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<mods:title id="36D71E9DD01B7F19BC45ABEA3EC2EE00">A new spider of the genus Scotinella Banks, 1911 (Araneae: Phrurolithidae) from Mexico and new combinations in Phrurolithidae</mods:title>
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<treatment id="03A287E3126EDD0FB98BFA37619FFB0C" ID-DOI="http://doi.org/10.5281/zenodo.6309246" ID-GBIF-Taxon="193379466" ID-Zenodo-Dep="6309246" LSID="urn:lsid:plazi:treatment:03A287E3126EDD0FB98BFA37619FFB0C" httpUri="http://treatment.plazi.org/id/03A287E3126EDD0FB98BFA37619FFB0C" lastPageId="2" lastPageNumber="288" pageId="1" pageNumber="287">
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<paragraph id="8BB436F5126EDD0CB98BFA376151FA68" blockId="1.[151,526,1414,1441]" box="[151,526,1414,1441]" pageId="1" pageNumber="287">
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<emphasis id="B97FEAE7126EDD0CB98BFA376151FA68" bold="true" box="[151,526,1414,1441]" pageId="1" pageNumber="287">
Genus:
<taxonomicName id="4C0B4D76126EDD0CB9E4FA376151FA68" ID-CoL="63T7B" ID-ENA="1109756" authority="Banks, 1911" authorityName="Banks" authorityYear="1911" box="[248,526,1414,1441]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126EDD0CB9E4FA37622EFA69" bold="true" box="[248,369,1414,1441]" italics="true" pageId="1" pageNumber="287">Scotinella</emphasis>
<bibRefCitation id="EF9A4B04126EDD0CB864FA366151FA68" author="Banks, N." box="[376,526,1414,1441]" pageId="1" pageNumber="287" pagination="440 - 456" refId="ref3995" refString="Banks, N. (1911) Some Arachnida from North Carolina. Proceedings of the Academy of Natural Sciences of Philadelphia, 63, 440 - 456." type="journal article" year="1911">Banks, 1911</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BB436F5126EDD0CB98BFA77613DFA16" blockId="1.[151,610,1478,1502]" box="[151,610,1478,1502]" pageId="1" pageNumber="287">
<emphasis id="B97FEAE7126EDD0CB98BFA77627CFA16" bold="true" box="[151,291,1478,1502]" pageId="1" pageNumber="287">
<typeStatus id="54B08857126EDD0CB98BFA776393FA16" box="[151,204,1478,1502]" pageId="1" pageNumber="287">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C0B4D76126EDD0CB836FA77613DFA16" ID-CoL="6XYJ7" authority="Banks, 1911" authorityName="Banks" authorityYear="1911" box="[298,610,1478,1502]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B97FEAE7126EDD0CB836FA776282FA16" box="[298,477,1478,1502]" italics="true" pageId="1" pageNumber="287">Scotinella pallida</emphasis>
<bibRefCitation id="EF9A4B04126EDD0CB8F8FA77613DFA16" author="Banks, N." box="[484,610,1478,1502]" pageId="1" pageNumber="287" pagination="440 - 456" refId="ref3995" refString="Banks, N. (1911) Some Arachnida from North Carolina. Proceedings of the Academy of Natural Sciences of Philadelphia, 63, 440 - 456." type="journal article" year="1911">Banks, 1911</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="8BB436F5126EDD0CB98BF9BD61D1F88F" blockId="1.[151,1436,1548,2042]" pageId="1" pageNumber="287">
<emphasis id="B97FEAE7126EDD0CB98BF9BD6255F9EE" bold="true" box="[151,266,1548,1574]" pageId="1" pageNumber="287">Diagnosis</emphasis>
(after
<bibRefCitation id="EF9A4B04126EDD0CB848F9BD6138F9EE" author="Dondale, C. D. &amp; Redner, J. H." box="[340,615,1548,1574]" pageId="1" pageNumber="287" pagination="1 - 194" refId="ref4024" refString="Dondale, C. D. &amp; Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194." type="journal article" year="1982">Dondale &amp; Redner 1982</bibRefCitation>
;
<bibRefCitation id="EF9A4B04126EDD0CBB6FF9BD607AF9EF" author="Penniman, A. J." box="[627,805,1548,1575]" pageId="1" pageNumber="287" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985</bibRefCitation>
). Femur of male palp with hooked ventral apophysis and retrolateral groove; strong retrolateral tibial apophysis (RTA) composed of two processes (bifid), the dorsal (dRTA) longer than the ventral (vRTA); tegulum rounded, usually without apophysis; embolus usually expanded at base, becoming slender, spiniform to filamentous distally (
<bibRefCitation id="EF9A4B04126EDD0CBBF6F9C9675AF95A" author="Dondale, C. D. &amp; Redner, J. H." box="[746,1029,1656,1682]" pageId="1" pageNumber="287" pagination="1 - 194" refId="ref4024" refString="Dondale, C. D. &amp; Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194." type="journal article" year="1982">Dondale &amp; Redner 1982</bibRefCitation>
: figs 262, 265;
<bibRefCitation id="EF9A4B04126EDD0CBDAFF9C96638F95B" author="Penniman, A. J." box="[1203,1383,1656,1683]" pageId="1" pageNumber="287" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985</bibRefCitation>
: figs 18, 22, 30, 35). Epigyne with a single or paired copulatory openings in atrial depressions; copulatory ducts short to long, with species-specific pathway; spermathecae ovoid or round in outline, located posteriorly; bursae smaller and slenderer than spermathecae, arising at the junction of copulatory tubes with spermathecae (
<bibRefCitation id="EF9A4B04126EDD0CBDDBF955638CF8EA" author="Dondale, C. D. &amp; Redner, J. H." pageId="1" pageNumber="287" pagination="1 - 194" refId="ref4024" refString="Dondale, C. D. &amp; Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194." type="journal article" year="1982">Dondale &amp; Redner 1982</bibRefCitation>
: figs 263264;
<bibRefCitation id="EF9A4B04126EDD0CB894F8B96162F8EB" author="Penniman, A. J." box="[392,573,1800,1827]" pageId="1" pageNumber="287" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985</bibRefCitation>
: fig. 17). Eight eyes in two rows, both straight. Opisthosoma of male (and sometimes female) with large dorsal scutum.
