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<document id="57E6D557E3B1EBA917403CB813CBE0E1" ID-DOI="10.11646/zootaxa.3609.2.5" ID-GBIF-Dataset="fad27fbd-389c-4c60-a927-77ac88afa267" ID-ISSN="1175-5326" ID-Zenodo-Dep="220831" ID-ZooBank="0B91082F-CCA0-4CE0-B221-AE62151CB985" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460571845318" checkinUser="plazi" docAuthor="Ban, Teruaki &amp; Ishikawa, Tadashi" docDate="2013" docId="03A287E3FFF91D15FF0DF8DF48F4F2E1" docLanguage="en" docName="zt03609p203.pdf" docOrigin="Zootaxa 3609 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sadoletus valdezi Bergroth" docType="treatment" docVersion="6" lastPageNumber="203" masterDocId="FF9BFF9BFFFF1D1DFF9AFFD04C1CF00E" masterDocTitle="The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan" masterLastPageNumber="203" masterPageNumber="195" pageNumber="201" updateTime="1698322288420" updateUser="plazi">
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<mods:title id="0C2D6816768D16CF6210C9CAD733D1EE">The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan</mods:title>
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<mods:name id="8F8A3189B79266B935342D1D3D36A3E5" type="personal">
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<mods:namePart id="A1FF0B7C12F82A3AC6245B96CEB485C1">Ban, Teruaki</mods:namePart>
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<mods:namePart id="643B5D424A97489B89FCD0FA5152F3AA">Ishikawa, Tadashi</mods:namePart>
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<treatment id="03A287E3FFF91D15FF0DF8DF48F4F2E1" ID-DOI="http://doi.org/10.5281/zenodo.6151273" ID-GBIF-Taxon="119558919" ID-Zenodo-Dep="6151273" LSID="urn:lsid:plazi:treatment:03A287E3FFF91D15FF0DF8DF48F4F2E1" httpUri="http://treatment.plazi.org/id/03A287E3FFF91D15FF0DF8DF48F4F2E1" lastPageId="8" lastPageNumber="203" pageId="6" pageNumber="201">
<subSubSection id="C311657EFFF91D1BFF0DF8DF4FB5F79E" pageId="6" pageNumber="201" type="nomenclature">
<paragraph id="8BB436F5FFF91D1BFF0DF8DF4DC3F727" blockId="6.[151,479,1807,1866]" box="[151,479,1807,1833]" pageId="6" pageNumber="201">
<heading id="D0FC8199FFF91D1BFF0DF8DF4DC3F727" bold="true" box="[151,479,1807,1833]" fontSize="11" level="1" pageId="6" pageNumber="201" reason="1">
<taxonomicName id="4C0B4D76FFF91D1BFF0DF8DF4DC3F727" authority="Bergroth" authorityName="Bergroth" box="[151,479,1807,1833]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFF0DF8DF4DC3F727" bold="true" box="[151,479,1807,1833]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF0DF8DF4D7AF727" bold="true" box="[151,358,1807,1833]" italics="true" pageId="6" pageNumber="201">Sadoletus valdezi</emphasis>
Bergroth
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF0DF8E24D7AF744" blockId="6.[151,479,1807,1866]" box="[151,358,1842,1866]" pageId="6" pageNumber="201">
(
