234 lines
25 KiB
XML
234 lines
25 KiB
XML
<document id="04AF6BA234299C85E6C33774FBFE78BD" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA783FFFA4FEECFCF698A1F89F" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra ubicki Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="44" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="43" updateTime="1698190273211" updateUser="plazi">
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<mods:mods id="9C845BB7D2DC9C33DA42878D99378AD3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6CBD4CDBBBB007C01685A8BF0BEE14D8">
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<mods:title id="66516413B254B6B7BB4CF22488A3A5CE">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="1A200226119449896EAD50B4930160FE" type="personal">
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<mods:role id="3C00D2C67A61D81A43AEBEADFD849E6C">
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<mods:roleTerm id="2045726FE6DD2FEE497CB5672C648F19">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2D504D7F85B503D27C739348D8824300">Erwin, Terry L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="14155B605D08092AFF7FB3C79E6B6D1F">text</mods:typeOfResource>
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<mods:relatedItem id="639F20D377ADA13A197C03BA5CAEC814" type="host">
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<mods:titleInfo id="B4F5F4B94AAB57156FA73050C4200BF6">
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<mods:title id="3DF8DAD6140E2D135B432B8083C4FC8A">Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part id="131868792F906EEC05FCA4C7297A3503">
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<mods:date id="C0ED0F07C0B37FE12DBA417527A1FCD6">2002</mods:date>
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<mods:detail id="3025D8C49D3CE6880DE0D3FC6F539F60" type="volume">
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||
<mods:number id="F7461C731637AA2FF1358DECB5EE7666">119</mods:number>
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</mods:detail>
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<mods:extent id="5734EC8A18F1EFEB5CE72C8DCC0615AC" unit="page">
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<mods:start id="20890F25248DA7B49A6E6C5762FCCC27">1</mods:start>
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<mods:end id="A91B1BF7BC22043FA341A3AA3DCCFD8B">68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="CA51F5D5508B6A611698417710DAAD35">journal article</mods:classification>
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<mods:identifier id="9A1235A818557F33EEBB10E543C7F1BF" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="317A44DCEF6EFE4D208C8F015F99EEA8" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="5DA6D59E23860BB92D55540232C25FEF" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="88261F70054EC789F4058DD29BC7B989" type="Zenodo-Dep">162803</mods:identifier>
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</mods:mods>
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<treatment id="03A287CA783FFFA4FEECFCF698A1F89F" ID-DOI="http://doi.org/10.5281/zenodo.6278001" ID-GBIF-Taxon="119330620" ID-Zenodo-Dep="6278001" LSID="urn:lsid:plazi:treatment:03A287CA783FFFA4FEECFCF698A1F89F" httpUri="http://treatment.plazi.org/id/03A287CA783FFFA4FEECFCF698A1F89F" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<subSubSection id="C3116557783FFFA5FEECFCF69E02FC49" pageId="42" pageNumber="43" type="nomenclature">
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||
<paragraph id="8BB436DC783FFFA5FEECFCF69EBEFC72" blockId="42.[264,650,883,950]" box="[264,650,883,910]" pageId="42" pageNumber="43">
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||
<heading id="D0FC81B0783FFFA5FEECFCF69EBEFC72" bold="true" box="[264,650,883,910]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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||
<emphasis id="B97FEACE783FFFA5FEECFCF69EBEFC72" bold="true" box="[264,650,883,910]" pageId="42" pageNumber="43">
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<taxonomicName id="4C0B4D5F783FFFA5FEECFCF69DDAFC72" ID-CoL="B322" authority="Erwin" authorityName="Erwin" box="[264,494,883,910]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="ubicki" status="sp. nov.">
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<emphasis id="B97FEACE783FFFA5FEECFCF69DA3FC71" bold="true" box="[264,407,884,910]" italics="true" pageId="42" pageNumber="43">Agra ubicki</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B5783FFFA5FE1FFCF19EBEFC72" box="[507,650,883,909]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC783FFFA5FEECFC1E9E02FC49" blockId="42.[264,650,883,950]" box="[264,566,924,950]" pageId="42" pageNumber="43">
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(
