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<document id="AE3D03A37E33E44F8FBBEA8FCF529044" ID="10.11646/zootaxa.4147.1.2" ID-DOI="10.11646/zootaxa.4147.1.2" ID-GBIF-Dataset="b0e2f296-95a9-4ffc-92d8-ca46a49997c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="262564" ID-ZooBank="4CBE20C2-2851-4B16-8626-26C373413E1A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470119269460" checkinUser="plazi" docAuthor="Chagas-Jr, Amazonas" docDate="2016" docId="03A287E1A031BF61FF14767EC1F1FE72" docLanguage="en" docName="zootaxa.4147.1.2.pdf" docOrigin="Zootaxa 4147 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otostigmus (Parotostigmus) clavifer Chamberlin 1921" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FF9BFF99A034BF66FF837443C37AFFBD" masterDocTitle="A systematic appraisal of the types of ten species of Otostigmus (Parotostigmus) (Scolopendromorpha, Scolopendridae, Otostigminae)" masterLastPageNumber="58" masterPageNumber="36" pageNumber="41" updateTime="1698366654858" updateUser="plazi">
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<mods:title id="FF149C288B489463C4806EE8819045C3">A systematic appraisal of the types of ten species of Otostigmus (Parotostigmus) (Scolopendromorpha, Scolopendridae, Otostigminae)</mods:title>
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<treatment id="03A287E1A031BF61FF14767EC1F1FE72" ID-DOI="http://doi.org/10.5281/zenodo.5625510" ID-GBIF-Taxon="123513995" ID-Zenodo-Dep="5625510" LSID="urn:lsid:plazi:treatment:03A287E1A031BF61FF14767EC1F1FE72" httpUri="http://treatment.plazi.org/id/03A287E1A031BF61FF14767EC1F1FE72" lastPageId="7" lastPageNumber="43" pageId="5" pageNumber="41">
<subSubSection id="C311657CA031BF63FF14767EC233FDC5" pageId="5" pageNumber="41" type="nomenclature">
<paragraph id="8BB436F7A031BF63FF14767EC049FDEA" blockId="5.[151,819,573,632]" box="[151,819,573,599]" pageId="5" pageNumber="41">
<heading id="D0FC819BA031BF63FF14767EC049FDEA" bold="true" box="[151,819,573,599]" fontSize="11" level="1" pageId="5" pageNumber="41" reason="1">
<taxonomicName id="4C0B4D74A031BF63FF14767EC049FDEA" ID-CoL="75DNT" authority="Chamberlin, 1921" authorityName="Chamberlin" authorityYear="1921" box="[151,819,573,599]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="clavifer" subGenus="Parotostigmus">
<emphasis id="B97FEAE5A031BF63FF14767EC049FDEA" bold="true" box="[151,819,573,599]" pageId="5" pageNumber="41">
<emphasis id="B97FEAE5A031BF63FF14767EC25AFDEA" bold="true" box="[151,288,573,599]" italics="true" pageId="5" pageNumber="41">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A031BF63FEB2767EC2A5FDEA" bold="true" box="[305,479,573,599]" italics="true" pageId="5" pageNumber="41">Parotostigmus</emphasis>
)
<emphasis id="B97FEAE5A031BF63FE73767EC131FDEA" bold="true" box="[496,587,573,599]" italics="true" pageId="5" pageNumber="41">clavifer</emphasis>
<bibRefCitation id="EF9A4B06A031BF63FDD1767EC049FDEA" author="Chamberlin" box="[594,819,573,599]" pageId="5" pageNumber="41" refString="Chamberlin, R. V. (1921) Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, 97, 1 - 28." type="journal article" year="1921">Chamberlin, 1921</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BB436F7A031BF63FF14761CC233FDC5" blockId="5.[151,819,573,632]" box="[151,329,607,632]" pageId="5" pageNumber="41">
(