<figureCitation id="13302A70FFF91D1BFF05F8E24CF3F744" box="[159,239,1842,1866]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="6" pageNumber="201">Figs. 4</figureCitation>
,
<figureCitation id="13302A70FFF91D1BFF60F8E34D16F744" box="[250,266,1843,1866]" captionStart="FIGURES 5 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="6" pageNumber="201">7</figureCitation>
,
<figureCitation id="13302A70FFF91D1BFE8CF8E24D42F744" box="[278,350,1842,1866]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="6" pageNumber="201">2330</figureCitation>
)
</paragraph>
<paragraph id="8BB436F5FFF91D1BFF0DF8AA4FB5F79E" blockId="6.[151,937,1914,1936]" box="[151,937,1914,1936]" pageId="6" pageNumber="201">
<taxonomicName id="4C0B4D76FFF91D1BFF0DF8AA4E01F79E" authority="Bergroth, 1918: 107" authorityName="Bergroth" authorityPageNumber="107" authorityYear="1918" box="[151,541,1914,1936]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF91D1BFF0DF8AA4D54F79E" box="[151,328,1914,1936]" italics="true" pageId="6" pageNumber="201">Sadoletus valdezi</emphasis>
Bergroth, 1918: 107
</taxonomicName>
; Slater, 1964: 776; Péricart, 2001: 105.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF91D1AFF0DF8664D08F1C1" lastPageId="7" lastPageNumber="202" pageId="6" pageNumber="201" type="materials_examined">
<paragraph id="8BB436F5FFF91D1BFF0DF8664980F7FC" blockId="6.[151,1436,1974,2035]" pageId="6" pageNumber="201">
<emphasis id="B97FEAE7FFF91D1BFF0DF8664D9CF7C1" bold="true" box="[151,384,1974,1999]" pageId="6" pageNumber="201">Material examined.</emphasis>
Non-types:
<date id="FFB51035FFF91D1BFD96F8664E45F7C0" box="[524,601,1974,1999]" pageId="6" pageNumber="201" value="1915-03-08">8 3 15</date>
Ƥ and
<specimenCount id="9D0DFD7CFFF91D1BFD36F8664F0FF7C0" box="[684,787,1974,1998]" pageId="6" pageNumber="201" type="adult">27 adults</specimenCount>
(ICMI, NSMT, TUA) —
<emphasis id="B97FEAE7FFF91D1BFBA2F86748D1F7C1" bold="true" box="[1080,1229,1975,1999]" pageId="6" pageNumber="201">the Ryukyus</emphasis>
[Miyako-jima Is.] 1 3, Higashi-sokobaru, Hirara,
<date id="FFB51035FFF91D1BFD8BF80A4E96F7FC" box="[529,650,2010,2034]" pageId="6" pageNumber="201" value="1999-10-30">30.X.1999</date>
, light trap, TI. [Ishigaki-jima Is.]
<specimenCount id="9D0DFD7CFFF91D1BFBBDF80A4894F7FC" box="[1063,1160,2010,2034]" pageId="6" pageNumber="201" type="adult">4 adults</specimenCount>
, Takeda,
<date id="FFB51035FFF91D1BFB67F80A4968F7FC" box="[1277,1396,2010,2034]" pageId="6" pageNumber="201" value="1991-02-27">27.II.1991</date>
, T.
</paragraph>
<paragraph id="8BB436F5FFF81D1AFF0DFF474D08F1C1" blockId="7.[151,1436,151,463]" pageId="7" pageNumber="202">
Kishimoto; 1 Ƥ, Tomino,
<date id="FFB51035FFF81D1AFE22FF474E27F0A1" box="[440,571,151,175]" pageId="7" pageNumber="202" value="2002-09-27">27.IX.2002</date>
, H. Yamamoto; 1 Ƥ, Mt. Omoto-dake,