</paragraph>
<paragraph id="8BB436F5126EDD0CB9DBF8E1601EF832" blockId="1.[151,1436,1548,2042]" pageId="1" pageNumber="287">
<emphasis id="B97FEAE7126EDD0CB9DBF8E1613BF8A2" bold="true" box="[199,612,1871,1898]" pageId="1" pageNumber="287">
Current composition of
<taxonomicName id="4C0B4D76126EDD0CB8F6F8FE6102F8A2" authorityName="Banks" authorityYear="1911" box="[490,605,1871,1898]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126EDD0CB8F6F8FE6102F8A2" bold="true" box="[490,605,1871,1898]" italics="true" pageId="1" pageNumber="287">Scotinella</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="4C0B4D76126EDD0CBB70F8E160B3F8A2" authority="(Petrunkevitch, 1910)" baseAuthorityName="Petrunkevitch" baseAuthorityYear="1910" box="[620,1004,1872,1898]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="britcheri">
<emphasis id="B97FEAE7126EDD0CBB70F8E161B4F8A2" box="[620,747,1872,1898]" italics="true" pageId="1" pageNumber="287">S. britcheri</emphasis>
(Petrunkevitch, 1910)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBAE5F8E1667AF8A2" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[1017,1317,1872,1898]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="brittoni">
<emphasis id="B97FEAE7126EDD0CBAE5F8E16735F8A2" box="[1017,1130,1872,1898]" italics="true" pageId="1" pageNumber="287">S. brittoni</emphasis>
(Gertsch, 1941)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBC2FF8E1624AF847" authority="Levi, 1951" authorityName="Levi" authorityYear="1951" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="custeri">
<emphasis id="B97FEAE7126EDD0CBC2FF8E166C3F8A2" box="[1331,1436,1872,1898]" italics="true" pageId="1" pageNumber="287">S. custeri</emphasis>
Levi, 1951
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CB83FF8C5611CF846" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[291,579,1908,1934]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="deleta">
<emphasis id="B97FEAE7126EDD0CB83FF8C562DAF846" box="[291,389,1908,1934]" italics="true" pageId="1" pageNumber="287">S. deleta</emphasis>
(Gertsch, 1941)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBB4EF8C56023F846" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[594,892,1908,1934]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="divesta">
<emphasis id="B97FEAE7126EDD0CBB4EF8C561E0F846" box="[594,703,1908,1934]" italics="true" pageId="1" pageNumber="287">S. divesta</emphasis>
(Gertsch, 1941)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBA97F8C5679DF846" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[907,1218,1908,1934]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="divinula">
<emphasis id="B97FEAE7126EDD0CBA97F8C5675AF846" box="[907,1029,1908,1934]" italics="true" pageId="1" pageNumber="287">S. divinula</emphasis>
(Gertsch, 1941)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBDCDF8C5638DF87B" authority="Roddy, 1957" authorityName="Roddy" authorityYear="1957" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="dixiana">
<emphasis id="B97FEAE7126EDD0CBDCDF8C5661CF846" box="[1233,1347,1908,1934]" italics="true" pageId="1" pageNumber="287">S. dixiana</emphasis>
Roddy, 1957
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CB9C2F829614DF87B" authority="(Gertsch, 1935)" baseAuthorityName="Gertsch" baseAuthorityYear="1935" box="[222,530,1944,1971]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="fratrella">
<emphasis id="B97FEAE7126EDD0CB9C2F8296208F87A" box="[222,343,1944,1970]" italics="true" pageId="1" pageNumber="287">S. fratrella</emphasis>
(Gertsch, 1935)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBB02F829606BF87B" authority="Levi, 1951" authorityName="Levi" authorityYear="1951" box="[542,820,1944,1971]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="madisonia">
<emphasis id="B97FEAE7126EDD0CBB02F82961EFF87A" box="[542,688,1944,1970]" italics="true" pageId="1" pageNumber="287">S. madisonia</emphasis>
Levi, 1951
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBA5DF8296762F87B" authority="Levi, 1951" authorityName="Levi" authorityYear="1951" box="[833,1085,1944,1971]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="manitou">
<emphasis id="B97FEAE7126EDD0CBA5DF82960E6F87A" box="[833,953,1944,1970]" italics="true" pageId="1" pageNumber="287">S. manitou</emphasis>
Levi, 1951
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBD56F829621FF81E" authority="(Chamberlin &amp; Gertsch, 1930)" baseAuthorityName="Chamberlin &amp; Gertsch" baseAuthorityYear="1930" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="minnetonka">
<emphasis id="B97FEAE7126EDD0CBD56F82967B5F87A" box="[1098,1258,1944,1970]" italics="true" pageId="1" pageNumber="287">S. minnetonka</emphasis>