<figureCitation id="13302A72A031BF63FF1D761CC23BFDC5" box="[158,321,607,632]" captionStart-0="FIGURES 11 12" captionStart-1="FIGURES 13 14" captionStartId-0="5.[151,264,1981,2004]" captionStartId-1="6.[151,264,1168,1191]" captionTargetBox-0="[151,1436,1193,1960]" captionTargetBox-1="[151,1435,193,1147]" captionTargetId-0="figure@5.[151,1436,1193,1960]" captionTargetId-1="figure@6.[151,1435,193,1147]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 11 12. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 11. Habitus, (dorsal view). 12. Habitus, (ventral view). Scale bars 5 mm." captionText-1="FIGURES 13 14. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 13. Cephalic plate and tergites 1 3. 14. Tergite of ultimate leg-bearing segment and prefemur of ultimate legs showing the digitiform appendix. DA—Digitiform appendix; TH—Tuft of hairs. Scale bars 1 mm." httpUri-0="https://zenodo.org/record/262569/files/figure.png" httpUri-1="https://zenodo.org/record/262570/files/figure.png" pageId="5" pageNumber="41">Figures 1114</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311657CA031BF63FF1476EAC09FFCA7" pageId="5" pageNumber="41" type="reference_group">
<paragraph id="8BB436F7A031BF63FF1476EAC1B1FD03" blockId="5.[151,997,680,794]" box="[151,715,680,702]" pageId="5" pageNumber="41">
<treatmentCitationGroup id="AB1B11D9A031BF63FF1476EAC1B1FD03" box="[151,715,680,702]" pageId="5" pageNumber="41">
<taxonomicName id="4C0B4D74A031BF63FF1476EAC139FD03" ID-CoL="75DNT" authority="Chamberlin, 1921: 13" authorityName="Chamberlin" authorityPageNumber="13" authorityYear="1921" box="[151,579,680,702]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A031BF63FF1476EAC227FD03" box="[151,349,680,702]" italics="true" pageId="5" pageNumber="41">Otostigmus clavifer</emphasis>
<treatmentCitation id="0AAA10E6A031BF63FEE776EBC139FD03" author="Chamberlin" box="[356,579,680,702]" page="13" pageId="5" pageNumber="41" year="1921">
<bibRefCitation id="EF9A4B06A031BF63FEE776EBC164FD03" author="Chamberlin" box="[356,542,680,702]" pageId="5" pageNumber="41" refString="Chamberlin, R. V. (1921) Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, 97, 1 - 28." type="journal article" year="1921">Chamberlin, 1921</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
, Pl. 1,
<figureCitation id="13302A72A031BF63FD0A76EBC1BDFD03" box="[649,711,680,702]" captionStart="FIGURES 3 4" captionStartId="2.[151,264,1409,1432]" captionTargetBox="[152,1435,437,1388]" captionTargetId="figure@2.[152,1435,437,1388]" captionTargetPageId="2" captionText="FIGURES 3 4. Otostigmus (P.) brunneus. Holotype MCZ IZ no. 2181 (31416), male. 3. Cephalic plate and tergites 1 4. 4. Tergites of antepenult, penultimate and ultimate leg-bearing segments and prefemora of ultimate legs. DA-Digitiform appendix. Scale bars 1 mm." httpUri="https://zenodo.org/record/262566/files/figure.png" pageId="5" pageNumber="41">Fig. 3</figureCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB436F7A031BF63FF147684C04DFD46" blockId="5.[151,997,680,794]" pageId="5" pageNumber="41">
<treatmentCitationGroup id="AB1B11D9A031BF63FF147684C04DFD46" pageId="5" pageNumber="41">
<taxonomicName id="4C0B4D74A031BF63FF147684C1B7FD61" ID-CoL="75DNT" authority="Attems, 1930: 165" authorityName="Attems" authorityPageNumber="165" authorityYear="1930" box="[151,717,710,733]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="clavifer" subGenus="Parotostigmus">
<emphasis id="B97FEAE5A031BF63FF147684C271FD61" box="[151,267,711,732]" italics="true" pageId="5" pageNumber="41">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A031BF63FE9A768BC2D6FD60" box="[281,428,712,733]" italics="true" pageId="5" pageNumber="41">Parotostigmus</emphasis>
)
<emphasis id="B97FEAE5A031BF63FE397685C17DFD61" box="[442,519,710,732]" italics="true" pageId="5" pageNumber="41">clavifer</emphasis>
