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<document id="E65E8FAC6ADEA562AB4E2D86B51B9F40" ID-CLB-Dataset="24132" ID-DOI="10.1080/00222933.2018.1554166" ID-GBIF-Dataset="bd9add32-702b-495c-9ed7-7ed98fb4c591" ID-ISSN="1464-5262" ID-Zenodo-Dep="3654165" ID-ZooBank="FA2EDE40-93E0-43B7-9ACF-ECCE12B0E671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581079052408" checkinUser="valdenar" docAuthor="Morozov, Grigori, Sabirov, Rushan &amp; Zimina, Olga" docDate="2019" docId="03A287D2FC6141539BC1F94734943147" docLanguage="en" docName="JNATHIST.52.4748.29612992.pdf.imf" docOrigin="Journal of Natural History 52 (47)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Suberites montalbidus Carter 1880" docType="treatment" docVersion="12" lastPageNumber="2992" masterDocId="FF9BFFAAFC6C415C9A7AFFC33570355D" masterDocTitle="Sponge fauna of the New Siberian Shoal: biodiversity and some features of formation" masterLastPageNumber="2992" masterPageNumber="2961" pageNumber="2961" updateTime="1698780954505" updateUser="ExternalLinkService">
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<mods:title id="60A339A40E5BC30A34FC4540AF86AC95">Sponge fauna of the New Siberian Shoal: biodiversity and some features of formation</mods:title>
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<taxonomicName id="4C0B4D47FC6141519BC1F947361733C0" authority="Carter, 1880" authorityName="Carter" authorityYear="1880" box="[443,871,1668,1693]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="13" phylum="Porifera" rank="species" species="montalbidus">
<emphasis id="B97FEAD6FC6141519BC1F94737BE33C0" bold="true" box="[443,718,1668,1693]" italics="true" pageId="13">Suberites montalbidus</emphasis>
<bibRefCitation id="EF9A4B35FC61415198A2F947361733C0" author="Carter HJ" box="[728,871,1668,1693]" pageId="13" pagination="253 - 277" refId="ref12337" refString="Carter HJ. 1880. Description of two new Sponges. Ann Mag Nat Hist. 5 (6): 253 - 277." type="journal article" year="1880">Carter, 1880</bibRefCitation>
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(
<figureCitation id="13302A41FC614151983BF96537D533E2" box="[577,677,1702,1727]" captionStart="Figure 8" captionStartId="14.[160,225,893,916]" captionTargetBox="[163,1153,142,862]" captionTargetId="figure@14.[163,1153,142,862]" captionTargetPageId="14" captionText="Figure 8. suberites montalbidus: (a, b) longitudinal sections through the surface (a) and inner part (b) of the sponge; (c, d) habitus; (eh) scanning electron microscope (SEM) images of spicules; (e) large tylostyle; (f) small tylostyle; (g, h) microxea and microstrongyles, respectively." figureDoi="http://doi.org/10.5281/zenodo.3654201" httpUri="https://zenodo.org/record/3654201/files/figure.png" pageId="13">Figure 8</figureCitation>
(a
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h))
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<emphasis id="B97FEAD6FC6241529ADAFBDA34E7316C" box="[160,407,1049,1073]" italics="true" pageId="14">suberites montalbidus</emphasis>
<treatmentCitation id="0AAA10D5FC6241529BDEFBDA37FB316C" box="[420,651,1049,1073]" pageId="14">