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<figureCitation id="13302A59783FFFA5FEF5FC1E9D9EFC49" box="[273,426,924,950]" captionStart="FIGURE 18. A" captionStartId="43.[264,368,1163,1187]" captionTargetBox="[419,1166,412,1134]" captionTargetId="figure@43.[418,1169,409,1136]" captionTargetPageId="43" captionText="FIGURE 18. A. ubicki, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0." httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="42" pageNumber="43">Figs. 18a, 18</figureCitation>
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b, 18c, 18d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557783FFFA5FEECFC699E97FBD1" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="8BB436DC783FFFA5FEECFC699E97FBD1" blockId="42.[264,1326,1003,1870]" pageId="42" pageNumber="43">
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<emphasis id="B97FEACE783FFFA5FEECFC699DB5FBFA" bold="true" box="[264,385,1003,1029]" pageId="42" pageNumber="43">
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<typeStatus id="54B0887E783FFFA5FEECFC699D48FBFA" box="[264,380,1003,1029]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male,
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<emphasis id="B97FEACE783FFFA5FE33FC699EA3FBFA" bold="true" box="[471,663,1003,1029]" pageId="42" pageNumber="43">
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<collectingCountry id="F31C764C783FFFA5FE33FC699EA6FBFA" box="[471,658,1003,1029]" name="Costa Rica" pageId="42" pageNumber="43">COSTA RICA</collectingCountry>
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:
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</emphasis>
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Puntarenas,
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<quantity id="4CF39B39783FFFA5FCD5FC6E9F5BFBF9" box="[817,879,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="42" pageNumber="43" unit="km" value="3.0">3 km</quantity>
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NE Golfito,
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<geoCoordinate id="EE3F501B783FFFA5FBE9FC6E9851FBF9" box="[1037,1125,1004,1030]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="8.65">8º 39'N</geoCoordinate>
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||
<geoCoordinate id="EE3F501B783FFFA5FB8AFC6E98E7FBF9" box="[1134,1235,1004,1030]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-83.166664">83º10'W</geoCoordinate>
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,
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<quantity id="4CF39B39783FFFA5FB04FC6E9912FBF9" box="[1248,1318,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="100.0">100m</quantity>
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, May, (D. Ubick)(
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<collectionCode id="ED1AAE19783FFFA5FE32FB969E25FBD1" LSID="urn:lsid:biocol.org:col:34700" box="[470,529,1044,1070]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="42" pageNumber="43">CAS</collectionCode>
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: ADP4553).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557783FFFA5FEDCFBB99E3DFB59" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="8BB436DC783FFFA5FEDCFBB99E3DFB59" blockId="42.[264,1326,1003,1870]" pageId="42" pageNumber="43">
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<emphasis id="B97FEACE783FFFA5FEDCFBB99D84FBAA" bold="true" box="[312,432,1083,1109]" pageId="42" pageNumber="43">Diagnosis</emphasis>
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. Color piceous. Head markedly impressed on the occiput, almost foveate. Prothorax without the numerous long setae found in adults of
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<taxonomicName id="4C0B4D5F783FFFA5FC19FBE6991EFB81" authority="Liebke. Elytral" authorityName="Liebke. Elytral" box="[1021,1322,1124,1150]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="hilaris">
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<emphasis id="B97FEACE783FFFA5FC19FBE6985AFB82" box="[1021,1134,1124,1149]" italics="true" pageId="42" pageNumber="43">A. hilaris</emphasis>
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Liebke. Elytral
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</taxonomicName>
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intervals
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<taxonomicName id="4C0B4D5F783FFFA5FE90FB0E9DADFB59" box="[372,409,1164,1190]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
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catenate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557783FFFA5FEDCFB319F37FA19" pageId="42" pageNumber="43" type="description">
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<paragraph id="8BB436DC783FFFA5FEDCFB319F37FA19" blockId="42.[264,1326,1003,1870]" pageId="42" pageNumber="43">
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<emphasis id="B97FEACE783FFFA5FEDCFB319DFDFB32" bold="true" box="[312,457,1203,1229]" pageId="42" pageNumber="43">Description</emphasis>
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. Color and luster: Color piceous. Surface shiny. Form: Head (