<treatmentCitation id="0AAA10E6A031BF63FD8D7684C1B7FD61" author="Attems" box="[526,717,710,732]" page="165" pageId="5" pageNumber="41" year="1930">
<bibRefCitation id="EF9A4B06A031BF63FD8D7684C1E0FD61" author="Attems" box="[526,666,710,732]" pageId="5" pageNumber="41" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich, Lief. 54, 1 - 308." type="journal article" year="1930">Attems, 1930</bibRefCitation>
: 165
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AAA10E6A031BF63FD547685C0E6FD61" author="Bucherl" box="[727,924,710,732]" page="112" pageId="5" pageNumber="41" year="1974">
<bibRefCitation id="EF9A4B06A031BF63FD547685C010FD61" author="Bucherl" box="[727,874,710,732]" pageId="5" pageNumber="41" refString="Bucherl, W. (1974) Die Scolopendromorpha der Neotropischen Region. Symposia of the Zoological Society of London, 32, 99 - 133." type="journal article" year="1974">Bücherl, 1974</bibRefCitation>
: 112
</treatmentCitation>
;
<taxonomicName id="4C0B4D74A031BF63FF1476A5C1B7FD46" ID-CoL="75DNT" authority="Bucherl, 1940: 223" authorityName="Bucherl" authorityPageNumber="223" authorityYear="1940" box="[151,717,741,763]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="clavifer" subGenus="Androstigmus">
<emphasis id="B97FEAE5A031BF63FF1476A5C271FD46" box="[151,267,742,763]" italics="true" pageId="5" pageNumber="41">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A031BF63FE9A76A6C2DFFD46" box="[281,421,741,763]" italics="true" pageId="5" pageNumber="41">Androstigmus</emphasis>
)
<emphasis id="B97FEAE5A031BF63FE3076A6C17AFD46" box="[435,512,741,763]" italics="true" pageId="5" pageNumber="41">clavifer</emphasis>
<treatmentCitation id="0AAA10E6A031BF63FD8476A6C1B7FD46" author="Bucherl" box="[519,717,741,763]" page="223" pageId="5" pageNumber="41" year="1940">
<bibRefCitation id="EF9A4B06A031BF63FD8476A6C1E0FD46" author="Bucherl" box="[519,666,741,763]" pageId="5" pageNumber="41" refString="Bucherl, W. (1940) Os quilopodos do Brasil. Memorias do Instituto Butantan, 13, 49 - 363." type="journal article" year="1940">Bücherl, 1940</bibRefCitation>
: 223
</treatmentCitation>
</taxonomicName>
; 1942: 60;
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB436F7A031BF63FF147747C09FFCA7" blockId="5.[151,997,680,794]" box="[151,997,771,794]" pageId="5" pageNumber="41">
<treatmentCitationGroup id="AB1B11D9A031BF63FF147747C09FFCA7" box="[151,997,771,794]" pageId="5" pageNumber="41">
<taxonomicName id="4C0B4D74A031BF63FF147747C185FCA7" ID-CoL="4B7X2" authority="Chamberlin, 1921: 13" authorityName="Chamberlin" authorityPageNumber="13" authorityYear="1921" box="[151,767,772,794]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Parotostigmus">
<emphasis id="B97FEAE5A031BF63FF147747C271FCA4" box="[151,267,772,793]" italics="true" pageId="5" pageNumber="41">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A031BF63FE9A7746C2D6FCA7" box="[281,428,773,794]" italics="true" pageId="5" pageNumber="41">Parotostigmus</emphasis>
)
<emphasis id="B97FEAE5A031BF63FE397747C162FCA7" box="[442,536,772,794]" italics="true" pageId="5" pageNumber="41">brunneus</emphasis>
<treatmentCitation id="0AAA10E6A031BF63FD9C7747C185FCA7" author="Chamberlin" box="[543,767,772,794]" page="13" pageId="5" pageNumber="41" year="1921">
<bibRefCitation id="EF9A4B06A031BF63FD9C7747C1A0FCA7" author="Chamberlin" box="[543,730,772,794]" pageId="5" pageNumber="41" refString="Chamberlin, R. V. (1921) Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, 97, 1 - 28." type="journal article" year="1921">Chamberlin, 1921</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
.