<date id="FFB51035FFF81D1AFC6BFF47487DF0A1" box="[1009,1121,151,175]" pageId="7" pageNumber="202" value="1997-04-07">7.IV.1997</date>
, TI. [Kohama-jima Is.] 3 Ƥ, Ufudaki,
<date id="FFB51035FFF81D1AFE9BFF6C4D6BF0DA" box="[257,375,188,212]" pageId="7" pageNumber="202" value="2007-05-14">14.V.2007</date>
, light trap, TB; 1 Ƥ, Ufudaki,
<date id="FFB51035FFF81D1AFD55FF6C4F58F0DA" box="[719,836,188,212]" pageId="7" pageNumber="202" value="2007-05-14">14.V.2007</date>
, light trap, S. Fujinuma. [Iriomote-jima Is.] 1 3 3 Ƥ, Ootomi,
<date id="FFB51035FFF81D1AFF66FF0F4D6DF0F9" box="[252,369,223,247]" pageId="7" pageNumber="202" value="2007-05-19">19.V.2007</date>
, light trap, TB;
<specimenCount id="9D0DFD7CFFF81D1AFDB3FF304E9AF0F9" box="[553,646,223,247]" pageId="7" pageNumber="202" type="adult">7 adults</specimenCount>
, Komi,
<date id="FFB51035FFF81D1AFD78FF0F4F78F0F9" box="[738,868,223,247]" pageId="7" pageNumber="202" value="1998-06-21">21.VI.1998</date>
, light trap, TI; 1 3 1 Ƥ, Funaura,
<date id="FFB51035FFF81D1AFB74FF0F4973F0F9" box="[1262,1391,223,247]" pageId="7" pageNumber="202" value="1984-11-11">11.XI.1984</date>
, M. Tomokuni;
<specimenCount id="9D0DFD7CFFF81D1AFE80FED44D71F112" box="[282,365,260,284]" pageId="7" pageNumber="202" type="adult">1 adult</specimenCount>
, Urauchi,
<date id="FFB51035FFF81D1AFE7FFED44E46F112" box="[485,602,260,284]" pageId="7" pageNumber="202" value="1999-05-29">29.V.1999</date>
, light trap, H. Mizushima; 2 3 1 Ƥ, Shirahama,
<date id="FFB51035FFF81D1AFB14FED4491FF112" box="[1166,1283,260,284]" pageId="7" pageNumber="202" value="2008-05-12">12.V.2008</date>
, TB;
<specimenCount id="9D0DFD7CFFF81D1AFADFFED44984F112" box="[1349,1432,260,284]" pageId="7" pageNumber="202" type="adult">1 adult</specimenCount>
, Shirahama,
<date id="FFB51035FFF81D1AFEB9FEF74D84F131" box="[291,408,295,319]" pageId="7" pageNumber="202" value="2008-05-10">10.V.2008</date>
, TB;
<specimenCount id="9D0DFD7CFFF81D1AFE79FEF74E59F131" box="[483,581,295,319]" pageId="7" pageNumber="202" type="adult">3 adults</specimenCount>
, Shirahama,
<date id="FFB51035FFF81D1AFD78FEF74F44F131" box="[738,856,295,319]" pageId="7" pageNumber="202" value="2003-05-29">29.V.2003</date>
, S. Nagashima; 1 3, Shirahama,
<date id="FFB51035FFF81D1AFB68FEF74968F131" box="[1266,1396,295,319]" pageId="7" pageNumber="202" value="1998-11-23">23.XI.1998</date>
, T. Shimada;
<specimenCount id="9D0DFD7CFFF81D1AFE95FE9D4D6DF16A" box="[271,369,332,356]" pageId="7" pageNumber="202" type="adult">7 adults</specimenCount>
, Shirahama-rindo,
<date id="FFB51035FFF81D1AFDCEFE9C4ECBF16A" box="[596,727,332,356]" pageId="7" pageNumber="202" value="1998-11-21">21.XI.1998</date>
, light trap, TI; 1 3 1 Ƥ, Shirahama-rindo,
<date id="FFB51035FFF81D1AFB68FE9C497BF16A" box="[1266,1383,332,356]" pageId="7" pageNumber="202" value="2000-05-30">30.V.2000</date>
, TI. [Yonaguni-jima Is.]