(Chamberlin &amp; Gertsch, 1930)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CB850F80D6114F81E" authority="Banks, 1911" authorityName="Banks" authorityYear="1911" box="[332,587,1980,2006]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B97FEAE7126EDD0CB850F80D62E8F81E" box="[332,439,1980,2006]" italics="true" pageId="1" pageNumber="287">S. pallida</emphasis>
<bibRefCitation id="EF9A4B04126EDD0CB8A2F80D6114F81E" author="Banks, N." box="[446,587,1980,2006]" pageId="1" pageNumber="287" pagination="440 - 456" refId="ref3995" refString="Banks, N. (1911) Some Arachnida from North Carolina. Proceedings of the Academy of Natural Sciences of Philadelphia, 63, 440 - 456." type="journal article" year="1911">Banks, 1911</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBB4BF80D670DF81E" authority="(Chamberlin &amp; Gertsch, 1930)" baseAuthorityName="Chamberlin &amp; Gertsch" baseAuthorityYear="1930" box="[599,1106,1980,2006]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="pelvicolens">
<emphasis id="B97FEAE7126EDD0CBB4BF80D61AEF81E" box="[599,753,1980,2006]" italics="true" pageId="1" pageNumber="287">S. pelvicolens</emphasis>
(Chamberlin &amp; Gertsch, 1930)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CBD41F80D66C7F81E" authority="(Emerton, 1890)" baseAuthorityName="Emerton" baseAuthorityYear="1890" box="[1117,1432,1980,2006]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="pugnata">
<emphasis id="B97FEAE7126EDD0CBD41F80D678AF81E" box="[1117,1237,1980,2006]" italics="true" pageId="1" pageNumber="287">S. pugnata</emphasis>
(Emerton, 1890)
</taxonomicName>
,
<taxonomicName id="4C0B4D76126EDD0CB98BF851628BF832" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[151,468,2016,2042]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="redempta">
<emphasis id="B97FEAE7126EDD0CB98BF8516243F832" box="[151,284,2016,2042]" italics="true" pageId="1" pageNumber="287">S. redempta</emphasis>
(Gertsch, 1941)
</taxonomicName>
, and
<taxonomicName id="4C0B4D76126EDD0CBB0CF8516061F832" authority="(Gertsch, 1941)" baseAuthorityName="Gertsch" baseAuthorityYear="1941" box="[528,830,2016,2042]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B97FEAE7126EDD0CBB0CF85161DCF832" box="[528,643,2016,2042]" italics="true" pageId="1" pageNumber="287">S. sculleni</emphasis>
(Gertsch, 1941)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C311657E126DDD0FB9DBFF27602BFCDC" pageId="2" pageNumber="288" type="discussion">
<paragraph id="8BB436F5126DDD0FB9DBFF276650FDD0" blockId="2.[151,1437,150,1221]" pageId="2" pageNumber="288">
<emphasis id="B97FEAE7126DDD0FB9DBFF276264FF78" bold="true" box="[199,315,150,176]" pageId="2" pageNumber="288">Remarks.</emphasis>
<bibRefCitation id="EF9A4B04126DDD0FB859FF27612AFF78" author="Zamani, A. &amp; Marusik, Y. M." box="[325,629,150,176]" pageId="2" pageNumber="288" pagination="311 - 329" refId="ref4647" refString="Zamani, A. &amp; Marusik, Y. M. (2020) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa, 4758 (2), 311 - 329. https: // doi. org / 10.11646 / zootaxa. 4758.2.6" type="journal article" year="2020">Zamani &amp; Marusik (2020)</bibRefCitation>
delimited the genus
<taxonomicName id="4C0B4D76126DDD0FBA74FF2760A9FF78" authorityName="C. L. Koch" authorityYear="1839" box="[872,1014,150,176]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBA74FF2760A9FF78" box="[872,1014,150,176]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
to include only those species whose males have a large unbranched retroventral tibial apophysis (referred to as RTA by
<bibRefCitation id="EF9A4B04126DDD0FBD5AFF0A6618FF1C" author="Komnenov, M. &amp; Pitta, E. &amp; Zografou, K. &amp; Chatzaki, M." box="[1094,1351,186,213]" pageId="2" pageNumber="288" pagination="1 - 66" refId="ref4187" refString="Komnenov, M., Pitta, E., Zografou, K. &amp; Chatzaki, M. (2016) Discovering the still unexplored arachnofauna of the National Park of Dadia-Lefkimi-Soufli, NE Greece: a taxonomic review with description of new species. Zootaxa, 4096 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4096.1.1" type="journal article" year="2016">
Komnenov
<emphasis id="B97FEAE7126DDD0FBDD6FF0A67A2FF1C" box="[1226,1277,186,212]" italics="true" pageId="2" pageNumber="288">et al</emphasis>
. 2016
</bibRefCitation>
;
<bibRefCitation id="EF9A4B04126DDD0FBC4FFF0B6256FF31" author="Penniman, A. J." pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985</bibRefCitation>
;
<bibRefCitation id="EF9A4B04126DDD0FB809FF6F62EDFF30" author="Platnick, N. I." box="[277,434,222,248]" pageId="2" pageNumber="288" pagination="1 - 29" refId="ref4413" refString="Platnick, N. I. (2019) The guardstone spiders of the Phrurotimpus palustris group (Araneae, Phrurolithidae). American Museum Novitates, 3944, 1 - 29. https: // doi. org / 10.1206 / 3944.1" type="journal article" year="2019">Platnick 2019</bibRefCitation>
;