<emphasis id="B97FEAE5A031BF63FC8A7740C010FCA4" bold="true" box="[777,874,771,794]" pageId="5" pageNumber="41">Syn. nov.</emphasis>
(see above)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C311657CA031BF63FF147705C65CFC3E" pageId="5" pageNumber="41" type="materials_examined">
<paragraph id="8BB436F7A031BF63FF147705C65CFC3E" blockId="5.[151,1436,838,1151]" pageId="5" pageNumber="41">
<materialsCitation id="3B633CAAA031BF63FF147705C65CFC3E" ID-GBIF-Occurrence="1290984915" collectingDate="1914-08-24" collectionCode="MCZ" collectorName="F. M. Gaige" country="Guyana" location="Sand Hill Forest" pageId="5" pageNumber="41" specimenCount="2" stateProvince="Michigan" typeStatus="holotype">
<emphasis id="B97FEAE5A031BF63FF147705C2B1FCE2" bold="true" box="[151,459,838,863]" pageId="5" pageNumber="41">
<typeStatus id="54B08855A031BF63FF147705C3A8FCE2" box="[151,210,838,863]" pageId="5" pageNumber="41">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54B08855A031BF63FE5B7705C13AFCE2" box="[472,576,838,863]" pageId="5" pageNumber="41" type="holotype">Holotype</typeStatus>
<collectionCode id="ED1AAE32A031BF63FDCD7704C1F1FCE2" box="[590,651,839,863]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="41">MCZ</collectionCode>
IZ no. 2180 (14285),
<collectingCountry id="F31C7667A031BF63FC217704C083FCE2" box="[930,1017,839,863]" name="Guyana" pageId="5" pageNumber="41">Guyana</collectingCountry>
(British Guiana),
<location id="8ED4602CA031BF63FB5B7705C6E6FCE3" LSID="urn:lsid:plazi:treatment:03A287E1A031BF61FF14767EC1F1FE72:8ED4602CA031BF63FB5B7705C6E6FCE3" box="[1240,1436,838,863]" country="Guyana" name="Sand Hill Forest" pageId="5" pageNumber="41" stateProvince="Michigan">Sand Hill Forest</location>
Expedition: Bryant Walker Expedition (Uni. of
<collectingRegion id="49CFF815A031BF63FD2F7729C066FC3E" box="[684,796,874,899]" country="United States of America" name="Michigan" pageId="5" pageNumber="41">Michigan</collectingRegion>
), 1914,
<collectorName id="26FE5321A031BF63FCF67728C08CFC3E" box="[885,1014,874,899]" pageId="5" pageNumber="41">F.M. Gaige</collectorName>
,
<date id="FFB51037A031BF63FB827729C7F3FC3E" box="[1025,1161,874,899]" pageId="5" pageNumber="41" value="1914-08-24">
<collectingDate id="EFF1E9DFA031BF63FB827729C7F3FC3E" box="[1025,1161,874,899]" pageId="5" pageNumber="41" value="1914-08-24">24.viii.1914</collectingDate>
</date>
(
<figureCitation id="13302A72A031BF63FB147729C663FC3E" box="[1175,1305,874,899]" captionStart="FIGURES 11 12" captionStartId="5.[151,264,1981,2004]" captionTargetBox="[151,1436,1193,1960]" captionTargetId="figure@5.[151,1436,1193,1960]" captionTargetPageId="5" captionText="FIGURES 11 12. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 11. Habitus, (dorsal view). 12. Habitus, (ventral view). Scale bars 5 mm." httpUri="https://zenodo.org/record/262569/files/figure.png" pageId="5" pageNumber="41">Figs 1112</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311657CA031BF63FF4477CDC64FFC52" pageId="5" pageNumber="41" type="diagnosis">
<paragraph id="8BB436F7A031BF63FF4477CDC64FFC52" blockId="5.[151,1436,838,1151]" pageId="5" pageNumber="41">
<emphasis id="B97FEAE5A031BF63FF4477CDC23BFC1A" bold="true" box="[199,321,910,935]" pageId="5" pageNumber="41">Diagnosis.</emphasis>
Tergites smooth. Sternites 220 with short incomplete paramedian sutures; sternites without depressions. Legs 520 with one tarsal spur. Prefemur of ultimate legs of males with a digitiform appendix the same length as the prefemur; digitiform appendix with a tuft of hairs located in the median side of the tip.