<bibRefCitation id="EF9A4B35FC6241529BDEFBDA375C316C" author="Carter HJ" box="[420,556,1049,1073]" pageId="14" pagination="253 - 277" refId="ref12337" refString="Carter HJ. 1880. Description of two new Sponges. Ann Mag Nat Hist. 5 (6): 253 - 277." type="journal article" year="1880">Carter 1880</bibRefCitation>
, p. 256
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;
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<bibRefCitation id="EF9A4B35FC62415298E1FBDA3644316C" author="Fristedt K" box="[667,820,1049,1073]" pageId="14" pagination="401 - 471" refId="ref12790" refString="Fristedt K. 1887. Sponges from the Atlantic and Arctic Oceans and the Behring Sea. Vega- Expeditionens Vetenskap Iakttagelser (Nordenskiold). 4: 401 - 471." type="journal article" year="1887">Fristedt 1887</bibRefCitation>
, p. 428
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429
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;
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<bibRefCitation id="EF9A4B35FC62415299A0FBDA31F3316C" author="Swartschewsky BA" box="[986,1155,1049,1073]" pageId="14" pagination="307 - 371" refId="ref14895" refString="Swartschewsky BA. 1906. Beitrage zur Spongien-Fauna des Weissen Meeres. Mem Soci Nat Kiew. 20: 307 - 371. German." type="journal article" year="1906">Swartschewsky</bibRefCitation>
1906, p. 318
<emphasis id="B97FEAD6FC6241529B36FBF83429310E" box="[332,345,1083,1107]" italics="true" pageId="14"></emphasis>
319, pl. XIII, fig. 3
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<emphasis id="B97FEAD6FC6241529ADAFB9D3471312B" box="[160,257,1118,1142]" italics="true" pageId="14">suberites</emphasis>
sp.:
<treatmentCitation id="0AAA10D5FC6241529B42FB9D36FD31C4" pageId="14">
<bibRefCitation id="EF9A4B35FC6241529B42FB9D34A9312B" author="Vosmaer GCJ" box="[312,473,1118,1142]" pageId="14" pagination="1 - 58" refId="ref15169" refString="Vosmaer GCJ. 1882. Report on the sponges dredged up in the Arctic Sea by the ' Willem Barents ' in the years 1878 and 1879. Niederlandisches Archiv fur Zoologie Supplement. 1 (3): 1 - 58." type="journal article" year="1882">Vosmaer 1882</bibRefCitation>
, p. 32
<emphasis id="B97FEAD6FC624152985AFB9D375D312B" box="[544,557,1118,1142]" italics="true" pageId="14"></emphasis>
33; pl. I, figs 22
<emphasis id="B97FEAD6FC62415298A1FB9D3798312B" box="[731,744,1118,1142]" italics="true" pageId="14"></emphasis>
23; pl. IV, figs 140
<emphasis id="B97FEAD6FC62415299C9FB9D36B0312B" box="[947,960,1118,1142]" italics="true" pageId="14"></emphasis>
144
<taxonomicName id="4C0B4D47FC6241529ADAFB42375D31C4" authority=": Lambe 1895" authorityName=": Lambe" authorityYear="1895" box="[160,557,1153,1177]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="14" phylum="Porifera" rank="species" species="montalbidus">
<emphasis id="B97FEAD6FC6241529ADAFB4234E231C4" box="[160,402,1153,1177]" italics="true" pageId="14">suberites montalbidus</emphasis>
:
<bibRefCitation id="EF9A4B35FC6241529BDBFB42375D31C4" box="[417,557,1153,1177]" pageId="14" refId="ref13725" type="journal article">Lambe 1895</bibRefCitation>
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(?), p. 127
