27fbd-389c-4c60-a927-77ac88afa267</mods:identifier>
<mods:identifier id="5FC759338B224A1CCC733582CC0C307A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4A4A3BFC577AB377C6EEF0B4B8F89181" type="Zenodo-Dep">220831</mods:identifier>
<mods:identifier id="671B6FD50A953505CDB8EFE0436459D4" type="ZooBank">0B91082F-CCA0-4CE0-B221-AE62151CB985</mods:identifier>
</mods:mods>
<treatment id="03A287E3FFFE1D1BFF0DF9D44E68F6B2" ID-DOI="http://doi.org/10.5281/zenodo.6151271" ID-GBIF-Taxon="119558918" ID-Zenodo-Dep="6151271" LSID="urn:lsid:plazi:treatment:03A287E3FFFE1D1BFF0DF9D44E68F6B2" httpUri="http://treatment.plazi.org/id/03A287E3FFFE1D1BFF0DF9D44E68F6B2" lastPageId="6" lastPageNumber="201" pageId="1" pageNumber="196">
<subSubSection id="C311657EFFFE1D1CFF0DF9D44F92F68A" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="8BB436F5FFFE1D1CFF0DF9D44DDBF613" blockId="1.[151,455,1539,1598]" box="[151,455,1539,1566]" pageId="1" pageNumber="196">
<heading id="D0FC8199FFFE1D1CFF0DF9D44DDBF613" bold="true" box="[151,455,1539,1566]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<taxonomicName id="4C0B4D76FFFE1D1CFF0DF9D44DDBF613" authority="Hidaka" authorityName="Hidaka" box="[151,455,1539,1566]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="196" phylum="Tracheophyta" rank="species" species="izzardi">
<emphasis id="B97FEAE7FFFE1D1CFF0DF9D44DDBF613" bold="true" box="[151,455,1539,1566]" pageId="1" pageNumber="196">
<emphasis id="B97FEAE7FFFE1D1CFF0DF9D44D7FF610" bold="true" box="[151,355,1540,1566]" italics="true" pageId="1" pageNumber="196">Sadoletus izzardi</emphasis>
Hidaka
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB436F5FFFE1D1CFF0DF9F64D4BF630" blockId="1.[151,455,1539,1598]" box="[151,343,1574,1598]" pageId="1" pageNumber="196">
(
<figureCitation id="13302A70FFFE1D1CFF05F9F64CF1F630" box="[159,237,1574,1598]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="1" pageNumber="196">Figs. 1</figureCitation>
, 
<figureCitation id="13302A70FFFE1D1CFF60F9F64D16F630" box="[250,266,1574,1598]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="1" pageNumber="196">5</figureCitation>
, 
<figureCitation id="13302A70FFFE1D1CFE8CF9F64D53F630" box="[278,335,1574,1598]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="1" pageNumber="196">8–14</figureCitation>
)
</paragraph>
<paragraph id="8BB436F5FFFE1D1CFF0DF9BF4F92F68A" blockId="1.[151,910,1646,1669]" box="[151,910,1646,1669]" pageId="1" pageNumber="196">
<taxonomicName id="4C0B4D76FFFE1D1CFF0DF9BF4E15F68A" authority="Hidaka, 1959: 199" authorityName="Hidaka" authorityPageNumber="199" authorityYear="1959" box="[151,521,1646,1669]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="196" phylum="Tracheophyta" rank="species" species="izzardi">
<emphasis id="B97FEAE7FFFE1D1CFF0DF9BF4D58F68B" box="[151,324,1647,1669]" italics="true" pageId="1" pageNumber="196">Sadoletus izzardi</emphasis>
Hidaka, 1959: 199
</taxonomicName>
; Slater, 1964: 776; Miyoshi, 1997: 43.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFE1D1CFF0DF9614CEAF757" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="8BB436F5FFFE1D1CFF0DF9614CEAF757" blockId="1.[151,1436,1713,2025]" pageId="1" pageNumber="196">
<emphasis id="B97FEAE7FFFE1D1CFF0DF9614D98F6C4" bold="true" box="[151,388,1713,1738]" pageId="1" pageNumber="196">Material examined.</emphasis>
Non-types: 
<date id="FFB51035FFFE1D1CFD8FF9614E6BF6C7" box="[533,631,1713,1737]" pageId="1" pageNumber="196" value="1914-03-22">22 3 14</date>
Ƥ (NSMT, TUA) — 
<emphasis id="B97FEAE7FFFE1D1CFCE6F9624FC5F6C4" bold="true" box="[892,985,1714,1738]" pageId="1" pageNumber="196">Honshu</emphasis>
[Yamaguchi Pref.] 1 3, Oomine Pass, Hikari-shi, 
<date id="FFB51035FFFE1D1CFE80F9054D8DF6E3" box="[282,401,1749,1773]" pageId="1" pageNumber="196" value="1997-02-12">12.II.1997</date>
, K. Miyoshi; 1 Ƥ, Hikari-shi, 
<date id="FFB51035FFFE1D1CFD6CF9054F71F6E3" box="[758,877,1749,1773]" pageId="1" pageNumber="196" value="1997-02-19">19.II.1997</date>
, K. Miyoshi; 5 3 1 Ƥ, Haginodaira, Hikari-shi, 
<date id="FFB51035FFFE1D1CFF0DF9294D3EF71F" box="[151,290,1785,1809]" pageId="1" pageNumber="196" value="2009-07-18">18.VII.2009</date>
, TB; 
<date id="FFB51035FFFE1D1CFEFDF9294DD5F71F" box="[359,457,1785,1809]" pageId="1" pageNumber="196" value="2010-03-15">15 3 10</date>
Ƥ, Mt. Senbou, Hikari-shi, 
<date id="FFB51035FFFE1D1CFC8FF9294FBDF71F" box="[789,929,1785,1809]" pageId="1" pageNumber="196" value="2009-07-18">18.VII.2009</date>
, TB. 