</paragraph>
</subSubSection>
<subSubSection id="C311657CA031BF60FF4477B9C799F830" lastPageId="6" lastPageNumber="42" pageId="5" pageNumber="41" type="description">
<paragraph id="8BB436F7A031BF63FF4477B9C160FBC2" blockId="5.[151,1436,838,1151]" pageId="5" pageNumber="41">
<emphasis id="B97FEAE5A031BF63FF4477B9C2BDFBAE" bold="true" box="[199,455,1018,1043]" pageId="5" pageNumber="41">Redescription (male).</emphasis>
Length:
<quantity id="4CF39B12A031BF63FDA877B9C100FBAF" box="[555,634,1018,1043]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="41" unit="mm" value="53.0">53 mm</quantity>
from the anterior margin of the cephalic plate to the posterior margin of tergite 21. Antennae: left antenna with 23 articles and right with 20 (20 articles in both antennae); two basal articles glabrous (third and fourth basal articles glabrous). Cephalic plate smooth, without sutures and depression, but with median sulcus anteriorly (
<figureCitation id="13302A72A031BF63FE3A7025C176FBC2" box="[441,524,1126,1151]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1168,1191]" captionTargetBox="[151,1435,193,1147]" captionTargetId="figure@6.[151,1435,193,1147]" captionTargetPageId="6" captionText="FIGURES 13 14. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 13. Cephalic plate and tergites 1 3. 14. Tergite of ultimate leg-bearing segment and prefemur of ultimate legs showing the digitiform appendix. DA—Digitiform appendix; TH—Tuft of hairs. Scale bars 1 mm." httpUri="https://zenodo.org/record/262570/files/figure.png" pageId="5" pageNumber="41">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF74667FA031BF63FF1473FEC29BF84F" httpUri="https://zenodo.org/record/262569/files/figure.png" pageId="5" pageNumber="41" targetBox="[151,1436,1193,1960]" targetPageId="5">
<paragraph id="8BB436F7A031BF63FF1473FEC29BF84F" blockId="5.[151,1436,1981,2034]" pageId="5" pageNumber="41">
<emphasis id="B97FEAE5A031BF63FF1473FEC236F86E" bold="true" box="[151,332,1981,2004]" pageId="5" pageNumber="41">FIGURES 1112</emphasis>
.
<taxonomicName id="4C0B4D74A031BF63FEDA73FCC133F869" box="[345,585,1982,2004]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="41" phylum="Arthropoda" rank="species" species="clavifer" subGenus="Parotostigmus">
<emphasis id="B97FEAE5A031BF63FEDA73FCC2B7F869" box="[345,461,1983,2004]" italics="true" pageId="5" pageNumber="41">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A031BF63FE5873FCC292F869" box="[475,488,1983,2004]" italics="true" pageId="5" pageNumber="41">P</emphasis>
.)
<emphasis id="B97FEAE5A031BF63FE7E73FDC133F869" box="[509,585,1982,2004]" italics="true" pageId="5" pageNumber="41">clavifer</emphasis>
</taxonomicName>
. Holotype MCZ IZ no.2180 (14285), male.
<emphasis id="B97FEAE5A031BF63FB8F73FDC758F86E" bold="true" box="[1036,1058,1982,2003]" pageId="5" pageNumber="41">11</emphasis>
. Habitus, (dorsal view).
<emphasis id="B97FEAE5A031BF63FAA373FDC643F86E" bold="true" box="[1312,1337,1982,2003]" pageId="5" pageNumber="41">12</emphasis>
. Habitus, (ventral view). Scale bars 5 mm.