<emphasis id="B97FEAD6FC62415298DEFB4237C131C4" box="[676,689,1153,1177]" italics="true" pageId="14"></emphasis>
128; pl. III, fig. 6a
<emphasis id="B97FEAD6FC624152990EFB4236F131C4" box="[884,897,1153,1177]" italics="true" pageId="14"></emphasis>
c
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<emphasis id="B97FEAD6FC6241529ADAFB6734CC31E1" box="[160,444,1187,1212]" italics="true" pageId="14">
<taxonomicName id="4C0B4D47FC6241529ADAFB6734F431E1" baseAuthorityName="Olivi" baseAuthorityYear="1792" box="[160,388,1188,1212]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="14" phylum="Porifera" rank="species" species="domuncula">suberites domuncula</taxonomicName>
<emphasis id="B97FEAD6FC6241529BF1FB6034E831E6" bold="true" box="[395,408,1187,1211]" italics="true" pageId="14">fi</emphasis>
cus
</emphasis>
:
<treatmentCitation id="0AAA10D5FC6241529BB3FB6734693183" pageId="14">
<bibRefCitation id="EF9A4B35FC6241529BB3FB67372131E1" author="Koltun VM" box="[457,593,1188,1212]" pageId="14" pagination="1 - 263" refId="ref13426" refString="Koltun VM. 1959. Kрiмнiрogovыi gubki (iviрныk i daльнivo (tocныk мoрiй CCCР [Siliceous horny sponges of the northern and fareastern seas of the U. S. S. R.]. In: Pavlovsky EN, editor. Identifiers of the USSR fauna issued by the Zoological Institute of the Academy of Sciences of the USSR. Vol. 67 Russian. Moscow - Leningrad: Nauka, Academy of Sciences of the USSR; p. 1 - 263" type="book chapter" year="1959">Koltun 1959</bibRefCitation>
(?), p. 95, figs 66
<emphasis id="B97FEAD6FC6241529977FB60366A31E6" box="[781,794,1187,1211]" italics="true" pageId="14"></emphasis>
67; pl. XXXIV, figs 1
<emphasis id="B97FEAD6FC624152998BFB60368E31E6" box="[1009,1022,1187,1211]" italics="true" pageId="14"></emphasis>
3; pl. XXXVI, figs 1
<emphasis id="B97FEAD6FC6241529A84FB05347B3183" box="[254,267,1222,1246]" italics="true" pageId="14"></emphasis>
2
</treatmentCitation>
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</paragraph>
<caption id="DF74664CFC6241529ADAFCBE36DB3692" ID-DOI="http://doi.org/10.5281/zenodo.3654201" ID-Zenodo-Dep="3654201" httpUri="https://zenodo.org/record/3654201/files/figure.png" pageId="14" startId="14.[160,225,893,916]" targetBox="[163,1153,142,862]" targetPageId="14">
<paragraph id="8BB436C4FC6241529ADAFCBE36DB3692" blockId="14.[160,1156,893,975]" pageId="14">
<emphasis id="B97FEAD6FC6241529ADAFCBE358A36C9" bold="true" box="[160,250,893,916]" pageId="14">Figure 8.</emphasis>
<taxonomicName id="4C0B4D47FC6241529B79FCBE34A536C9" authorityName="Carter" authorityYear="1880" box="[259,469,893,916]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="14" phylum="Porifera" rank="species" species="montalbidus">
<emphasis id="B97FEAD6FC6241529B79FCBE34A536C9" box="[259,469,893,916]" italics="true" pageId="14">suberites montalbidus</emphasis>
</taxonomicName>
: (a, b) longitudinal sections through the surface (a) and inner part (b) of the sponge; (c, d) habitus; (eh) scanning electron microscope (SEM) images of spicules; (e) large tylostyle; (f) small tylostyle; (g, h) microxea and microstrongyles, respectively.