<emphasis id="B97FEAE7FFFE1D1CFC7FF9294855F71C" bold="true" box="[997,1097,1785,1810]" pageId="1" pageNumber="196">Shikoku</emphasis>
[Ehime Pref.] 1 3, Sugitate, 
<date id="FFB51035FFFE1D1CFF0DF8CD4D10F73B" box="[151,268,1821,1845]" pageId="1" pageNumber="196" value="1951-05-16">16.V.1951</date>
, M. Miyatake; 1 Ƥ, Sugitate, Matsuyama, 
<date id="FFB51035FFFE1D1CFD76F8CD4F71F73B" box="[748,877,1821,1845]" pageId="1" pageNumber="196" value="1953-03-31">31.III.1953</date>
, T. Mohri; 1 Ƥ, Matsuyama, Fumoto, 
<date id="FFB51035FFFE1D1CFA80F8CD498BF73B" box="[1306,1431,1821,1845]" pageId="1" pageNumber="196" value="1998-07-09">9.VII.1998</date>
, S. Yano.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFE1D1EFF5DF8B548EBF7E8" lastPageId="3" lastPageNumber="198" pageId="1" pageNumber="196" type="diagnosis">
<paragraph id="8BB436F5FFFE1D1EFF5DF8B548EBF7E8" blockId="1.[151,1436,1713,2025]" lastBlockId="3.[151,1437,1926,2022]" lastPageId="3" lastPageNumber="198" pageId="1" pageNumber="196">
<emphasis id="B97FEAE7FFFE1D1CFF5DF8B54D5DF770" bold="true" box="[199,321,1893,1918]" pageId="1" pageNumber="196">Diagnosis.</emphasis>
Recognized among congeners of 
<emphasis id="B97FEAE7FFFE1D1CFD4AF8B54F21F773" box="[720,829,1893,1917]" italics="true" pageId="1" pageNumber="196">Sadoletus</emphasis>
by a combination of the following characters: head about two-thirds as long as width across eyes (
<figureCitation id="13302A70FFFE1D1CFD3FF8594EFBF7AF" box="[677,743,1929,1953]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
); antennae almost brownish yellow or orange, with segments III and IV often darker than basal two segments (
<figureCitation id="13302A70FFFE1D1CFD40F87D4F02F7CB" box="[730,798,1965,1989]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
); prothorax and scutellum blackish and shiny (
<figureCitation id="13302A70FFFE1D1CFADFF87D4996F7CB" box="[1349,1418,1965,1989]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
); humeral angles strongly prominent posteriad (
<figureCitation id="13302A70FFFE1D1CFD35F8014EEDF7E7" box="[687,753,2001,2025]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
); scutellum with Y-shaped carina; hemelytra not exceeding apex of abdomen (
<figureCitation id="13302A70FFFC1D1EFEE1F8564DDFF790" box="[379,451,1926,1950]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="3" pageNumber="198">Fig. 1</figureCitation>
); femora and tibiae pale orange to brownish yellow (
<figureCitation id="13302A70FFFC1D1EFBCEF8564880F790" box="[1108,1180,1926,1950]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="3" pageNumber="198">Fig. 1</figureCitation>
); tibiae without dark annulations (
<figureCitation id="13302A70FFFC1D1EFEBCF87A4D7BF7CC" box="[294,359,1962,1986]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="3" pageNumber="198">Fig. 1</figureCitation>
); paramere strongly curved throughout its length, with projection at base (
<figureCitation id="13302A70FFFC1D1EFB39F87A48FAF7CC" box="[1187,1254,1962,1986]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="3" pageNumber="198">Fig. 9</figureCitation>
); and ovipositor more than 1.0 mm, nearly reaching sternite IV in rest (
<figureCitation id="13302A70FFFC1D1EFD63F81E4F21F7E8" box="[761,829,1998,2022]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="3" pageNumber="198">Fig. 5</figureCitation>
). Body length 
<quantity id="4CF39B10FFFC1D1EFC79F81E489AF7E8" box="[995,1158,1998,2022]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.55" metricValueMin="4.25" pageId="3" pageNumber="198" unit="mm" value="4.9" valueMax="5.55" valueMin="4.25">4.25–5.55 mm</quantity>
in length.
</paragraph>
</subSubSection>
<caption id="DF74667DFFFD1D1FFF0DF87D4DD0F7EC" httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="2" pageNumber="197" targetBox="[233,1367,201,1927]" targetPageId="2">
<paragraph id="8BB436F5FFFD1D1FFF0DF87D4DD0F7EC" blockId="2.[151,1436,1965,2018]" pageId="2" pageNumber="197">
<emphasis id="B97FEAE7FFFD1D1FFF0DF87D4D26F7CD" bold="true" box="[151,314,1965,1988]" pageId="2" pageNumber="197">FIGURES 1–4.</emphasis>
Habitus of Japanese 
<emphasis id="B97FEAE7FFFD1D1FFD8EF87E4E6AF7CA" box="[532,630,1966,1988]" italics="true" pageId="2" pageNumber="197">Sadoletus</emphasis>
. 1, 
<taxonomicName id="4C0B4D76FFFD1D1FFD07F87E4EE6F7CA" box="[669,762,1966,1988]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="197" phylum="Tracheophyta" rank="species" species="izzardi">
<emphasis id="B97FEAE7FFFD1D1FFD07F87E4EE6F7CA" box="[669,762,1966,1988]" italics="true" pageId="2" pageNumber="197">S. izzardi</emphasis>
</taxonomicName>
, male; 2–3, 
<taxonomicName id="4C0B4D76FFFD1D1FFCE0F87E4817F7CA" box="[890,1035,1966,1988]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="197" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFFD1D1FFCE0F87E4817F7CA" box="[890,1035,1966,1988]" italics="true" pageId="2" pageNumber="197">S. ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C579CFFFD1D1FFB88F87E487FF7CD" box="[1042,1123,1966,1987]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B97FEAE7FFFD1D1FFB88F87E487FF7CD" bold="true" box="[1042,1123,1966,1987]" pageId="2" pageNumber="197">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (2) and female (3); 4, 
<taxonomicName id="4C0B4D76FFFD1D1FFA13F87E4CC3F7EC" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="197" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFFD1D1FFA13F87E4CC3F7EC" italics="true" pageId="2" pageNumber="197">S. valdezi</emphasis>
</taxonomicName>
, male. Scales: 1.0 mm.
</paragraph>
</caption>
<caption id="DF74667DFFFC1D1EFF0DFDB34E66F296" httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="3" pageNumber="198" targetBox="[177,1390,213,560]" targetPageId="3">
<paragraph id="8BB436F5FFFC1D1EFF0DFDB34E66F296" blockId="3.[151,1435,611,664]" pageId="3" pageNumber="198">
<emphasis id="B97FEAE7FFFC1D1EFF0DFDB34D20F277" bold="true" box="[151,316,611,634]" pageId="3" pageNumber="198">FIGURES 5–7.</emphasis>
Apical part of female abdomen, ventral view. 5, 
<taxonomicName id="4C0B4D76FFFC1D1EFCA5FDB44FF2F274" box="[831,1006,612,634]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="198" phylum="Tracheophyta" rank="species" species="izzardi">
<emphasis id="B97FEAE7FFFC1D1EFCA5FDB44FF2F274" box="[831,1006,612,634]" italics="true" pageId="3" pageNumber="198">Sadoletus izzardi</emphasis>
</taxonomicName>
; 6, 
<taxonomicName id="4C0B4D76FFFC1D1EFB83FDB548B7F274" box="[1049,1195,612,634]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="198" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFFC1D1EFB83FDB548B7F274" box="[1049,1195,612,634]" italics="true" pageId="3" pageNumber="198">S. ryukyuensis</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7FFFC1D1EFB2EFDB4491BF277" bold="true" box="[1204,1287,612,633]" pageId="3" pageNumber="198">
<taxonomicNameLabel id="A24C579CFFFC1D1EFB2EFDB4491BF277" box="[1204,1287,612,633]" pageId="3" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 7, 
<taxonomicName id="4C0B4D76FFFC1D1EFAA8FDB54989F274" box="[1330,1429,612,634]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="198" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFFC1D1EFAA8FDB54989F274" box="[1330,1429,612,634]" italics="true" pageId="3" pageNumber="198">S. valdezi</emphasis>
</taxonomicName>
. Scales: 0.5 mm. abbreviations: st7, sternite VII.