<specimenCount id="9D0DFD7CFFF81D1AFEE2FEBF4DCEF189" box="[376,466,367,391]" pageId="7" pageNumber="202" type="adult">2 adults</specimenCount>
, Tarumai,
<date id="FFB51035FFF81D1AFDDDFEBF4EA0F189" box="[583,700,367,391]" pageId="7" pageNumber="202" value="2000-05-20">20.V.2000</date>
, light trap, H. Mizushima; 1 Ƥ, Tarumai,
<date id="FFB51035FFF81D1AFB12FEBF48E0F189" box="[1160,1276,367,391]" pageId="7" pageNumber="202" value="2005-06-09">9.VI.2005</date>
, light trap, TI;
<specimenCount id="9D0DFD7CFFF81D1AFF0DFE444CEDF1A2" box="[151,241,404,428]" pageId="7" pageNumber="202" type="adult">2 adults</specimenCount>
, Mt. Urabu-dake,
<date id="FFB51035FFF81D1AFE26FE444E27F1A2" box="[444,571,404,428]" pageId="7" pageNumber="202" value="1997-04-13">13.IV.1997</date>
, TI; 2 Ƥ, Mt. Urabu-dake,
<date id="FFB51035FFF81D1AFCFDFE444FF5F1A2" box="[871,1001,404,428]" pageId="7" pageNumber="202" value="1998-11-16">16.XI.1998</date>
, TI; 1 3, Mt. Urabu-dake,
<date id="FFB51035FFF81D1AFA8FFE44498BF1A2" box="[1301,1431,404,428]" pageId="7" pageNumber="202" value="2001-06-18">18.VI.2001</date>
, K. Toyoda.
</paragraph>
</subSubSection>
<caption id="DF74667DFFF81D1AFF0DF9B04988F6BD" httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="7" pageNumber="202" targetBox="[191,1407,541,1586]" targetPageId="7">
<paragraph id="8BB436F5FFF81D1AFF0DF9B04988F6BD" blockId="7.[151,1436,1632,1716]" pageId="7" pageNumber="202">
<emphasis id="B97FEAE7FFF81D1AFF0DF9B04D48F67B" bold="true" box="[151,340,1632,1655]" pageId="7" pageNumber="202">FIGURES 2330.</emphasis>
<taxonomicName id="4C0B4D76FFF81D1AFEC3F9B04E16F678" box="[345,522,1632,1654]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="202" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF81D1AFEC3F9B04E16F678" box="[345,522,1632,1654]" italics="true" pageId="7" pageNumber="202">Sadoletus valdezi</emphasis>
</taxonomicName>
. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 2627, phallus (endosoma not everted), ventral (26) and lateral (27) views; 2829, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 2430.
</paragraph>
</caption>
<subSubSection id="C311657EFFF81D1AFF5DF930481DF7FA" pageId="7" pageNumber="202" type="diagnosis">
<paragraph id="8BB436F5FFF81D1AFF5DF930481DF7FA" blockId="7.[151,1437,1760,2036]" pageId="7" pageNumber="202">
<emphasis id="B97FEAE7FFF81D1AFF5DF9304D5DF6F7" bold="true" box="[199,321,1760,1785]" pageId="7" pageNumber="202">Diagnosis.</emphasis>
This new species can be recognized among congeners of
<emphasis id="B97FEAE7FFF81D1AFB98F9304873F6F6" box="[1026,1135,1760,1784]" italics="true" pageId="7" pageNumber="202">Sadoletus</emphasis>
by a combination of the following characters: head, anterior lobe of pronotum, mesepisternum, metepisternum, and basal half of scutellum reddish brown to blackish (
<figureCitation id="13302A70FFF81D1AFE5CF8F84E10F74E" box="[454,524,1832,1856]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Fig. 4</figureCitation>
); antennae brownish yellow (
<figureCitation id="13302A70FFF81D1AFCCEF8F84F85F74E" box="[852,921,1832,1856]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Fig. 4</figureCitation>
); evaporatrium blackish; peritreme yellowish; scutellum weakly tumid in basal one-third (
<figureCitation id="13302A70FFF81D1AFD17F89C4EC8F76A" box="[653,724,1868,1892]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Fig. 4</figureCitation>
); corium whitish to brownish yellow, with dark small spot on apex (
<figureCitation id="13302A70FFF81D1AFF40F8A04D03F786" box="[218,287,1904,1928]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Fig. 4</figureCitation>
); tibiae with dark annulations (
<figureCitation id="13302A70FFF81D1AFDE3F8A04EA2F786" box="[633,702,1904,1928]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Fig. 4</figureCitation>
); each laterotergite pale brown anteriorly and blackish posteriorly (