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<figureCitation id="13302A59783FFFA5FB5AFB369916FB31" box="[1214,1314,1204,1230]" captionStart="FIGURE 18. A" captionStartId="43.[264,368,1163,1187]" captionTargetBox="[419,1166,412,1134]" captionTargetId="figure@43.[418,1169,409,1136]" captionTargetPageId="43" captionText="FIGURE 18. A. ubicki, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0." httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="42" pageNumber="43">Fig. 18a</figureCitation>
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) behind eyes moderately long, markedly rounded, occiput markedly impressed on the occiput, almost foveate. Prothorax widest at basal third, pronotum somewhat flattened with moderately dense coarse punctures. Elytron with apex (
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<figureCitation id="13302A59783FFFA5FC2AFAAE9816FAB9" box="[974,1058,1324,1350]" captionStart="FIGURE 18. A" captionStartId="43.[264,368,1163,1187]" captionTargetBox="[419,1166,412,1134]" captionTargetId="figure@43.[418,1169,409,1136]" captionTargetPageId="43" captionText="FIGURE 18. A. ubicki, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0." httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="42" pageNumber="43">Fig. 18</figureCitation>
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b) almost truncate, lateral tooth small, sutural angle rounded,
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<taxonomicName id="4C0B4D5F783FFFA5FD38FAD69F35FA91" box="[732,769,1364,1390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
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projected acutely. Sternum VI (
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<figureCitation id="13302A59783FFFA5FB9EFAD698FAFA91" box="[1146,1230,1364,1390]" captionStart="FIGURE 18. A" captionStartId="43.[264,368,1163,1187]" captionTargetBox="[419,1166,412,1134]" captionTargetId="figure@43.[418,1169,409,1136]" captionTargetPageId="43" captionText="FIGURE 18. A. ubicki, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0." httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="42" pageNumber="43">Fig. 18</figureCitation>
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c) moderately emarginate and vnotched in both sexes. Aedeagus (
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<figureCitation id="13302A59783FFFA5FC3CFAFE981AFA69" box="[984,1070,1404,1430]" captionStart="FIGURE 18. A" captionStartId="43.[264,368,1163,1187]" captionTargetBox="[419,1166,412,1134]" captionTargetId="figure@43.[418,1169,409,1136]" captionTargetPageId="43" captionText="FIGURE 18. A. ubicki, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0." httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="42" pageNumber="43">Fig. 18</figureCitation>
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d) with short ostium, apex of phallus a very broad triangular lobe, parameres glabrous. Size: small, 8.0 to
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<quantity id="4CF39B39783FFFA5FEECFA4E9D6EFA19" box="[264,346,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="42" pageNumber="43" unit="mm" value="9.5">9.5mm</quantity>
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in length,
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<quantity id="4CF39B39783FFFA5FE3CFA4E9EA7FA19" box="[472,659,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="2.86" metricValueMax="3.24" metricValueMin="2.48" pageId="42" pageNumber="43" unit="mm" value="2.86" valueMax="3.24" valueMin="2.48">2.48 to 3.24mm</quantity>
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in width.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3116557783FFFA5FEA6FA719EF2F8B1" pageId="42" pageNumber="43" type="materials_examined">
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||
<paragraph id="8BB436DC783FFFA5FEA6FA719EFFF929" blockId="42.[264,1326,1003,1870]" pageId="42" pageNumber="43">
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<emphasis id="B97FEACE783FFFA5FEA6FA719E94F9F2" bold="true" box="[322,672,1523,1549]" pageId="42" pageNumber="43">Other specimens examined.</emphasis>
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<collectingCountry id="F31C764C783FFFA5FD4EFA769F06F9F1" box="[682,818,1524,1550]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
|
||
: 4
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||
<typeStatus id="54B0887E783FFFA5FCBDFA769FF8F9F1" box="[857,972,1524,1550]" pageId="42" pageNumber="43" type="paratype">paratypes</typeStatus>
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||
,
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<specimenCount id="9D0DFD55783FFFA5FC3FFA769873F9F1" box="[987,1095,1524,1550]" pageId="42" pageNumber="43" type="female">1 female</specimenCount>
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||
,
|
||
<specimenCount id="9D0DFD55783FFFA5FBB1FA769882F9F1" box="[1109,1206,1524,1550]" pageId="42" pageNumber="43" type="male">3 males</specimenCount>
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||
, from the Osa Conservation Area.