</paragraph>
</caption>
<caption id="DF74667DFFFC1D1EFF0DF93C4EB5F74E" httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="3" pageNumber="198" targetBox="[169,1407,740,1726]" targetPageId="3">
<paragraph id="8BB436F5FFFC1D1EFF0DF93C4EB5F74E" blockId="3.[151,1436,1772,1857]" pageId="3" pageNumber="198">
<emphasis id="B97FEAE7FFFC1D1EFF0DF93C4D56F70C" bold="true" box="[151,330,1772,1795]" pageId="3" pageNumber="198">FIGURES 8–14.</emphasis>
<taxonomicName id="4C0B4D76FFFC1D1EFEC8F93D4E1DF70D" box="[338,513,1773,1795]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="198" phylum="Tracheophyta" rank="species" species="izzardi">
<emphasis id="B97FEAE7FFFC1D1EFEC8F93D4E1DF70D" box="[338,513,1773,1795]" italics="true" pageId="3" pageNumber="198">Sadoletus izzardi</emphasis>
</taxonomicName>
. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10–11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12–13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C311657EFFFB1D19FF5DFF474E67F189" pageId="4" pageNumber="199" type="description">
<paragraph id="8BB436F5FFFB1D19FF5DFF474924F131" blockId="4.[151,1436,151,860]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DFF474DF6F0BE" bold="true" box="[199,490,151,176]" pageId="4" pageNumber="199">Description of genitalia.</emphasis>
Male: Pygophore (
<figureCitation id="13302A70FFFB1D19FD50FF474F13F0A1" box="[714,783,151,175]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="4" pageNumber="199">Fig. 8</figureCitation>
) nearly globe-like in dorsal view, covered with suberect setae in apical half; dorsal aperture wide, more than one-third as wide at middle as pygophore, with collar along anterior margin. Paramere (
<figureCitation id="13302A70FFFB1D19FE4BFF0F4E0AF0F9" box="[465,534,223,247]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="4" pageNumber="199">Fig. 9</figureCitation>
) strongly curved in apical two-thirds, covered with several short to long, erect setae in basal one-third, and with projection near base. Phallosoma (
<figureCitation id="13302A70FFFB1D19FC29FED44821F112" box="[947,1085,260,284]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="4" pageNumber="199">Figs. 10, 11</figureCitation>
) twice as long as its width in ventral view. Endosoma (
<figureCitation id="13302A70FFFB1D19FE2EFEF74E25F131" box="[436,569,295,319]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="4" pageNumber="199">Figs. 12, 13</figureCitation>
) a little twisted in apical part, lacking definite sclerotized processes.
</paragraph>
<paragraph id="8BB436F5FFFB1D19FF5DFE9C4E67F189" blockId="4.[151,1436,151,860]" pageId="4" pageNumber="199">
Females: Ovipositor (
<figureCitation id="13302A70FFFB1D19FE23FE9C4DE7F16A" box="[441,507,332,356]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="4" pageNumber="199">Fig. 5</figureCitation>
) 
<quantity id="4CF39B10FFFB1D19FD90FE9C4EB0F16A" box="[522,684,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" metricValueMax="1.6" metricValueMin="1.38" pageId="4" pageNumber="199" unit="mm" value="1.49" valueMax="1.6" valueMin="1.38">1.38–1.60 mm</quantity>
in length, nearly reaching sternite IV in rest. Spermatheca (
<figureCitation id="13302A70FFFB1D19FAD9FE9C498FF16A" box="[1347,1427,332,356]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1772,1795]" captionTargetBox="[169,1407,740,1726]" captionTargetId="figure@3.[151,1436,714,1751]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Sadoletus izzardi. 8, Pygophore, dorsal view (setae omitted from left half); 9, left paramere, dorsal view; 10 – 11, phallus (endosoma not everted), ventral (10) and lateral (11) views; 12 – 13, phallus (endosoma everted), ventral (12) and lateral (13) views; 14, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/220834/files/figure.png" pageId="4" pageNumber="199">Fig. 14</figureCitation>
) almost globe, with rounded lobe near apex.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D19FF5DFE434F61F1A2" box="[199,893,403,428]" pageId="4" pageNumber="199" type="distribution">
<paragraph id="8BB436F5FFFB1D19FF5DFE434F61F1A2" blockId="4.[151,1436,151,860]" box="[199,893,403,428]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DFE434D7CF1A2" bold="true" box="[199,352,403,428]" pageId="4" pageNumber="199">Distribution.</emphasis>
<collectingCountry id="F31C7665FFFB1D19FEFDFE454DB7F1A2" box="[359,427,405,428]" name="Japan" pageId="4" pageNumber="199">Japan</collectingCountry>
: Honshu, Shodoshima, Shikoku, Kyushu.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D19FF5DFE684FCFF352" pageId="4" pageNumber="199" type="discussion">
<paragraph id="8BB436F5FFFB1D19FF5DFE684DB3F232" blockId="4.[151,1436,151,860]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DFE684D27F1DE" bold="true" box="[199,315,440,464]" pageId="4" pageNumber="199">Remarks.</emphasis>
In addition to the characters in the key mentioned above, this species is separable from other Japanese congeners of 
<emphasis id="B97FEAE7FFFB1D19FE01FE0C4E14F1FA" box="[411,520,476,500]" italics="true" pageId="4" pageNumber="199">Sadoletus</emphasis>
by having an almost concolorous antenna, legs, and abdomen; the hemelytra not exceeding the apex of the abdomen; and the female abdominal sternites V and VI with the posterior margins strongly curved anteriad.
</paragraph>
<paragraph id="8BB436F5FFFB1D19FF5DFD974E7CF2C2" blockId="4.[151,1436,151,860]" pageId="4" pageNumber="199">
This species is more similar in the male and female genital structures to species of other heterogastrid genera such as 
<taxonomicName id="4C0B4D76FFFB1D19FF61FDBC4DE7F28A" authority="Scudder" authorityName="Scudder" box="[251,507,620,644]" class="Insecta" family="Heterogastridae" genus="Parathyginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7FFFB1D19FF61FDBC4D8EF28A" box="[251,402,620,644]" italics="true" pageId="4" pageNumber="199">Parathyginus</emphasis>
Scudder
</taxonomicName>
and 
<taxonomicName id="4C0B4D76FFFB1D19FDA6FDBD4F5DF28A" authority="Schilling" authorityName="Schilling" box="[572,833,620,644]" class="Insecta" family="Heterogastridae" genus="Heterogaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7FFFB1D19FDA6FDBD4ED3F28A" box="[572,719,621,644]" italics="true" pageId="4" pageNumber="199">Heterogaster</emphasis>
Schilling
</taxonomicName>
than to other Japanese members of 
<emphasis id="B97FEAE7FFFB1D19FB66FDBC4975F28A" box="[1276,1385,620,644]" italics="true" pageId="4" pageNumber="199">Sadoletus</emphasis>
(cf. Servadei 1951; our observation). It may suggest that a taxonomic revision of 
<emphasis id="B97FEAE7FFFB1D19FC66FD404875F2A6" box="[1020,1129,656,680]" italics="true" pageId="4" pageNumber="199">Sadoletus</emphasis>
with a careful examination of all the species of this genus is needed.