<figureCitation id="13302A70FFF81D1AFF05F8444CEDF7A2" box="[159,241,1940,1964]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1965,1988]" captionTargetBox="[233,1367,201,1927]" captionTargetId="figure@2.[192,1397,171,1960]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Japanese Sadoletus. 1, S. izzardi, male; 2 3, S. ryukyuensis sp. nov., male (2) and female (3); 4, S. valdezi, male. Scales: 1.0 mm." httpUri="https://zenodo.org/record/220832/files/figure.png" pageId="7" pageNumber="202">Figs. 4</figureCitation>
,
<figureCitation id="13302A70FFF81D1AFE9AF8444D03F7A2" box="[256,287,1940,1964]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="7" pageNumber="202">23</figureCitation>
); paramere widened at apical one-third, with acute at apex (
<figureCitation id="13302A70FFF81D1AFC4CF8444830F7A2" box="[982,1068,1940,1964]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="7" pageNumber="202">Fig. 25</figureCitation>
); female abdominal sternite VII with posterior margin entirely concave toward midpoint (cleft on midline) (
<figureCitation id="13302A70FFF81D1AFC45F868483FF7DE" box="[991,1059,1976,2000]" captionStart="FIGURES 5 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="7" pageNumber="202">Fig. 7</figureCitation>
); and ovipositor less than 1.0 mm in length, not reaching sternite VI in rest (
<figureCitation id="13302A70FFF81D1AFDF6F80C4EADF7FA" box="[620,689,2012,2036]" captionStart="FIGURES 5 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="7" pageNumber="202">Fig. 7</figureCitation>
). Body length
<quantity id="4CF39B10FFF81D1AFCCDF80C4FE0F7FA" box="[855,1020,2012,2036]" metricMagnitude="-3" metricUnit="m" metricValue="4.875" metricValueMax="5.5" metricValueMin="4.25" pageId="7" pageNumber="202" unit="mm" value="4.875" valueMax="5.5" valueMin="4.25">4.255.50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF71D15FF5DFF474CE6F189" pageId="8" pageNumber="203" type="description">
<paragraph id="8BB436F5FFF71D15FF5DFF474817F131" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="203">
<emphasis id="B97FEAE7FFF71D15FF5DFF474DF4F0BE" bold="true" box="[199,488,151,176]" pageId="8" pageNumber="203">Description of genitalia.</emphasis>
Male: Pygophore (
<figureCitation id="13302A70FFF71D15FD5CFF474F05F0A1" box="[710,793,151,175]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="8" pageNumber="203">Fig. 24</figureCitation>
) nearly globous in dorsal view, covered with fine setae; dorsal aperture more than one-third as wide at middle as pygophore, with collar along anterior margin. Paramere (
<figureCitation id="13302A70FFF71D15FF05FF0F4CEFF0F9" box="[159,243,223,247]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="8" pageNumber="203">Fig. 25</figureCitation>
) twisted at basal one-third, widened at apical one-third, acute and curved at apex, covered with several erect setae in basal one-third and a few short setae apically. Phallosoma (
<figureCitation id="13302A70FFF71D15FC5EFED44856F112" box="[964,1098,260,284]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="8" pageNumber="203">Figs. 26, 27</figureCitation>
) 1.5 times as long as its width in ventral view. Endosoma (
<figureCitation id="13302A70FFF71D15FE4AFEF74E4BF131" box="[464,599,295,319]" captionStart="FIGURES 23 30" captionStartId="7.[151,264,1632,1655]" captionTargetBox="[191,1407,541,1586]" captionTargetId="figure@7.[151,1436,517,1611]" captionTargetPageId="7" captionText="FIGURES 23 30. Sadoletus valdezi. 23, Laterotergites, dorsal view; 24, pygophore, dorsal view (setae omitted from left half); 25, left paramere, dorsal view; 26 27, phallus (endosoma not everted), ventral (26) and lateral (27) views; 28 29, phallus (endosoma everted), ventral (28) and lateral (29) views; 30, spermatheca. Scales: 0.5 mm for Fig. 23, 0.1 mm for Figs. 24 30." httpUri="https://zenodo.org/record/220836/files/figure.png" pageId="8" pageNumber="203">Figs. 28, 29</figureCitation>
) covered with fine spines near middle.