</paragraph>
</caption>
<caption id="DF74667FA032BF60FF1470D3C10CFB59" httpUri="https://zenodo.org/record/262570/files/figure.png" pageId="6" pageNumber="42" targetBox="[151,1435,193,1147]" targetPageId="6">
<paragraph id="8BB436F7A032BF60FF1470D3C10CFB59" blockId="6.[151,1435,1168,1252]" pageId="6" pageNumber="42">
<emphasis id="B97FEAE5A032BF60FF1470D3C237FB1B" bold="true" box="[151,333,1168,1191]" pageId="6" pageNumber="42">FIGURES 1314</emphasis>
.
<taxonomicName id="4C0B4D74A032BF60FED970D1C131FB1A" box="[346,587,1169,1191]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="clavifer" subGenus="Parotostigmus">
<emphasis id="B97FEAE5A032BF60FED970D1C2B7FB1A" box="[346,461,1170,1191]" italics="true" pageId="6" pageNumber="42">Otostigmus</emphasis>
(
<emphasis id="B97FEAE5A032BF60FE5E70D1C290FB1A" box="[477,490,1170,1191]" italics="true" pageId="6" pageNumber="42">P</emphasis>
.)
<emphasis id="B97FEAE5A032BF60FE7C70D2C131FB1A" box="[511,587,1169,1191]" italics="true" pageId="6" pageNumber="42">clavifer</emphasis>
</taxonomicName>
. Holotype MCZ IZ no.2180 (14285), male.
<emphasis id="B97FEAE5A032BF60FB8D70D2C75DFB1B" bold="true" box="[1038,1063,1169,1190]" pageId="6" pageNumber="42">13</emphasis>
. Cephalic plate and tergites 13.
<emphasis id="B97FEAE5A032BF60FAFF70D2C6EFFB1B" bold="true" box="[1404,1429,1169,1190]" pageId="6" pageNumber="42">14</emphasis>
. Tergite of ultimate leg-bearing segment and prefemur of ultimate legs showing the digitiform appendix. DA—Digitiform appendix; TH—Tuft of hairs. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BB436F7A032BF60FF447153C1EBFA04" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">Coxosternal tooth-plates wider than long with 4+4 teeth and with a strong seta on each plate; the two inner teeth closer to each other than to the external ones. Coxosternite with a transverse suture at the base of the toothplates and a very short longitudinal suture (sulcus at the base of dental plates at an obtuse angle at middle, from which a short median line extends caudad). Margin of the forcipular trochanteroprefemoral process with four short denticles, and the tip pointed (not described).</paragraph>
<paragraph id="8BB436F7A032BF60FF447186C224F9F4" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">
Tergites smooth (all smooth and shining); Tergites 620 with complete paramedian sutures (from 5); tergites 25 with incomplete very short anterior and posterior sutures; tergites 921 with poorly-developed margination, not clearly visible (only 21 marginate); tergite 21 with posterior margin convex, angled (obtusely angular at middle) (
<figureCitation id="13302A72A032BF60FF7E7273C22BF9F4" box="[253,337,1584,1609]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1168,1191]" captionTargetBox="[151,1435,193,1147]" captionTargetId="figure@6.[151,1435,193,1147]" captionTargetPageId="6" captionText="FIGURES 13 14. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 13. Cephalic plate and tergites 1 3. 14. Tergite of ultimate leg-bearing segment and prefemur of ultimate legs showing the digitiform appendix. DA—Digitiform appendix; TH—Tuft of hairs. Scale bars 1 mm." httpUri="https://zenodo.org/record/262570/files/figure.png" pageId="6" pageNumber="42">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BB436F7A032BF60FF447216C045F90B" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">Sternites 220 with short incomplete paramedian sutures; those of the anterior and posterior sternites shorter than in the mid body (generally without distinct furrows); sternites without depressions; sternite 21 shorter than preceding, with posterior margin straight (slightly incurved).</paragraph>