</paragraph>
<paragraph id="8BB436F5FFFB1D19FF5DFD074FCFF352" blockId="4.[151,1436,151,860]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DFD074D95F2FE" bold="true" box="[199,393,727,752]" pageId="4" pageNumber="199">Biological notes.</emphasis>
According to Miyoshi (1997), this species was collected from 
<taxonomicName id="4C0B4D76FFFB1D19FBD6FD084D34F31A" authority="Matsumura" authorityName="Matsumura" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="199" phylum="Tracheophyta" rank="species" species="sieboldiana">
<emphasis id="B97FEAE7FFFB1D19FBD6FD084909F2FE" box="[1100,1301,728,752]" italics="true" pageId="4" pageNumber="199">Alnus sieboldiana</emphasis>
Matsumura (Betulaceae)
</taxonomicName>
. In addition, 
<taxonomicName id="4C0B4D76FFFB1D19FE24FD2C4809F31A" authority="Sieb. et Zucc." authorityName="Sieb. et Zucc." box="[446,1045,764,788]" class="Magnoliopsida" family="Juglandaceae" genus="Platycarya" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="199" phylum="Tracheophyta" rank="species" species="strobilacea">
<emphasis id="B97FEAE7FFFB1D19FE24FD2C4EA3F31A" box="[446,703,764,788]" italics="true" pageId="4" pageNumber="199">Platycarya strobilacea</emphasis>
Sieb. et Zucc. (Juglandaceae)
</taxonomicName>
was also confirmed as a new host plant of this heterogastrid by our field research in Yamaguchi, western Honshu. A number of adults and nymphs, often hiding in chinks of the cone-like nuts, were collected from the trees.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D19FF0DFC7F4D8FF45E" pageId="4" pageNumber="199" type="description">
<paragraph id="8BB436F5FFFB1D19FF0DFC7F4D9EF3E2" blockId="4.[151,712,943,1004]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF0DFC7F4ED4F3C6" bold="true" box="[151,712,943,968]" pageId="4" pageNumber="199">
<taxonomicName id="4C0B4D76FFFB1D19FF0DFC7F4DD3F3C6" authority="Ban" authorityName="Ban" box="[151,463,943,968]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="199" phylum="Tracheophyta" rank="species" species="ryukyuensis">
<emphasis id="B97FEAE7FFFB1D19FF0DFC7F4D85F3C6" bold="true" box="[151,409,943,968]" italics="true" pageId="4" pageNumber="199">Sadoletus ryukyuensis</emphasis>
Ban
</taxonomicName>
et Ishikawa, sp. nov.
</emphasis>
(
<figureCitation id="13302A70FFFB1D19FF05FC044D17F3E2" box="[159,267,980,1004]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2–3</figureCitation>
, 
<figureCitation id="13302A70FFFB1D19FE8DFC044D3BF3E2" box="[279,295,980,1004]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="4" pageNumber="199">6</figureCitation>
, 
<figureCitation id="13302A70FFFB1D19FEA8FC044D66F3E2" box="[306,378,980,1004]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="4" pageNumber="199">15–22</figureCitation>
)
</paragraph>
<paragraph id="8BB436F5FFFB1D19FF0DFBCC4D8FF45E" blockId="4.[151,1435,1051,1104]" pageId="4" pageNumber="199">
<taxonomicName id="4C0B4D76FFFB1D19FF0DFBCC4D50F43C" box="[151,332,1052,1074]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="199" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFFB1D19FF0DFBCC4D50F43C" box="[151,332,1052,1074]" italics="true" pageId="4" pageNumber="199">Sadoletus valdezi</emphasis>
</taxonomicName>
(non Bergroth, 1918): Hidaka, 1959: 200; Miyamoto and Yasunaga, 1989: 175; Tomokuni, 1989: 191; Hayashi, 2002: 141.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D19FF0DFBAD48D0F667" pageId="4" pageNumber="199" type="materials_examined">
<paragraph id="8BB436F5FFFB1D19FF0DFBAD48D0F667" blockId="4.[151,1437,1149,2038]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF0DFBAD4D3EF498" bold="true" box="[151,290,1149,1174]" pageId="4" pageNumber="199">
<typeStatus id="54B08857FFFB1D19FF0DFBAD4CCEF49B" box="[151,210,1149,1173]" pageId="4" pageNumber="199">Type</typeStatus>
series.
</emphasis>
<typeStatus id="54B08857FFFB1D19FEB2FBAD4D89F49B" box="[296,405,1149,1173]" pageId="4" pageNumber="199" type="holotype">Holotype</typeStatus>
: 3, Shirahama, Iriomote-jima Is., the Ryukyus, 
<date id="FFB51035FFFB1D19FC32FBAD4802F49B" box="[936,1054,1149,1173]" pageId="4" pageNumber="199" value="2008-05-13">13.V.2008</date>
, TB (TUA). 