<bibRefCitation id="EF9A4B04126DDD0FB8A2FF6E6121FF31" author="Wang, L. Y. &amp; Chen, H. M. &amp; Zhou, K. X. &amp; Zhang, F. &amp; Zhang, Z. S." box="[446,638,222,249]" pageId="2" pageNumber="288" pagination="447 - 464" refId="ref4526" refString="Wang, L. Y., Chen, H. M., Zhou, K. X., Zhang, F. &amp; Zhang, Z. S. (2015) Diversity of spiders in Fanjing Mountain Nature Reserve, Guizhou, China, I: Six new species of Phrurolithidae (Araneae). Zootaxa, 4012 (3), 447 - 464. https: // doi. org / 10.11646 / zootaxa. 4012.3.2" type="journal article" year="2015">
Wang
<emphasis id="B97FEAE7126DDD0FBB19FF6E6169FF30" box="[517,566,222,248]" italics="true" pageId="2" pageNumber="288">et al</emphasis>
. 2015
</bibRefCitation>
) and a small RTA. Although
<bibRefCitation id="EF9A4B04126DDD0FBADFFF6F67B4FF30" author="Zamani, A. &amp; Marusik, Y. M." box="[963,1259,222,248]" pageId="2" pageNumber="288" pagination="311 - 329" refId="ref4647" refString="Zamani, A. &amp; Marusik, Y. M. (2020) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa, 4758 (2), 311 - 329. https: // doi. org / 10.11646 / zootaxa. 4758.2.6" type="journal article" year="2020">Zamani &amp; Marusik (2020)</bibRefCitation>
did not provide diagnostic characteristics for female genitalia,
<taxonomicName id="4C0B4D76126DDD0FBB83FEB260B7FED5" authority="(C. L. Koch, 1835)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1835" box="[671,1000,258,285]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B97FEAE7126DDD0FBB83FEB26051FED4" box="[671,782,258,284]" italics="true" pageId="2" pageNumber="288">P. festivus</emphasis>
(C. L. Koch, 1835)
</taxonomicName>
and other Old World
<taxonomicName id="4C0B4D76126DDD0FBDC5FEB3639CFE88" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97FEAE7126DDD0FBDC5FEB36638FED4" box="[1241,1383,258,284]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
species
</taxonomicName>
have additional slightly sclerotized receptacles located anteriorly, which are larger than spermathecae. These structures are referred to as secondary receptacles by
<bibRefCitation id="EF9A4B04126DDD0FBBF7FEFB60FAFEAD" author="Almquist, S." box="[747,933,330,357]" pageId="2" pageNumber="288" pagination="285 - 601" refId="ref3881" refString="Almquist, S. (2006) Swedish Araneae, part 2 - families Dictynidae to Salticidae. Insect Systematics &amp; Evolution, Supplement, 63, 285 - 601." type="journal article" year="2006">Almquist (2006)</bibRefCitation>
and
<bibRefCitation id="EF9A4B04126DDD0FBAC7FEFB6658FEAC" author="Zamani, A. &amp; Marusik, Y. M." box="[987,1287,330,356]" pageId="2" pageNumber="288" pagination="311 - 329" refId="ref4647" refString="Zamani, A. &amp; Marusik, Y. M. (2020) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa, 4758 (2), 311 - 329. https: // doi. org / 10.11646 / zootaxa. 4758.2.6" type="journal article" year="2020">Zamani &amp; Marusik (2020)</bibRefCitation>
; as bursae by
<bibRefCitation id="EF9A4B04126DDD0FB98BFEDF6206FE41" author="Penniman, A. J." box="[151,345,366,393]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman (1985)</bibRefCitation>
and
<bibRefCitation id="EF9A4B04126DDD0FB88DFEDE611FFE40" author="Kamura, T." box="[401,576,366,393]" pageId="2" pageNumber="288" pagination="117 - 130" refId="ref4113" refString="Kamura, T. (2021) Three new genera of the family Phrurolithidae (Araneae) from East Asia. Acta Arachnologica, 70, 117 - 130. https: // doi. org / 10.2476 / asjaa. 70.117" type="journal article" year="2021">Kamura (2021)</bibRefCitation>
, and as anterior part of spermathecae by
<bibRefCitation id="EF9A4B04126DDD0FBD13FEDE667EFE41" author="Komnenov, M. &amp; Pitta, E. &amp; Zografou, K. &amp; Chatzaki, M." box="[1039,1313,366,393]" pageId="2" pageNumber="288" pagination="1 - 66" refId="ref4187" refString="Komnenov, M., Pitta, E., Zografou, K. &amp; Chatzaki, M. (2016) Discovering the still unexplored arachnofauna of the National Park of Dadia-Lefkimi-Soufli, NE Greece: a taxonomic review with description of new species. Zootaxa, 4096 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4096.1.1" type="journal article" year="2016">
Komnenov
<emphasis id="B97FEAE7126DDD0FBD8FFEDE679AFE40" box="[1171,1221,366,392]" italics="true" pageId="2" pageNumber="288">et al</emphasis>
. (2016)
</bibRefCitation>
. Nevertheless, such receptacles may not be homologous with
<taxonomicName id="4C0B4D76126DDD0FBBC8FE23601BFE64" authorityName="Banks" authorityYear="1911" box="[724,836,402,428]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBBC8FE23601BFE64" box="[724,836,402,428]" italics="true" pageId="2" pageNumber="288">Scotinella</emphasis>
</taxonomicName>
bursae (referred to as spermathecal organ by
<bibRefCitation id="EF9A4B04126DDD0FBC21FE23623BFE18" author="Dondale, C. D. &amp; Redner, J. H." pageId="2" pageNumber="288" pagination="1 - 194" refId="ref4024" refString="Dondale, C. D. &amp; Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194." type="journal article" year="1982">Dondale &amp; Redner (1982))</bibRefCitation>