</paragraph>
</caption>
<paragraph id="8BB436C4FC6241529ADAFADC34563064" blockId="14.[160,1156,1311,1614]" box="[160,294,1311,1337]" pageId="14">
<emphasis id="B97FEAD6FC6241529ADAFADC34563064" bold="true" box="[160,294,1311,1337]" italics="true" pageId="14">Description</emphasis>
</paragraph>
<paragraph id="8BB436C4FC6241529ADAFA8737773313" blockId="14.[160,1156,1311,1614]" pageId="14">
(
<figureCitation id="13302A41FC6241529AD2FA87347A3001" box="[168,266,1348,1372]" captionStart="Figure 8" captionStartId="14.[160,225,893,916]" captionTargetBox="[163,1153,142,862]" captionTargetId="figure@14.[163,1153,142,862]" captionTargetPageId="14" captionText="Figure 8. suberites montalbidus: (a, b) longitudinal sections through the surface (a) and inner part (b) of the sponge; (c, d) habitus; (eh) scanning electron microscope (SEM) images of spicules; (e) large tylostyle; (f) small tylostyle; (g, h) microxea and microstrongyles, respectively." figureDoi="http://doi.org/10.5281/zenodo.3654201" httpUri="https://zenodo.org/record/3654201/files/figure.png" pageId="14">Figure 8</figureCitation>
(c
<emphasis id="B97FEAD6FC6241529B64FA87345B3001" box="[286,299,1348,1372]" italics="true" pageId="14"></emphasis>
d)). Sponge elongated, pear shaped or conical, sometimes laterally flattened, up to
<quantity id="4CF39B21FC6241529B40FAA534F63023" box="[314,390,1382,1406]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="14" unit="cm" value="5.5">5.5 cm</quantity>
in height and
<quantity id="4CF39B21FC624152984CFAA537F33023" box="[566,643,1382,1406]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="14" unit="cm" value="4.5">4.5 cm</quantity>
in width. Body slightly narrows downwards forming a short peduncle, by which the sponge attaches on stones or on polychaetes tubes. The surface is rugose, sometimes nearly smooth (only juvenile forms). The consistency is soft and elastic. The single osculum of roundish or slit-like form (about
<quantity id="4CF39B21FC6241529ADAFA3235983354" box="[160,232,1521,1545]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" unit="mm" value="2.0">2 mm</quantity>
in diameter) is placed at the summit and surrounded by the short (about
<quantity id="4CF39B21FC6241529ADAF9D7358A3371" box="[160,250,1556,1580]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" unit="mm" value="0.8">0.8 mm</quantity>
in height) spicular collar (not always). Colour (in alcohol) ash grey or pale pink. Six specimens examined.
</paragraph>
<paragraph id="8BB436C4FC6341539ADAFF4E347535FA" blockId="15.[160,1156,141,410]" box="[160,261,141,167]" pageId="15">
<emphasis id="B97FEAD6FC6341539ADAFF4E347535FA" bold="true" box="[160,261,141,167]" italics="true" pageId="15">Skeleton</emphasis>
</paragraph>
<paragraph id="8BB436C4FC6341539ADAFF713472346F" blockId="15.[160,1156,141,410]" pageId="15">
(
<figureCitation id="13302A41FC6341539AD2FF71347B3597" box="[168,267,178,202]" captionStart="Figure 8" captionStartId="14.[160,225,893,916]" captionTargetBox="[163,1153,142,862]" captionTargetId="figure@14.[163,1153,142,862]" captionTargetPageId="14" captionText="Figure 8. suberites montalbidus: (a, b) longitudinal sections through the surface (a) and inner part (b) of the sponge; (c, d) habitus; (eh) scanning electron microscope (SEM) images of spicules; (e) large tylostyle; (f) small tylostyle; (g, h) microxea and microstrongyles, respectively." figureDoi="http://doi.org/10.5281/zenodo.3654201" httpUri="https://zenodo.org/record/3654201/files/figure.png" pageId="15">Figure 8</figureCitation>
(a
<emphasis id="B97FEAD6FC6341539B5AFF71345D3597" box="[288,301,178,202]" italics="true" pageId="15"></emphasis>
b)). The main skeleton consists of a quite loose or diffuse network of spicule bundles and single spicules. Only near the surface, spicules (mostly small tylostyles) become more or less regularly arranged, and grouped in the radial bundles.