<typeStatus id="54B08857FFFB1D19FB36FBAE4901F498" box="[1196,1309,1150,1174]" pageId="4" pageNumber="199" type="paratype">Paratypes</typeStatus>
: 39 3 34 Ƥ — 
<emphasis id="B97FEAE7FFFB1D19FF21FB724D4DF4B4" bold="true" box="[187,337,1186,1210]" pageId="4" pageNumber="199">the Ryukyus</emphasis>
[Yaku-shima Is.] 1 3, Hirauchi, 
<date id="FFB51035FFFB1D19FD50FB714F23F4B7" box="[714,831,1185,1209]" pageId="4" pageNumber="199" value="1960-05-25">25.V.1960</date>
, Y. Kimura (OMNH). [Tokara Islands] 1 Ƥ, Nakanoshima Is., 
<date id="FFB51035FFFB1D19FE8BFB154D8EF4D3" box="[273,402,1221,1245]" pageId="4" pageNumber="199" value="1953-06-13">13.VI.1953</date>
, T. Nakane (ELKU). [Tokuno-shima Is.] 5 3 3 Ƥ, Matsubara, Amagi-chô, 
<date id="FFB51035FFFB1D19FA98FB154971F4D3" box="[1282,1389,1221,1245]" pageId="4" pageNumber="199" value="1988-10-02">2.X.1988</date>
, M. Tomokuni (NSMT). [Okinawa Is.] 1 Ƥ, Matsuda, Ginoza-son, 
<date id="FFB51035FFFB1D19FCC9FB394FCBF50F" box="[851,983,1257,1281]" pageId="4" pageNumber="199" value="1995-05-04" valueMax="1995-05-08" valueMin="1995-05-04">4–8.V.1995</date>
, M. Sugimoto (TUA). [Miyako-jima Is.] 1 Ƥ, Higashi-sokobaru, Hirara, 
<date id="FFB51035FFFB1D19FD9EFADD4E60F52B" box="[516,636,1293,1317]" pageId="4" pageNumber="199" value="1999-10-30">30.X.1999</date>
, light trap, TI (TUA). [Ishigaki-jima Is.] 2 3, Kabira, 
<date id="FFB51035FFFB1D19FB6FFADD4964F52B" box="[1269,1400,1293,1317]" pageId="4" pageNumber="199" value="1966-06-15">15.VI.1966</date>
, S. Tachikawa (TUA); 1 3 2 Ƥ, Banna Park, 
<date id="FFB51035FFFB1D19FDFCFAE14EFAF547" box="[614,742,1329,1353]" pageId="4" pageNumber="199" value="1997-03-25">25.III.1997</date>
, H. Yoshitake (TUA); 1 Ƥ, Sokobaru, 
<date id="FFB51035FFFB1D19FB08FAE14914F547" box="[1170,1288,1329,1353]" pageId="4" pageNumber="199" value="2000-05-24">24.V.2000</date>
, TI (TUA); 1 3, Tomino, 
<date id="FFB51035FFFB1D19FEBAFA854DBFF563" box="[288,419,1365,1389]" pageId="4" pageNumber="199" value="2002-09-27">27.IX.2002</date>
, H. Yamamoto (ICMI); 1 Ƥ, Omoto, 
<date id="FFB51035FFFB1D19FCD3FA854FA8F563" box="[841,948,1365,1389]" pageId="4" pageNumber="199" value="2002-10-01">1.X.2002</date>
, light trap, H. Yamamoto (ICMI); 1 3, Mt. Omoto-dake, 
<date id="FFB51035FFFB1D19FEADFAA94DB6F59F" box="[311,426,1401,1425]" pageId="4" pageNumber="199" value="2003-06-04">4.VI.2003</date>
, S. Nagashima (ICMI). [Kuro-shima Is.] 1 Ƥ, Hori, 
<date id="FFB51035FFFB1D19FBB1FAA948BCF59F" box="[1067,1184,1401,1425]" pageId="4" pageNumber="199" value="2001-05-28">28.V.2001</date>
, T. Shimada (ICMI). [Iriomote-jima Is.] 7 3 2 Ƥ, Funaura, 
<date id="FFB51035FFFB1D19FDFEFA4D4EFAF5BB" box="[612,742,1437,1461]" pageId="4" pageNumber="199" value="1984-11-12">12.XI.1984</date>
, M. Tomokuni (NSMT); 2 Ƥ, Toyohara, 
<date id="FFB51035FFFB1D19FB49FA4D496DF5BB" box="[1235,1393,1437,1461]" pageId="4" pageNumber="199" value="1997-05-10" valueMax="1997-05-11" valueMin="1997-05-10">10–11.V.1997</date>
, TI (TUA); 
<date id="FFB51035FFFB1D19FF6EFA114D5AF5D7" box="[244,326,1473,1497]" pageId="4" pageNumber="199" value="1914-02-03">14 3 2</date>
Ƥ, Komi, 
<date id="FFB51035FFFB1D19FE5FFA114E5BF5D7" box="[453,583,1473,1497]" pageId="4" pageNumber="199" value="1998-06-21">21.VI.1998</date>
, light trap, TI (TUA); 1 3, Shirahama-rindo, 
<date id="FFB51035FFFB1D19FBCDFA1148C5F5D7" box="[1111,1241,1473,1497]" pageId="4" pageNumber="199" value="1998-11-21">21.XI.1998</date>
, TI (TUA); 1 Ƥ, Shirahama-rindo, 
<date id="FFB51035FFFB1D19FEF2FA354DC1F5F3" box="[360,477,1509,1533]" pageId="4" pageNumber="199" value="2000-05-30">30.V.2000</date>
, TI (TUA); 1 3, Shirahama, 
<date id="FFB51035FFFB1D19FCD7FA354FD3F5F3" box="[845,975,1509,1533]" pageId="4" pageNumber="199" value="1998-11-23">23.XI.1998</date>
, T. Shimada (TUA); 1 Ƥ, Shirahama, 
<date id="FFB51035FFFB1D19FF0DF9D94D05F62F" box="[151,281,1545,1569]" pageId="4" pageNumber="199" value="2003-06-19">19.VI.2003</date>
, T. Tsuru (TUA); 1 3 2 Ƥ, Shirahama, 
<date id="FFB51035FFFB1D19FD42F9D94F51F62F" box="[728,845,1545,1569]" pageId="4" pageNumber="199" value="2008-05-12">12.V.2008</date>
, TB (TUA); 2 3, Otomi-rindo, 
<date id="FFB51035FFFB1D19FB2EF9D9492FF62F" box="[1204,1331,1545,1569]" pageId="4" pageNumber="199" value="2004-04-24">24.IV.2004</date>
, T. Tsuru (TUA); 2 3 4 Ƥ, Otomi, 
<date id="FFB51035FFFB1D19FE29F9FD4E3BF64B" box="[435,551,1581,1605]" pageId="4" pageNumber="199" value="2007-05-19">19.V.2007</date>
, TB (TUA). [Yonaguni-jima Is.] 4 Ƥ, Mt. Urabu-dake, 
<date id="FFB51035FFFB1D19FB02F9FD4906F64B" box="[1176,1306,1581,1605]" pageId="4" pageNumber="199" value="1998-11-16">16.XI.1998</date>
, TI (TUA); 1 Ƥ, Mt. Urabu-dake, 
<date id="FFB51035FFFB1D19FE0AF9814E1FF667" box="[400,515,1617,1641]" pageId="4" pageNumber="199" value="2004-09-09">9.IX.2004</date>
, S. Arai (TUA); 4 Ƥ, Tarumai, 
<date id="FFB51035FFFB1D19FCF9F9814FCBF667" box="[867,983,1617,1641]" pageId="4" pageNumber="199" value="2005-06-09">9.VI.2005</date>
, light trap, TI (TUA).