</paragraph>
<paragraph id="8BB436F5FFF71D15FF5DFE9C4CE6F189" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="203">
Females: Ovipositor (
<figureCitation id="13302A70FFF71D15FE5EFE9C4E10F16A" box="[452,524,332,356]" captionStart="FIGURES 5 7" captionStartId="3.[151,264,611,634]" captionTargetBox="[177,1390,213,560]" captionTargetId="figure@3.[151,1436,193,590]" captionTargetPageId="3" captionText="FIGURES 5 7. Apical part of female abdomen, ventral view. 5, Sadoletus izzardi; 6, S. ryukyuensis sp. nov.; 7, S. valdezi. Scales: 0.5 mm. abbreviations: st 7, sternite VII." httpUri="https://zenodo.org/record/220833/files/figure.png" pageId="8" pageNumber="203">Fig. 7</figureCitation>
)
<quantity id="4CF39B10FFF71D15FDBAFE9C4EF4F16A" box="[544,744,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="8" pageNumber="203" unit="mm" value="0.8" valueMax="0.85" valueMin="0.75">0.75 to 0.85 mm</quantity>
in length, not reaching sternite VI. Spermatheca (
<figureCitation id="13302A70FFF71D15FAA7FE9C4988F16A" box="[1341,1428,332,356]" captionStart="FIGURES 15 22" captionStartId="5.[151,264,1526,1549]" captionTargetBox="[175,1399,573,1474]" captionTargetId="figure@5.[151,1436,553,1505]" captionTargetPageId="5" captionText="FIGURES 15 22. Sadoletus ryukyuensis sp. nov. 15, Laterotergites, dorsal view; 16, pygophore, dorsal view (setae omitted from left half); 17, left paramere, dorsal view; 18 19, phallus (endosoma not everted), ventral (18) and lateral (19) views; 20 21, phallus (endosoma everted), ventral (20) and lateral (21) views; 22, spermatheca. Scales: 0.5 mm for Fig. 15, 0.1 mm for Figs. 16 22." httpUri="https://zenodo.org/record/220835/files/figure.png" pageId="8" pageNumber="203">Fig. 22</figureCitation>
) rounded.
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF71D15FF5DFE434CE5F219" pageId="8" pageNumber="203" type="distribution">
<paragraph id="8BB436F5FFF71D15FF5DFE434CE5F219" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="203">
<emphasis id="B97FEAE7FFF71D15FF5DFE434D7CF1A2" bold="true" box="[199,352,403,428]" pageId="8" pageNumber="203">Distribution.</emphasis>
<collectingCountry id="F31C7665FFF71D15FEF3FE454DB1F1A2" box="[361,429,405,428]" name="Japan" pageId="8" pageNumber="203">Japan</collectingCountry>
: the Ryukyus (Miyako-jima, Ishigaki-jima, Kohama-jima, Iriomote-jima, Yonaguni-jima Islands); the
<collectingCountry id="F31C7665FFF71D15FEB4FE674DACF1C1" box="[302,432,439,463]" name="Philippines" pageId="8" pageNumber="203">Philippines</collectingCountry>
. Previous records as
<taxonomicName id="4C0B4D76FFF71D15FD3CFE684F09F1DE" box="[678,789,440,464]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="203" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF71D15FD3CFE684EA8F1C1" box="[678,692,440,463]" italics="true" pageId="8" pageNumber="203">S</emphasis>
.
<emphasis id="B97FEAE7FFF71D15FD5FFE684F09F1DE" box="[709,789,440,464]" italics="true" pageId="8" pageNumber="203">valdezi</emphasis>
</taxonomicName>
from Nakano-shima Island (Hidaka 1959) and Tokunoshima Islands (Tomokuni 1989) arguably correspond to
<taxonomicName id="4C0B4D76FFF71D15FC9DFE0D4FBBF1FA" box="[775,935,476,500]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="203" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFF71D15FC9DFE0D4F09F1FA" box="[775,789,477,500]" italics="true" pageId="8" pageNumber="203">S</emphasis>
.