</paragraph>
<paragraph id="8BB436C4FC6341539ABAFEFF377234C7" blockId="15.[160,1156,141,410]" pageId="15">The cortex is composed of a thin layer of microscleres. In the underlying tissues (choanosome) microscleres are absent. Only in rare exceptions were they found in the walls of the aquiferous system.</paragraph>
<paragraph id="8BB436C4FC6341539ADAFE08358F34B8" blockId="15.[160,1156,459,658]" box="[160,255,459,485]" pageId="15">
<emphasis id="B97FEAD6FC6341539ADAFE08358F34B8" bold="true" box="[160,255,459,485]" italics="true" pageId="15">Spicules</emphasis>
</paragraph>
<paragraph id="8BB436C4FC6341539ADAFE2C344337CF" blockId="15.[160,1156,459,658]" pageId="15">
(
<figureCitation id="13302A41FC6341539AD2FE2C347E375A" box="[168,270,495,519]" captionStart="Figure 8" captionStartId="14.[160,225,893,916]" captionTargetBox="[163,1153,142,862]" captionTargetId="figure@14.[163,1153,142,862]" captionTargetPageId="14" captionText="Figure 8. suberites montalbidus: (a, b) longitudinal sections through the surface (a) and inner part (b) of the sponge; (c, d) habitus; (eh) scanning electron microscope (SEM) images of spicules; (e) large tylostyle; (f) small tylostyle; (g, h) microxea and microstrongyles, respectively." figureDoi="http://doi.org/10.5281/zenodo.3654201" httpUri="https://zenodo.org/record/3654201/files/figure.png" pageId="15">Figure 8</figureCitation>
(e
<emphasis id="B97FEAD6FC6341539B59FE2C3440375A" box="[291,304,495,519]" italics="true" pageId="15"></emphasis>
h)). Styles (tylo- and subtylostyles) straight, rather short-pointed (spicules with blunt apical ends occasionally found), dimensions: 167.7
<emphasis id="B97FEAD6FC6341539E53FDD131463777" box="[1065,1078,530,554]" italics="true" pageId="15"></emphasis>
387.8
<emphasis id="B97FEAD6FC6341539E0CFDD131F33777" box="[1142,1155,530,554]" italics="true" pageId="15"></emphasis>
538.9 × 5.4
<emphasis id="B97FEAD6FC6341539B50FDF734473711" box="[298,311,564,588]" italics="true" pageId="15"></emphasis>
11 (
<emphasis id="B97FEAD6FC6341539B13FDF634073710" box="[361,375,565,589]" italics="true" pageId="15">n</emphasis>
= 200) µm; microxea and microstrongyles centrotylote, spined, dimensions 16.7
<emphasis id="B97FEAD6FC6341539B16FD9434093732" box="[364,377,599,623]" italics="true" pageId="15"></emphasis>
38.29
<emphasis id="B97FEAD6FC6341539BC3FD9434B63732" box="[441,454,599,623]" italics="true" pageId="15"></emphasis>
62.9 (
<emphasis id="B97FEAD6FC634153986CFD9437543732" box="[534,548,599,623]" italics="true" pageId="15">n</emphasis>
= 60) µm and 8.2
<emphasis id="B97FEAD6FC6341539947FD94363A3732" box="[829,842,599,623]" italics="true" pageId="15"></emphasis>
19.57
<emphasis id="B97FEAD6FC63415399F3FD9436E63732" box="[905,918,599,623]" italics="true" pageId="15"></emphasis>
36 (
<emphasis id="B97FEAD6FC63415399A8FD9436903732" box="[978,992,599,623]" italics="true" pageId="15">n</emphasis>
= 60) µm, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C311654FFC6341539ADAFD0036D7367F" pageId="15" type="distribution">
<paragraph id="8BB436C4FC6341539ADAFD00345C3780" blockId="15.[160,1157,707,802]" box="[160,300,707,733]" pageId="15">
<emphasis id="B97FEAD6FC6341539ADAFD00345C3780" bold="true" box="[160,300,707,733]" italics="true" pageId="15">Distribution</emphasis>
</paragraph>
<paragraph id="8BB436C4FC6341539ADAFD2436D7367F" blockId="15.[160,1157,707,802]" pageId="15">
Bering Sea, West and East
<collectingCountry id="F31C7654FC6341539BA9FD24373737A2" box="[467,583,743,767]" name="Greenland" pageId="15">Greenland</collectingCountry>
Shelf. Chukchi Sea,