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D19FF5DF9A54893F7A3" pageId="4" pageNumber="199" type="diagnosis">
<paragraph id="8BB436F5FFFB1D19FF5DF9A54893F7A3" blockId="4.[151,1437,1149,2038]" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DF9A54D23F680" bold="true" box="[199,319,1653,1678]" pageId="4" pageNumber="199">Diagnosis.</emphasis>
Recognized among congeners of 
<emphasis id="B97FEAE7FFFB1D19FD57F9A54F24F683" box="[717,824,1653,1677]" italics="true" pageId="4" pageNumber="199">Sadoletus</emphasis>
by a combination of the following characters: head, anterior lobe of pronotum, and basal half of scutellum blackish (
<figureCitation id="13302A70FFFB1D19FCC9F9494FA6F6BF" box="[851,954,1689,1713]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2, 3</figureCitation>
); male antenna brownish yellow in segment I and basal half of segment II and blackish in remaining parts (
<figureCitation id="13302A70FFFB1D19FCCFF96D4F85F6DB" box="[853,921,1725,1749]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Fig. 2</figureCitation>
); posterior lobe of pronotum yellowish brown with pale stripe along meson and often with 4 dark markings arranged transversely near posterior margin (
<figureCitation id="13302A70FFFB1D19FABEF9314990F6F7" box="[1316,1420,1761,1785]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2, 3</figureCitation>
); scutellum with pale stripe along meson in apical half (
<figureCitation id="13302A70FFFB1D19FD74F8D54F44F713" box="[750,856,1797,1821]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2, 3</figureCitation>
); hemelytra well surpassing apex of abdomen (
<figureCitation id="13302A70FFFB1D19FAFFF8D54CDEF74F" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2, 3</figureCitation>
); corium blackish in apical one-third to apical half in male (
<figureCitation id="13302A70FFFB1D19FCC6F8F94FBCF74F" box="[860,928,1833,1857]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Fig. 2</figureCitation>
); tibiae with blackish annulations (
<figureCitation id="13302A70FFFB1D19FAB9F8F94990F74F" box="[1315,1420,1833,1857]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Figs. 2, 3</figureCitation>
); laterotergites concolor (
<figureCitation id="13302A70FFFB1D19FE06F89D4DF2F76B" box="[412,494,1869,1893]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="4" pageNumber="199">Fig. 15</figureCitation>
); pygophore horizontally elliptical in dorsal view (
<figureCitation id="13302A70FFFB1D19FB84F89D486CF76B" box="[1054,1136,1869,1893]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="4" pageNumber="199">Fig. 16</figureCitation>
); paramere narrow, twisted at basal one-third, expanded at apical one-third (
<figureCitation id="13302A70FFFB1D19FD2DF8A14F16F787" box="[695,778,1905,1929]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="4" pageNumber="199">Fig. 17</figureCitation>
); and female abdominal sternite VII with posterior margin roundly projected at both sides of midpoint (cleft on midline). Body length 
<quantity id="4CF39B10FFFB1D19FC49F8454897F7A3" box="[979,1163,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.25" metricValueMin="3.65" pageId="4" pageNumber="199" unit="mm" value="4.45" valueMax="5.25" valueMin="3.65">3.65 to 5.25 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFFB1D1BFF5DF8694F35F3AA" lastPageId="6" lastPageNumber="201" pageId="4" pageNumber="199" type="description">
<paragraph id="8BB436F5FFFB1D18FF5DF8694D8AF1FA" blockId="4.[151,1437,1149,2038]" lastBlockId="5.[151,1437,151,500]" lastPageId="5" lastPageNumber="200" pageId="4" pageNumber="199">
<emphasis id="B97FEAE7FFFB1D19FF5DF8694D45F7DC" bold="true" box="[199,345,1977,2002]" pageId="4" pageNumber="199">Description.</emphasis>
Male (
<typeStatus id="54B08857FFFB1D19FE22F8694E3DF7DF" box="[440,545,1977,2001]" pageId="4" pageNumber="199" type="holotype">holotype</typeStatus>
): Venter almost blackish. Setae on body silvery. Punctures on pronotum, scutellum, and coria dark brown. Head (
<figureCitation id="13302A70FFFB1D19FDF5F80D4EAAF7FB" box="[623,694,2013,2037]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Fig. 2</figureCitation>
) blackish; clypeus brownish yellow. Antenna (
<figureCitation id="13302A70FFFB1D19FB40F80D493DF7FB" box="[1242,1313,2013,2037]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="4" pageNumber="199">Fig. 2</figureCitation>
) brownish yellow in segment I and basal half of segment II and blackish in remaining parts. Rostrum pale brown, with segment IV dark brown apically. Pronotum (
<figureCitation id="13302A70FFFA1D18FD1DFF6C4ED6F0DA" box="[647,714,188,212]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) blackish in anterior lobe and yellowish brown in posterior lobe, with pale stripe along meson on posterior lobe and 4 dark markings arranged transversely near posterior margin. Area above acetabula and posterior margin of metepisternum brownish. Evaporatrium dark gray; peritreme brownish yellow. Scutellum (
<figureCitation id="13302A70FFFA1D18FE44FEF74E3CF131" box="[478,544,295,319]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) blackish in basal half and brownish in apical half, with pale stripe along meson in apical half. Corium (
<figureCitation id="13302A70FFFA1D18FE22FE9C4E1EF16A" box="[440,514,332,356]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) whitish in basal two-thirds and blackish in remaining part; clavus whitish; membrane hyaline. Coxae brownish, except apical margin pale yellow; trochanter pale yellow; femora and tibiae brownish yellow, with dark brown annulations; tarsomere I pale yellow, except apical margin dark brown; tarsomeres II and III dark brown. Abdominal sternites VI and VII dark brown at middle; laterotergites (
<figureCitation id="13302A70FFFA1D18FADAFE674988F1C1" box="[1344,1428,439,463]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="5" pageNumber="200">Fig. 15</figureCitation>
) uniformly dark brown.
</paragraph>
<caption id="DF74667DFFFA1D18FF0DFA264D26F666" httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="5" pageNumber="200" targetBox="[175,1399,573,1474]" targetPageId="5">
<paragraph id="8BB436F5FFFA1D18FF0DFA264D26F666" blockId="5.[151,1436,1526,1640]" pageId="5" pageNumber="200">
<emphasis id="B97FEAE7FFFA1D18FF0DFA264D4AF605" bold="true" box="[151,342,1526,1549]" pageId="5" pageNumber="200">FIGURES 15–22.</emphasis>
<taxonomicName id="4C0B4D76FFFA1D18FEC7FA264E23F602" box="[349,575,1526,1548]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="200" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFFA1D18FEC7FA264E23F602" box="[349,575,1526,1548]" italics="true" pageId="5" pageNumber="200">Sadoletus ryukyuensis</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7FFFA1D18FDDDFA274E85F602" bold="true" box="[583,665,1527,1548]" pageId="5" pageNumber="200">
<taxonomicNameLabel id="A24C579CFFFA1D18FDDDFA274E85F602" box="[583,665,1527,1548]" pageId="5" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18–19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20–21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16–22.
</paragraph>
</caption>
<paragraph id="8BB436F5FFFA1D18FF5DF95F4937F739" blockId="5.[151,1437,1679,2027]" pageId="5" pageNumber="200">
Body (
<figureCitation id="13302A70FFFA1D18FE8EF95F4D45F6A9" box="[276,345,1679,1703]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) 3.3 times as long as width across humeri, covered with pubescence. Head (
<figureCitation id="13302A70FFFA1D18FB25F95F4918F6A9" box="[1215,1284,1679,1703]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) 1.4 times as wide as its length, covered with both rough and fine punctures; tylus tumid upwards; clypeus weakly projecting anteriad; juga with decumbent setae. Eyes large, occupying nearly half area of head in lateral view, not reaching anterior margin of pronotum. Antennae covered with suberect setae; proportional lengths of segments I to IV 1.0: 2.3: 1.8: 2.2. Rostrum attaining to mesocoxae; proportional lengths of segments I to IV 1.6: 1.8: 1.3: 1.0.