, because
<bibRefCitation id="EF9A4B04126DDD0FB8CCFE0761CDFE19" author="Penniman, A. J." box="[464,658,438,465]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman (1985)</bibRefCitation>
used the term in a broad sense to include those “blind sac-like or tubular structures” with different sizes in relation to the spermathecae, that “may arise near the copulatory openings, near the middle of copulatory tubes, or at the junction of the copulatory tubes with the spermathecae”.
</paragraph>
<paragraph id="8BB436F5126DDD0FB9DBFD92602BFCDC" blockId="2.[151,1437,150,1221]" pageId="2" pageNumber="288">
We agree with the opinions of
<bibRefCitation id="EF9A4B04126DDD0FBB3CFD92619CFDF4" author="Kaston, B. J." box="[544,707,546,573]" pageId="2" pageNumber="288" refId="ref4155" refString="Kaston, B. J. (1972) How to know the spiders. 2 nd Edition. W. C. Brown Co., Dubuque, 289 pp." type="book" year="1972">Kaston (1972)</bibRefCitation>
,
<bibRefCitation id="EF9A4B04126DDD0FBBD3FD9360CCFDF4" author="Penniman, A. J." box="[719,915,546,572]" pageId="2" pageNumber="288" pagination="125 - 132" refId="ref4336" refString="Penniman, A. J. (1978) Taxonomic and natural history notes on Phrurolithus fratrellus Gertsch (Araneae: Clubionidae). Journal of Arachnology, 6, 125 - 132." type="journal article" year="1978">Penniman (1978)</bibRefCitation>
,
<bibRefCitation id="EF9A4B04126DDD0FBA83FD936798FDF4" author="Dondale, C. D. &amp; Redner, J. H." box="[927,1223,546,572]" pageId="2" pageNumber="288" pagination="1 - 194" refId="ref4024" refString="Dondale, C. D. &amp; Redner, J. H. (1982) The insects and arachnids of Canada, Part 9. The sac spiders of Canada and Alaska, Araneae: Clubionidae and Anyphaenidae. Research Branch Agriculture Canada Publication, 1724, 1 - 194." type="journal article" year="1982">Dondale &amp; Redner (1982)</bibRefCitation>
,
<bibRefCitation id="EF9A4B04126DDD0FBDCFFD9366C8FDF5" author="Penniman, A. J." box="[1235,1431,546,573]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman (1985)</bibRefCitation>
,
<bibRefCitation id="EF9A4B04126DDD0FB98BFDF66233FDA9" author="Wang, L. Y. &amp; Chen, H. M. &amp; Zhou, K. X. &amp; Zhang, F. &amp; Zhang, Z. S." box="[151,364,582,609]" pageId="2" pageNumber="288" pagination="447 - 464" refId="ref4526" refString="Wang, L. Y., Chen, H. M., Zhou, K. X., Zhang, F. &amp; Zhang, Z. S. (2015) Diversity of spiders in Fanjing Mountain Nature Reserve, Guizhou, China, I: Six new species of Phrurolithidae (Araneae). Zootaxa, 4012 (3), 447 - 464. https: // doi. org / 10.11646 / zootaxa. 4012.3.2" type="journal article" year="2015">
Wang
<emphasis id="B97FEAE7126DDD0FB9C3FDF6624EFDA8" box="[223,273,582,608]" italics="true" pageId="2" pageNumber="288">et al</emphasis>
. (2015)
</bibRefCitation>
,
<bibRefCitation id="EF9A4B04126DDD0FB864FDF76177FDA8" author="Platnick, N. I." box="[376,552,582,608]" pageId="2" pageNumber="288" pagination="1 - 29" refId="ref4413" refString="Platnick, N. I. (2019) The guardstone spiders of the Phrurotimpus palustris group (Araneae, Phrurolithidae). American Museum Novitates, 3944, 1 - 29. https: // doi. org / 10.1206 / 3944.1" type="journal article" year="2019">Platnick (2019)</bibRefCitation>
, and
<bibRefCitation id="EF9A4B04126DDD0FBB7FFDF760D2FDA8" author="Zamani, A. &amp; Marusik, Y. M." box="[611,909,582,608]" pageId="2" pageNumber="288" pagination="311 - 329" refId="ref4647" refString="Zamani, A. &amp; Marusik, Y. M. (2020) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa, 4758 (2), 311 - 329. https: // doi. org / 10.11646 / zootaxa. 4758.2.6" type="journal article" year="2020">Zamani &amp; Marusik (2020)</bibRefCitation>
that the North American
<taxonomicName id="4C0B4D76126DDD0FBDB0FDF76665FDA8" authorityName="C. L. Koch" authorityYear="1839" box="[1196,1338,582,608]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBDB0FDF76665FDA8" box="[1196,1338,582,608]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
are misplaced. Furthermore, the Mexican species currently placed in
<taxonomicName id="4C0B4D76126DDD0FBA57FDDB6086FD4C" authorityName="C. L. Koch" authorityYear="1839" box="[843,985,618,644]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBA57FDDB6086FD4C" box="[843,985,618,644]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
do not correspond to Old World species in several features. For example, all Mexican
<taxonomicName id="4C0B4D76126DDD0FBBBFFD3F606EFD60" authorityName="C. L. Koch" authorityYear="1839" box="[675,817,654,680]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBBBFFD3F606EFD60" box="[675,817,654,680]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
have a bifid RTA, hooked femoral apophysis, and the bulb lacking a tegular “knob” (referred to as a subterminal apophysis by