<location id="8ED4601FFC634153994BFD24368537A2" LSID="urn:lsid:plazi:treatment:03A287D2FC6141539BC1F94734943147:8ED4601FFC634153994BFD24368537A2" box="[817,1013,743,767]" municipality="North Sea" name="East Siberian Sea" pageId="15">East Siberian Sea</location>
, Laptev Sea, Kara Sea, Barents Sea, White Sea,
<collectingMunicipality id="6BD0ACBEFC6341539858FCC937E5367F" box="[546,661,778,802]" pageId="15">North Sea</collectingMunicipality>
. Depth range:
<quantity id="4CF39B21FC6341539944FCC936D3367F" box="[830,931,778,802]" metricMagnitude="1" metricUnit="m" metricValue="5.9" metricValueMax="11.3" metricValueMin="0.5" pageId="15" unit="m" value="59.0" valueMax="113.0" valueMin="5.0">
5
<emphasis id="B97FEAD6FC6341539936FCC93629367F" box="[844,857,778,802]" italics="true" pageId="15"></emphasis>
<specimenCount id="9D0DFD4DFC6341539923FCC936D3367F" box="[857,931,778,802]" pageId="15" type="generic">113 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C311654FFC6341539ADAFC9034943147" pageId="15" type="discussion">
<paragraph id="8BB436C4FC6341539ADAFC9034763630" blockId="15.[160,1157,851,1050]" box="[160,262,851,877]" pageId="15">
<emphasis id="B97FEAD6FC6341539ADAFC9034763630" bold="true" box="[160,262,851,877]" italics="true" pageId="15">Remarks</emphasis>
</paragraph>
<paragraph id="8BB436C4FC6341539ADAFCB434943147" blockId="15.[160,1157,851,1050]" pageId="15">
Widespread circumpolar boreal-Arctic species. Quite polymorphic, and is represented by at least two morphologically close species. It is possible that some specimens of
<taxonomicName id="4C0B4D47FC6341539B28FC7E37263688" authorityName=": Lambe" authorityYear="1895" box="[338,598,957,981]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="15" phylum="Porifera" rank="species" species="montalbidus">
<emphasis id="B97FEAD6FC6341539B28FC7E37263688" box="[338,598,957,981]" italics="true" pageId="15">suberites montalbidus</emphasis>
</taxonomicName>
examined by
<bibRefCitation id="EF9A4B35FC634153996FFC7E36B73688" author="Fristedt K" box="[789,967,957,981]" pageId="15" pagination="401 - 471" refId="ref12790" refString="Fristedt K. 1887. Sponges from the Atlantic and Arctic Oceans and the Behring Sea. Vega- Expeditionens Vetenskap Iakttagelser (Nordenskiold). 4: 401 - 471." type="journal article" year="1887">Fristedt (1887)</bibRefCitation>
, Lambe (1895) and
<bibRefCitation id="EF9A4B35FC6341539AA3FC1C340C36AA" author="Koltun VM" box="[217,380,991,1015]" pageId="15" pagination="1 - 263" refId="ref13426" refString="Koltun VM. 1959. Kрiмнiрogovыi gubki (iviрныk i daльнivo (tocныk мoрiй CCCР [Siliceous horny sponges of the northern and fareastern seas of the U. S. S. R.]. In: Pavlovsky EN, editor. Identifiers of the USSR fauna issued by the Zoological Institute of the Academy of Sciences of the USSR. Vol. 67 Russian. Moscow - Leningrad: Nauka, Academy of Sciences of the USSR; p. 1 - 263" type="book chapter" year="1959">Koltun (1959)</bibRefCitation>
are actually represented by a similar, newly allocated species,
<taxonomicName id="4C0B4D47FC6341539ADAFBC1340C3147" authority="Morozov &amp; Sabirov &amp; Zimina, 2019" authorityName="Morozov &amp; Sabirov &amp; Zimina" authorityYear="2019" box="[160,380,1026,1050]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="15" phylum="Porifera" rank="species" species="cebriones" status="sp. nov.">
<emphasis id="B97FEAD6FC6341539ADAFBC1340C3147" box="[160,380,1026,1050]" italics="true" pageId="15">suberites cebriones</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57ADFC6341539BFCFBC134943147" box="[390,484,1026,1050]" pageId="15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>