</paragraph>
<paragraph id="8BB436F5FFFA1D1BFF5DF8944FD4F0F9" blockId="5.[151,1437,1679,2027]" lastBlockId="6.[151,1437,151,1724]" lastPageId="6" lastPageNumber="201" pageId="5" pageNumber="200">
Pronotum (
<figureCitation id="13302A70FFFA1D18FEDCF8934D97F755" box="[326,395,1859,1883]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) a little more than 1.4 time as wide as its length, nearly trapezoidal, lacking collar, densely covered with punctures, and with lateral margin of anterior lobe weakly serrate; posterior lobe 1.25 times as long as anterior lobe, a little more convex than anterior lobe; posterior margin almost straight. Venter of prothorax, mesepisternum and metepisternum covered with rough punctures. Scutellum (
<figureCitation id="13302A70FFFA1D18FBBFF87F4870F7C9" box="[1061,1132,1967,1991]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) triangular, weakly tumid transversely in basal half, entirely covered with punctures. Hemelytra (
<figureCitation id="13302A70FFFA1D18FC2AF8034FEFF7E5" box="[944,1011,2003,2027]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="5" pageNumber="200">Fig. 2</figureCitation>
) exceeding apex of abdomen, lacking punctures from outside along vein R+M; apical margin of corium as long as claval suture. Profemur moderately expanded, armed with 2 spines ventrally at about apical one-third; one of these spines large and another small; tibiae covered with long setae, becoming gradually thicker in apical half.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFED44DE7F131" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
Abdomen (
<figureCitation id="13302A70FFF91D1BFED3FED44D8DF112" box="[329,401,260,284]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Fig. 2</figureCitation>
) as wide as hemelytra; venter covered with fine punctures, except sternites VI and VII glabrous and polished medially.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFE9C4DA6F219" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
Genital structures of 
<typeStatus id="54B08857FFF91D1BFE2DFE9D4E3BF16B" box="[439,551,333,357]" pageId="6" pageNumber="201" type="paratype">paratypes</typeStatus>
: Pygophore (
<figureCitation id="13302A70FFF91D1BFD27FE9C4F0CF16A" box="[701,784,332,356]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="6" pageNumber="201">Fig. 16</figureCitation>
) horizontally elliptical in dorsal view, covered with erect and suberect setae; dorsal aperture about one-third as wide at middle as pygophore, lacking collar. Paramere (
<figureCitation id="13302A70FFF91D1BFAF4FEBF4CA8F1A2" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="6" pageNumber="201">Fig. 17</figureCitation>
) narrow, twisted at basal one-third, widened at apical one-third, spatula-shaped in apical part, covered with a few setae. Phallosoma (
<figureCitation id="13302A70FFF91D1BFE3EFE674E31F1C1" box="[420,557,439,463]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="6" pageNumber="201">Figs. 18, 19</figureCitation>
) 1.5 times as long as its width in ventral view, roundly concave along apical margin; endosoma (
<figureCitation id="13302A70FFF91D1BFE1AFE0C4E0EF1FA" box="[384,530,476,500]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="6" pageNumber="201">Figs. 20, 21</figureCitation>
) with 2 areas of sclerotized processes; basal one of these areas with larger processes than apical one.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFDF44E17F2C2" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
Females (
<typeStatus id="54B08857FFF91D1BFEACFDF44DB5F232" box="[310,425,548,572]" pageId="6" pageNumber="201" type="paratype">paratypes</typeStatus>
): resembles male in general appearance (
<figureCitation id="13302A70FFF91D1BFC1FFDF44FD0F232" box="[901,972,548,572]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Fig. 3</figureCitation>
). Body evidently wider; antennae paler; coria entirely whitish to brownish yellow (
<figureCitation id="13302A70FFF91D1BFD1FFD974ED7F251" box="[645,715,583,607]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Fig. 3</figureCitation>
); annulations on femora paler; abdomen obviously wider than hemelytra (
<figureCitation id="13302A70FFF91D1BFE8DFDBC4D47F28A" box="[279,347,620,644]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Fig. 3</figureCitation>
); abdominal sternite VII with posterior margin roundly projected at both sides of midpoint (cleft on midline) (
<figureCitation id="13302A70FFF91D1BFEBDFD5F4D76F2A9" box="[295,362,655,679]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="6" pageNumber="201">Fig. 6</figureCitation>
); ovipositor 
<quantity id="4CF39B10FFF91D1BFE62FD5F4EB3F2A9" box="[504,687,655,679]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="6" pageNumber="201" unit="mm" value="0.65" valueMax="0.75" valueMin="0.55">0.55 to 0.75 mm</quantity>
in length, nearly reaching sternite VI (
<figureCitation id="13302A70FFF91D1BFBC6FD5F48BCF2A9" box="[1116,1184,655,679]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="6" pageNumber="201">Fig. 6</figureCitation>
); spermatheca narrow, turned upwards at apex (
<figureCitation id="13302A70FFF91D1BFE30FD644DE3F2C2" box="[426,511,692,716]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 – 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 – 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 – 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="6" pageNumber="201">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFD074D63F31A" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
Variations: body length varies from 
<quantity id="4CF39B10FFF91D1BFDC7FD074F05F2E1" box="[605,793,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.25" metricValueMin="3.65" pageId="6" pageNumber="201" unit="mm" value="4.45" valueMax="5.25" valueMin="3.65">3.65 to 5.25 mm</quantity>
. Blackish parts of the pronotum, scutellum, and coria are also variable in area.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFCF04F35F3AA" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF5DFCF04D6AF336" bold="true" box="[199,374,800,824]" pageId="6" pageNumber="201">Measurements</emphasis>
(
<typeStatus id="54B08857FFF91D1BFE10FCCF4DF3F339" box="[394,495,799,823]" pageId="6" pageNumber="201" type="holotype">holotype</typeStatus>
). Body length 4.76 (
<typeStatus id="54B08857FFF91D1BFD71FCCF4F44F339" box="[747,856,799,823]" pageId="6" pageNumber="201" type="paratype">paratypes</typeStatus>
3.65–5.25), width 1.43; head length 0.80, width across eyes 1.13; lengths of antennal segments I, II, III and IV 0.31, 0.70, 0.56 and 0.68, respectively; lengths of rostral segments I, II, III and IV 0.63, 0.71, 0.50 and 0.40, respectively; pronotum length 1.00, width 1.43; hemelytron length 3.06; scutellum length 0.80, width 0.78.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF91D1BFF5DFC7F4865F3E2" pageId="6" pageNumber="201" type="distribution">
<paragraph id="8BB436F5FFF91D1BFF5DFC7F4865F3E2" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF5DFC7F4D7CF3C6" bold="true" box="[199,352,943,968]" pageId="6" pageNumber="201">Distribution.</emphasis>
<collectingCountry id="F31C7665FFF91D1BFEF3FC614DB1F3C6" box="[361,429,945,968]" name="Japan" pageId="6" pageNumber="201">Japan</collectingCountry>
: the Ryukyus (Yaku-shima Is., Nakano-shima Is., Tokuno-shima Is., Okinawa-honto Is., Miyako-jima Is., Ishigaki-jima Is., Kuro-shima Is., Iriomote-jima Is., Yonaguni-jima Is.).