<bibRefCitation id="EF9A4B04126DDD0FBAD5FD0367AAFD04" author="Zamani, A. &amp; Marusik, Y. M." box="[969,1269,690,716]" pageId="2" pageNumber="288" pagination="311 - 329" refId="ref4647" refString="Zamani, A. &amp; Marusik, Y. M. (2020) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa, 4758 (2), 311 - 329. https: // doi. org / 10.11646 / zootaxa. 4758.2.6" type="journal article" year="2020">Zamani &amp; Marusik (2020)</bibRefCitation>
, and as tegular apophysis by
<bibRefCitation id="EF9A4B04126DDD0FB82CFD6762B5FD39" author="Almquist, S." box="[304,490,726,753]" pageId="2" pageNumber="288" pagination="285 - 601" refId="ref3881" refString="Almquist, S. (2006) Swedish Araneae, part 2 - families Dictynidae to Salticidae. Insect Systematics &amp; Evolution, Supplement, 63, 285 - 601." type="journal article" year="2006">Almquist (2006)</bibRefCitation>
and
<bibRefCitation id="EF9A4B04126DDD0FBB3DFD666186FD38" author="Kamura, T." box="[545,729,726,753]" pageId="2" pageNumber="288" pagination="117 - 130" refId="ref4113" refString="Kamura, T. (2021) Three new genera of the family Phrurolithidae (Araneae) from East Asia. Acta Arachnologica, 70, 117 - 130. https: // doi. org / 10.2476 / asjaa. 70.117" type="journal article" year="2021">Kamura (2021))</bibRefCitation>
, and the bursa (
<emphasis id="B97FEAE7126DDD0FBA97FD666096FD38" box="[907,969,727,752]" italics="true" pageId="2" pageNumber="288">sensu</emphasis>
<bibRefCitation id="EF9A4B04126DDD0FBACCFD6767DFFD39" author="Penniman, A. J." box="[976,1152,726,753]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985</bibRefCitation>
) is a small tubular protuberance at the junction of the copulatory ducts and spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C311657E126DDD0FB9DBFCAF619FFB0C" pageId="2" pageNumber="288" type="diagnosis">
<paragraph id="8BB436F5126DDD0FB9DBFCAF619FFB0C" blockId="2.[151,1437,150,1221]" pageId="2" pageNumber="288">
Based on similarities in the genitalic and somatic morphology of the Mexican
<taxonomicName id="4C0B4D76126DDD0FBD24FCAF6799FCF0" authorityName="C. L. Koch" authorityYear="1839" box="[1080,1222,798,824]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBD24FCAF6799FCF0" box="[1080,1222,798,824]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
to the
<typeStatus id="54B08857126DDD0FBC08FCAE661BFCF1" box="[1300,1348,799,825]" pageId="2" pageNumber="288">type</typeStatus>
species of
<taxonomicName id="4C0B4D76126DDD0FB9AAFCF36279FC94" authorityName="Banks" authorityYear="1911" box="[182,294,834,860]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FB9AAFCF36279FC94" box="[182,294,834,860]" italics="true" pageId="2" pageNumber="288">Scotinella</emphasis>
</taxonomicName>
and other North American
<taxonomicName id="4C0B4D76126DDD0FBB45FCF36196FC94" authorityName="Banks" authorityYear="1911" box="[601,713,834,860]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBB45FCF36196FC94" box="[601,713,834,860]" italics="true" pageId="2" pageNumber="288">Scotinella</emphasis>
</taxonomicName>
(see diagnosis), the following species are transferred:
<taxonomicName id="4C0B4D76126DDD0FBC30FCF36148FC48" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="adjacens" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBC30FCF363A4FC48" italics="true" pageId="2" pageNumber="288">Scotinella adjacens</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FB815FCD76150FC48" author="Gertsch, W. J. &amp; Davis, L. I." box="[265,527,870,896]" pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBB02FCD761C0FC48" bold="true" box="[542,671,870,896]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBB02FCD761C0FC48" box="[542,671,870,896]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0B4D76126DDD0FBBB1FCD66720FC48" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[685,1151,870,897]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="approximatus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBBB1FCD6603DFC48" box="[685,866,871,897]" italics="true" pageId="2" pageNumber="288">S. approximatus</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FBA6DFCD76729FC48" author="Gertsch, W. J. &amp; Davis, L. I." box="[881,1142,870,896]" pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBD9AFCD76658FC48" bold="true" box="[1158,1287,870,896]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBD9AFCD76658FC48" box="[1158,1287,870,896]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0B4D76126DDD0FBC09FCD6628AFC6C" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="coahuilanus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBC09FCD663E1FC6C" italics="true" pageId="2" pageNumber="288">S. coahuilanus</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FB9D7FC3B6293FC6C" author="Gertsch, W. J. &amp; Davis, L. I." box="[203,460,906,932]" pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FB8C6FC3B6106FC6C" bold="true" box="[474,601,906,932]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FB8C6FC3B6106FC6C" box="[474,601,906,932]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0B4D76126DDD0FBB7AFC3A60BDFC6C" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[614,994,906,933]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="debilis" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBB7AFC3A6193FC6C" box="[614,716,906,933]" italics="true" pageId="2" pageNumber="288">S. debilis</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FBBC4FC3B6086FC6C" author="Gertsch, W. J. &amp; Davis, L. I." box="[728,985,906,932]" pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBAFBFC3B6739FC6C" bold="true" box="[999,1126,906,932]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBAFBFC3B6739FC6C" box="[999,1126,906,932]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0B4D76126DDD0FBD6FFC3A6258FC00" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="diversus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBD6FFC3A67B4FC6C" box="[1139,1259,906,933]" italics="true" pageId="2" pageNumber="288">S. diversus</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FBDE4FC3B63A0FC00" author="Gertsch, W. J. &amp; Davis, L. I." pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FB812FC1F62D1FC00" bold="true" box="[270,398,942,968]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FB812FC1F62D1FC00" box="[270,398,942,968]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0B4D76126DDD0FB887FC1E6034FC00" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[411,875,942,969]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="tamaulipanus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FB887FC1E6110FC00" box="[411,591,942,969]" italics="true" pageId="2" pageNumber="288">S. tamaulipanus</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FBB42FC1F603DFC00" author="Gertsch, W. J. &amp; Davis, L. I." box="[606,866,942,968]" pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBA6DFC1F60AEFC00" bold="true" box="[881,1009,942,968]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBA6DFC1F60AEFC00" box="[881,1009,942,968]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C0B4D76126DDD0FBD32FC1E6386FC24" authority="(Gertsch &amp; Davis, 1940)" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="tepejicanus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBD32FC1E6795FC00" box="[1070,1226,943,969]" italics="true" pageId="2" pageNumber="288">S. tepejicanus</emphasis>
(
<bibRefCitation id="EF9A4B04126DDD0FBDC4FC1F638FFC24" author="Gertsch, W. J. &amp; Davis, L. I." pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis, 1940</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FB9C2FC636202FC24" bold="true" box="[222,349,978,1004]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FB9C2FC636202FC24" box="[222,349,978,1004]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Although the males of
<taxonomicName id="4C0B4D76126DDD0FBB41FC62619DFC24" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[605,706,978,1005]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="debilis" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBB41FC62619DFC24" box="[605,706,978,1005]" italics="true" pageId="2" pageNumber="288">S. debilis</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBBD4FC636018FC24" bold="true" box="[712,839,978,1004]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBBD4FC636018FC24" box="[712,839,978,1004]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0B4D76126DDD0FBA66FC6260ADFC24" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[890,1010,978,1005]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="diversus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBA66FC6260ADFC24" box="[890,1010,978,1005]" italics="true" pageId="2" pageNumber="288">S. diversus</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBAE4FC636728FC24" bold="true" box="[1016,1143,978,1004]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBAE4FC636728FC24" box="[1016,1143,978,1004]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
remain unknown,
<bibRefCitation id="EF9A4B04126DDD0FBC59FC636217FBD8" author="Gertsch, W. J. &amp; Davis, L. I." pageId="2" pageNumber="288" pagination="1 - 22" refId="ref4076" refString="Gertsch, W. J. &amp; Davis, L. I. (1940) Report on a collection of spiders from Mexico. III. American Museum Novitates, 1069, 1 - 22." type="journal article" year="1940">Gertsch &amp; Davis (1940)</bibRefCitation>
considered these two species congeneric with