<paragraph id="8BB436F7A032BF60FF447283C380F943" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">Coxopleuron without spines and with pore-field that covers almost all the coxopleuron, only the posterior end poreless.</paragraph>
<paragraph id="8BB436F7A032BF60FF44734AC799F830" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">
Legs 2 with two tarsal spurs (1 to 4); right legs 10, 11, 18, 19 and 20 with one tarsal spur, and 21 without; left legs 8 and 19 with one tarsal spur, and 20 and 21 without (5 to 20). The remaining legs are missing. Prefemur of ultimate legs with a digitiform appendix the same length as the prefemur; the tip of the digitiform appendix has a tuft of hairs located in the median side of the tip (distal end of dorsal or submedian side) (
<figureCitation id="13302A72A032BF60FB007336C7ACF830" box="[1155,1238,1909,1934]" captionStart="FIGURES 13 14" captionStartId="6.[151,264,1168,1191]" captionTargetBox="[151,1435,193,1147]" captionTargetId="figure@6.[151,1435,193,1147]" captionTargetPageId="6" captionText="FIGURES 13 14. Otostigmus (P.) clavifer. Holotype MCZ IZ no. 2180 (14285), male. 13. Cephalic plate and tergites 1 3. 14. Tergite of ultimate leg-bearing segment and prefemur of ultimate legs showing the digitiform appendix. DA—Digitiform appendix; TH—Tuft of hairs. Scale bars 1 mm." httpUri="https://zenodo.org/record/262570/files/figure.png" pageId="6" pageNumber="42">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C311657CA032BF61FF4473DBC1F1FE72" lastPageId="7" lastPageNumber="43" pageId="6" pageNumber="42" type="discussion">
<paragraph id="8BB436F7A032BF60FF4473DBC6E6F869" blockId="6.[151,1436,1296,2006]" pageId="6" pageNumber="42">
<emphasis id="B97FEAE5A032BF60FF4473DBC241F80C" bold="true" box="[199,315,1944,1969]" pageId="6" pageNumber="42">Remarks.</emphasis>
<taxonomicName id="4C0B4D74A032BF60FEC973DAC2B3F80C" box="[330,457,1945,1969]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A032BF60FEC973DAC2B3F80C" box="[330,457,1945,1969]" italics="true" pageId="6" pageNumber="42">O. clavifer</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EF9A4B06A032BF60FD3973DBC0E2F80C" author="Chamberlin" box="[698,920,1944,1969]" pageId="6" pageNumber="42" refString="Chamberlin, R. V. (1921) Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, 97, 1 - 28." type="journal article" year="1921">Chamberlin (1921)</bibRefCitation>
as a species of
<taxonomicName id="4C0B4D74A032BF60FBF273DAC78BF80D" box="[1137,1265,1945,1968]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEAE5A032BF60FBF273DAC78BF80D" box="[1137,1265,1945,1968]" italics="true" pageId="6" pageNumber="42">Otostigmus</emphasis>
</taxonomicName>
related to
<taxonomicName id="4C0B4D74A032BF60FA0273DAC258F868" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="42" phylum="Arthropoda" rank="species" species="scabricauda">
<emphasis id="B97FEAE5A032BF60FA0273DAC258F868" italics="true" pageId="6" pageNumber="42">O. scabricauda</emphasis>
</taxonomicName>
by the digitiform appendix of the prefemur of the ultimate legs. However, it differs from
<emphasis id="B97FEAE5A032BF60FA0273FEC6E6F869" box="[1409,1436,1981,2004]" italics="true" pageId="6" pageNumber="42">O.</emphasis>
</paragraph>
<paragraph id="8BB436F7A033BF61FF1474DBC1F1FE72" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="43">
<taxonomicName id="4C0B4D74A033BF61FF1474DBC258FF0D" box="[151,290,152,176]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="scabricauda">
<emphasis id="B97FEAE5A033BF61FF1474DBC258FF0D" box="[151,290,152,176]" italics="true" pageId="7" pageNumber="43">scabricauda</emphasis>
</taxonomicName>
by the smooth tergites, absence of keels on tergites, and the absence of a median depression on sternites.