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF91D1BFF5DFC284E51F43A" pageId="6" pageNumber="201" type="etymology">
<paragraph id="8BB436F5FFF91D1BFF5DFC284E51F43A" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF5DFC284D52F41E" bold="true" box="[199,334,1016,1040]" pageId="6" pageNumber="201">Etymology.</emphasis>
Named after the Ryukyus, an archipelago lying between Kyushu and 
<collectingCountry id="F31C7665FFF91D1BFBE2FC2748D1F401" box="[1144,1229,1015,1039]" name="Taiwan" pageId="6" pageNumber="201">Taiwan</collectingCountry>
, the distributional range of this new species; an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF91D1BFF5DFB904E68F6B2" pageId="6" pageNumber="201" type="discussion">
<paragraph id="8BB436F5FFF91D1BFF5DFB9048C1F53E" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF5DFB904D27F456" bold="true" box="[199,315,1088,1112]" pageId="6" pageNumber="201">Remarks.</emphasis>
This new species is most similar in general appearance to 
<taxonomicName id="4C0B4D76FFF91D1BFC47FB904931F456" authority="Bergroth. However" authorityName="Bergroth. However" box="[989,1325,1087,1112]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFC47FB904856F456" box="[989,1098,1088,1112]" italics="true" pageId="6" pageNumber="201">S. valdezi</emphasis>
Bergroth. However
</taxonomicName>
, it can be distinguished from the latter by a combination of the following characters: head and antennae darker, laterotergites uniformly dark brown, apical one-third of corium blackish in male, and posterior margin of female abdominal sternite VII roundly projected at both sides of midpoint. Male of this new species resembles in general habitus 
<emphasis id="B97FEAE7FFF91D1BFA1DFB7D4CFEF4E6" italics="true" pageId="6" pageNumber="201">S. corvus</emphasis>
Distant, 
<emphasis id="B97FEAE7FFF91D1BFED6FB004DAFF4E6" box="[332,435,1232,1256]" italics="true" pageId="6" pageNumber="201">S. barkei</emphasis>
Bergroth, and 
<emphasis id="B97FEAE7FFF91D1BFDFBFB004EE3F4E6" box="[609,767,1232,1256]" italics="true" pageId="6" pageNumber="201">S. voluptarius</emphasis>
Bergroth, but it can be distinguished from the latter three species by the antennae being brownish yellow in segment I and basal half of segment II and blackish apically, the brownish yellow femora with dark brown annulations, and the smaller blackish areas of the coria.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DFAEC4F31F65E" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
Hidaka (1959) first reported this species as 
<taxonomicName id="4C0B4D76FFF91D1BFD34FAED4F98F55A" authority="Bergroth" authorityName="Bergroth" box="[686,900,1340,1364]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFD34FAED4EA0F55A" box="[686,700,1341,1364]" italics="true" pageId="6" pageNumber="201">S</emphasis>
. 
<emphasis id="B97FEAE7FFF91D1BFD51FAEC4F06F55A" box="[715,794,1340,1364]" italics="true" pageId="6" pageNumber="201">valdezi</emphasis>
Bergroth
</taxonomicName>
from 
<collectingCountry id="F31C7665FFF91D1BFC52FAED4814F55A" box="[968,1032,1341,1364]" name="Japan" pageId="6" pageNumber="201">Japan</collectingCountry>
on the basis of four specimens (one male and two females from Nakano-shima Island and one male from Ishigaki-jima Island). Of these specimens, one female (Nakano-shima Is.) examined in this present study and another specimen shown in Hidaka’s paper (1959, 
<figureCitation id="13302A70FFF91D1BFF7CFA784D34F5CE" box="[230,296,1448,1472]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 – 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Fig. 2</figureCitation>
) (apparently male; locality not indicated) belongs to 
<taxonomicName id="4C0B4D76FFF91D1BFCE0FA784806F5CE" box="[890,1050,1448,1472]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFF91D1BFCE0FA784F94F5B1" box="[890,904,1448,1471]" italics="true" pageId="6" pageNumber="201">S</emphasis>
. 
<emphasis id="B97FEAE7FFF91D1BFC0CFA784806F5CE" box="[918,1050,1448,1472]" italics="true" pageId="6" pageNumber="201">ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C579CFFF91D1BFBBBFA784866F5CE" box="[1057,1146,1448,1472]" pageId="6" pageNumber="201" rank="species">
<emphasis id="B97FEAE7FFF91D1BFBBBFA784866F5CE" bold="true" box="[1057,1146,1448,1472]" pageId="6" pageNumber="201">sp. nov.</emphasis>
</taxonomicNameLabel>
without doubt. Although, unfortunately, the other two specimens (one male and one female) were not located by us, they are strongly supposed to be identical with this new species because the male has definite coloration in the hemelytron which is much different from that of 
<taxonomicName id="4C0B4D76FFF91D1BFE4AF9C54E27F622" box="[464,571,1556,1580]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFE4AF9C54DC2F622" box="[464,478,1557,1580]" italics="true" pageId="6" pageNumber="201">S</emphasis>
. 
<emphasis id="B97FEAE7FFF91D1BFE76F9C44E27F622" box="[492,571,1556,1580]" italics="true" pageId="6" pageNumber="201">valdezi</emphasis>
</taxonomicName>
, and all of the females were collected from the same locality (Nakano-shima Is.), which is far from the distributional range of 
<taxonomicName id="4C0B4D76FFF91D1BFD21F9E84F3AF65E" box="[699,806,1592,1616]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFD21F9E84ED5F641" box="[699,713,1592,1615]" italics="true" pageId="6" pageNumber="201">S</emphasis>
. 
<emphasis id="B97FEAE7FFF91D1BFD4DF9E84F3AF65E" box="[727,806,1592,1616]" italics="true" pageId="6" pageNumber="201">valdezi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF5DF98B4E68F6B2" blockId="6.[151,1437,151,1724]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF5DF98B4D91F67A" bold="true" box="[199,397,1627,1652]" pageId="6" pageNumber="201">Biological notes.</emphasis>
A number of adults of 
<taxonomicName id="4C0B4D76FFF91D1BFD3CF98D4F56F67A" box="[678,842,1628,1652]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFF91D1BFD3CF98D4EA8F67A" box="[678,692,1629,1652]" italics="true" pageId="6" pageNumber="201">S</emphasis>
. 
<emphasis id="B97FEAE7FFF91D1BFD5CF98C4F56F67A" box="[710,842,1628,1652]" italics="true" pageId="6" pageNumber="201">ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C579CFFF91D1BFCCEF98C4FAEF67A" box="[852,946,1628,1652]" pageId="6" pageNumber="201" rank="species">
<emphasis id="B97FEAE7FFF91D1BFCCEF98C4FAEF67A" bold="true" box="[852,946,1628,1652]" pageId="6" pageNumber="201">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected from trees of 
<taxonomicName id="4C0B4D76FFF91D1BFA99F98D4DFFF696" authority="Reinw. ex Blume" authorityName="Reinw. ex Blume" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="201" phylum="Mollusca" rank="species" species="virgata">
<emphasis id="B97FEAE7FFF91D1BFA99F98D4980F67A" box="[1283,1436,1629,1652]" italics="true" pageId="6" pageNumber="201">Ficus virgata</emphasis>
Reinw. ex Blume (Moraceae)
</taxonomicName>
bearing a number of fruits, which may be a host plant of this new species. The new species is often attracted by artificial light.
</paragraph>
</subSubSection>
</treatment>
</document>