<emphasis id="B97FEAE7FFF71D15FCB9FE0C4FBBF1FA" box="[803,935,476,500]" italics="true" pageId="8" pageNumber="203">ryukyuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C579CFFF71D15FC37FE0C481AF1FA" box="[941,1030,476,500]" pageId="8" pageNumber="203" rank="species">
<emphasis id="B97FEAE7FFF71D15FC37FE0C481AF1FA" bold="true" box="[941,1030,476,500]" pageId="8" pageNumber="203">sp. nov.</emphasis>
</taxonomicNameLabel>
(see also “Remarks” in the previous species).
</paragraph>
</subSubSection>
<subSubSection id="C311657EFFF71D15FF5DFDF448F4F2E1" pageId="8" pageNumber="203" type="discussion">
<paragraph id="8BB436F5FFF71D15FF5DFDF44F76F2A9" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="203">
<emphasis id="B97FEAE7FFF71D15FF5DFDF44D27F232" bold="true" box="[199,315,548,572]" pageId="8" pageNumber="203">Remarks.</emphasis>
This species is similar in general appearance to
<taxonomicName id="4C0B4D76FFF71D15FC1BFDF5483AF232" box="[897,1062,548,572]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="203" phylum="Tracheophyta" rank="species" species="ryukyuensis" status="sp. nov.">
<emphasis id="B97FEAE7FFF71D15FC1BFDF5483AF232" box="[897,1062,548,572]" italics="true" pageId="8" pageNumber="203">S. ryukyuensis</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7FFF71D15FBA8FDF44893F232" bold="true" box="[1074,1167,548,572]" pageId="8" pageNumber="203">
<taxonomicNameLabel id="A24C579CFFF71D15FBA8FDF44893F232" box="[1074,1167,548,572]" pageId="8" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the paler, whitish to brownish yellow coria, the bicolored laterotergites, the globe pygophore in dorsal view, the smaller parameres with the apex acute and curved, and posterior margin of female abdominal sternite VII entirely concave toward midpoint (cleft on midline).
</paragraph>
<paragraph id="8BB436F5FFF71D15FF5DFD6348F4F2E1" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="203">
<emphasis id="B97FEAE7FFF71D15FF5DFD634D92F2C2" bold="true" box="[199,398,691,716]" pageId="8" pageNumber="203">Biological notes.</emphasis>
Adults of
<taxonomicName id="4C0B4D76FFF71D15FD89FD654E9FF2C2" box="[531,643,692,716]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="203" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF71D15FD89FD654E3DF2C2" box="[531,545,693,716]" italics="true" pageId="8" pageNumber="203">S</emphasis>
.
<emphasis id="B97FEAE7FFF71D15FDA9FD644E9FF2C2" box="[563,643,692,716]" italics="true" pageId="8" pageNumber="203">valdezi</emphasis>
</taxonomicName>
have often been found on trees of
<taxonomicName id="4C0B4D76FFF71D15FBB6FD654D04F2E1" authority="Reinw. ex Blume" authorityName="Reinw. ex Blume" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="203" phylum="Mollusca" rank="species" species="virgata">
<emphasis id="B97FEAE7FFF71D15FBB6FD6548D9F2C2" box="[1068,1221,693,716]" italics="true" pageId="8" pageNumber="203">Ficus virgata</emphasis>
Reinw. ex Blume (Moraceae)
</taxonomicName>
with fruits. This tree species might be considered to be one of host plants of
<taxonomicName id="4C0B4D76FFF71D15FBECFD0848FDF2FE" box="[1142,1249,728,752]" class="Magnoliopsida" family="Juglandaceae" genus="Sadoletus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="203" phylum="Tracheophyta" rank="species" species="valdezi">
<emphasis id="B97FEAE7FFF71D15FBECFD084898F2E1" box="[1142,1156,728,751]" italics="true" pageId="8" pageNumber="203">S</emphasis>
.
<emphasis id="B97FEAE7FFF71D15FB08FD0848FDF2FE" box="[1170,1249,728,752]" italics="true" pageId="8" pageNumber="203">valdezi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>