<bibRefCitation id="EF9A4B06A033BF61FE8674FFC2A4FF69" author="Chamberlin" box="[261,478,188,213]" pageId="7" pageNumber="43" refString="Chamberlin, R. V. (1921) Results of the Bryant Walker Expeditions of the University of the Michigan to Colombia, 1913, and British Guiana, 1914. Occasional Papers of the Museum of Zoology, 97, 1 - 28." type="journal article" year="1921">Chamberlin (1921)</bibRefCitation>
also noticed that the digitiform appendix in the prefemur of the ultimate legs was different from the digitiform appendix of
<taxonomicName id="4C0B4D74A033BF61FDE674A3C06AFF45" box="[613,784,224,248]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="scabricauda">
<emphasis id="B97FEAE5A033BF61FDE674A3C06AFF45" box="[613,784,224,248]" italics="true" pageId="7" pageNumber="43">O. scabricauda</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C0B4D74A033BF61FCBF74A3C0C8FF45" box="[828,946,224,248]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A033BF61FCBF74A3C0C8FF45" box="[828,946,224,248]" italics="true" pageId="7" pageNumber="43">O. clavifer</emphasis>
</taxonomicName>
the digitiform appendix is slightly flattened dorsoventrally and the tuft of hair emerges on the median side of the tip. In
<taxonomicName id="4C0B4D74A033BF61FC627547C7F4FEA1" box="[993,1166,260,284]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="scabricauda">
<emphasis id="B97FEAE5A033BF61FC627547C7F4FEA1" box="[993,1166,260,284]" italics="true" pageId="7" pageNumber="43">O. scabricauda</emphasis>
</taxonomicName>
the digitiform appendix is also slightly flattened dorsoventrally, but the tuft of hairs emerges at the tip, not on the median side of the tip. Therefore, the
<typeStatus id="54B08855A033BF61FEBE750EC217FED8" box="[317,365,333,357]" pageId="7" pageNumber="43">type</typeStatus>
of
<taxonomicName id="4C0B4D74A033BF61FE10750FC176FED9" box="[403,524,332,356]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A033BF61FE10750FC176FED9" box="[403,524,332,356]" italics="true" pageId="7" pageNumber="43">O. clavifer</emphasis>
</taxonomicName>
is more related to
<taxonomicName id="4C0B4D74A033BF61FD5D750FC013FED9" box="[734,873,332,356]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B97FEAE5A033BF61FD5D750FC013FED9" box="[734,873,332,356]" italics="true" pageId="7" pageNumber="43">O. brunneus</emphasis>
</taxonomicName>
than
<taxonomicName id="4C0B4D74A033BF61FC2B750FC72FFED9" box="[936,1109,332,356]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="scabricauda">
<emphasis id="B97FEAE5A033BF61FC2B750FC72FFED9" box="[936,1109,332,356]" italics="true" pageId="7" pageNumber="43">O. scabricauda</emphasis>
</taxonomicName>
, the only difference between them being the length of the digitiform appendix of the prefemur of the ultimate legs. As
<taxonomicName id="4C0B4D74A033BF61FBFC7533C78DFE35" box="[1151,1271,368,392]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A033BF61FBFC7533C78DFE35" box="[1151,1271,368,392]" italics="true" pageId="7" pageNumber="43">O. clavifer</emphasis>
</taxonomicName>
was described in the same work, but one page before
<taxonomicName id="4C0B4D74A033BF61FDEE75D7C186FE11" box="[621,764,404,428]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B97FEAE5A033BF61FDEE75D7C186FE11" box="[621,764,404,428]" italics="true" pageId="7" pageNumber="43">O. brunneus</emphasis>
</taxonomicName>
, and both are from the same
<typeStatus id="54B08855A033BF61FBE275D6C7EBFE10" box="[1121,1169,405,429]" pageId="7" pageNumber="43">type</typeStatus>
locality, I consider
<taxonomicName id="4C0B4D74A033BF61FA0275D7C385FE6D" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B97FEAE5A033BF61FA0275D7C385FE6D" italics="true" pageId="7" pageNumber="43">O. brunneus</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C0B4D74A033BF61FD8E75FBC1FEFE6D" box="[525,644,440,464]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="43" phylum="Arthropoda" rank="species" species="clavifer">
<emphasis id="B97FEAE5A033BF61FD8E75FBC1FEFE6D" box="[525,644,440,464]" italics="true" pageId="7" pageNumber="43">O. clavifer</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>