<taxonomicName id="4C0B4D76126DDD0FBA44FC4660A2FBD8" authorityName="Chamé-Vázquez &amp; Jiménez" authorityYear="2022" baseAuthorityName="Gertsch &amp; Davis" baseAuthorityYear="1940" box="[856,1021,1014,1041]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="coahuilanus" status="comb. nov.">
<emphasis id="B97FEAE7126DDD0FBA44FC4660A2FBD8" box="[856,1021,1014,1041]" italics="true" pageId="2" pageNumber="288">S. coahuilanus</emphasis>
</taxonomicName>
<emphasis id="B97FEAE7126DDD0FBD19FC4767D9FBD8" bold="true" box="[1029,1158,1014,1040]" pageId="2" pageNumber="288">
<taxonomicNameLabel id="A24C579C126DDD0FBD19FC4767D9FBD8" box="[1029,1158,1014,1040]" pageId="2" pageNumber="288" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, which has an endogyne similar to other
<taxonomicName id="4C0B4D76126DDD0FB859FBAB6153FBFC" box="[325,524,1050,1076]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97FEAE7126DDD0FB859FBAB62EAFBFC" box="[325,437,1050,1076]" italics="true" pageId="2" pageNumber="288">Scotinella</emphasis>
species
</taxonomicName>
(see
<bibRefCitation id="EF9A4B04126DDD0FBB58FBAB6074FBFC" author="Penniman, A. J." box="[580,811,1050,1077]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman 1985: 100</bibRefCitation>
, fig. 99). There are several American phrurolithids listed under
<taxonomicName id="4C0B4D76126DDD0FB9C1FB8F6234FB90" authorityName="C. L. Koch" authorityYear="1839" box="[221,363,1086,1112]" class="Arachnida" family="Phrurolithidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FB9C1FB8F6234FB90" box="[221,363,1086,1112]" italics="true" pageId="2" pageNumber="288">Phrurolithus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B4D76126DDD0FB8BCFB8F616CFB90" authorityName="Chamberlin" authorityYear="1921" box="[416,563,1086,1112]" class="Arachnida" family="Phrurolithidae" genus="Phruronellus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FB8BCFB8F616CFB90" box="[416,563,1086,1112]" italics="true" pageId="2" pageNumber="288">Phruronellus</emphasis>
</taxonomicName>
, all of them likely misplaced. Moreover, we agree with the opinions of
<bibRefCitation id="EF9A4B04126DDD0FBC51FB8E63BDFBB4" author="Kaston, B. J." pageId="2" pageNumber="288" refId="ref4155" refString="Kaston, B. J. (1972) How to know the spiders. 2 nd Edition. W. C. Brown Co., Dubuque, 289 pp." type="book" year="1972">Kaston (1972)</bibRefCitation>
and
<bibRefCitation id="EF9A4B04126DDD0FB807FBD36281FBB5" author="Penniman, A. J." box="[283,478,1122,1149]" pageId="2" pageNumber="288" refId="ref4370" refString="Penniman, A. J. (1985) Revision of the britcheri and pugnata groups of Scotinella (Araneae Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders. The Ohio State University, Columbus, Ohio, 247 pp." type="book" year="1985">Penniman (1985)</bibRefCitation>
that
<taxonomicName id="4C0B4D76126DDD0FBB04FBD361D7FBB4" authorityName="Banks" authorityYear="1911" box="[536,648,1122,1148]" class="Arachnida" family="Phrurolithidae" genus="Scotinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBB04FBD361D7FBB4" box="[536,648,1122,1148]" italics="true" pageId="2" pageNumber="288">Scotinella</emphasis>
</taxonomicName>
is a senior synonym of
<taxonomicName id="4C0B4D76126DDD0FBA80FBD36770FBB4" authorityName="Chamberlin" authorityYear="1921" box="[924,1071,1122,1148]" class="Arachnida" family="Phrurolithidae" genus="Phruronellus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBA80FBD36770FBB4" box="[924,1071,1122,1148]" italics="true" pageId="2" pageNumber="288">Phruronellus</emphasis>
</taxonomicName>
; see also
<bibRefCitation id="EF9A4B04126DDD0FBDBCFBD36610FBB4" author="Roth, V. D." box="[1184,1359,1122,1149]" pageId="2" pageNumber="288" refId="ref4456" refString="Roth, V. D. (1985) Spider genera of North America. American Arachnological Society, Gainesville, Florida, 176, pp." type="book" year="1985">Roth (1985: 10</bibRefCitation>
;
<bibRefCitation id="EF9A4B04126DDD0FBC40FBD363EBFB68" author="Roth, V. D." pageId="2" pageNumber="288" refId="ref4483" refString="Roth, V. D. (1994) Spider Genera of North America, with Keys to Families and Genera, and a Guide to Literature. 3 rd Edition. American Arachnological Society, Gainesville, Florida, 203 pp." type="book" year="1994">1994: 83</bibRefCitation>
). However, reassessment of the
<typeStatus id="54B08857126DDD0FBB34FB36613CFB69" box="[552,611,1159,1185]" pageId="2" pageNumber="288">types</typeStatus>
of all
<taxonomicName id="4C0B4D76126DDD0FBBACFB37601CFB68" authorityName="Chamberlin" authorityYear="1921" box="[688,835,1158,1184]" class="Arachnida" family="Phrurolithidae" genus="Phruronellus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE7126DDD0FBBACFB37601CFB68" box="[688,835,1158,1184]" italics="true" pageId="2" pageNumber="288">Phruronellus</emphasis>
</taxonomicName>
are particularly needed, because many details of the male and female copulatory organs are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>