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<collectingCountry id="F31C764C783FFFA5FDD5F99E9E83F9C9" box="[561,695,1564,1590]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
|
||
, Puntarenas,
|
||
<quantity id="4CF39B39783FFFA5FCB3F99E9F4BF9C9" box="[855,895,1564,1590]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1.0">1m</quantity>
|
||
, P.N. Corcovado, Madrigal, 3.0 km W,
|
||
<geoCoordinate id="EE3F501B783FFFA5FED5F9C69DADF9A1" box="[305,409,1604,1630]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-83.6">83°36'W</geoCoordinate>
|
||
<geoCoordinate id="EE3F501B783FFFA5FE46F9C69DC3F9A1" box="[418,503,1604,1630]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="8.483334">8°29'N</geoCoordinate>
|
||
, May, (D. Ubick)(
|
||
<collectionCode id="ED1AAE19783FFFA5FD32F9C69F25F9A1" LSID="urn:lsid:biocol.org:col:34700" box="[726,785,1604,1630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="42" pageNumber="43">CAS</collectionCode>
|
||
: ADP4554),
|
||
<quantity id="4CF39B39783FFFA5FC49F9C69FE1F9A1" box="[941,981,1604,1630]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1.0">1m</quantity>
|
||
, Peninsula de Osa,
|
||
<geoCoordinate id="EE3F501B783FFFA5FB25F9C6991CF9A1" box="[1217,1320,1604,1630]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-83.6">83°36'W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3F501B783FFFA5FEECF9EE9D69F979" box="[264,349,1644,1670]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="8.483334">8°29'N</geoCoordinate>
|
||
, August, (D.R. Whitehead)(
|
||
<collectionCode id="ED1AAE19783FFFA5FD54F9EE9F33F979" LSID="urn:lsid:biocol.org:col:34871" box="[688,775,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
|
||
: ADP59571),
|
||
<quantity id="4CF39B39783FFFA5FC57F9EE9FEFF979" box="[947,987,1644,1670]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="1.0">1m</quantity>
|
||
, Golfito, 3.0 km NE,
|
||
<quantity id="4CF39B39783FFFA5FB07F9EE991DF979" box="[1251,1321,1644,1670]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="43" unit="m" value="100.0">100m</quantity>
|
||
,
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||
<geoCoordinate id="EE3F501B783FFFA5FEECF9169D5BF951" box="[264,367,1684,1710]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-83.166664">83°10'W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3F501B783FFFA5FE99F9169DE5F951" box="[381,465,1684,1710]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="8.65">8°39'N</geoCoordinate>
|
||
, May, (D. Ubick)(
|
||
<collectionCode id="ED1AAE19783FFFA5FD54F9169EDFF951" LSID="urn:lsid:biocol.org:col:34700" box="[688,747,1684,1710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="42" pageNumber="43">CAS</collectionCode>
|
||
: ADP4545), shaking foliage, tropical wet forest, 1f, May, (D. Ubick)(
|
||
<collectionCode id="ED1AAE19783FFFA5FE1AF93E9E0DF929" LSID="urn:lsid:biocol.org:col:34700" box="[510,569,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="42" pageNumber="43">CAS</collectionCode>
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||
: ADP4552).
|
||
</paragraph>
|
||
<paragraph id="8BB436DC783FFFA5FEDCF9619EF2F8B1" blockId="42.[264,1326,1003,1870]" pageId="42" pageNumber="43">
|
||
<emphasis id="B97FEACE783FFFA5FEDCF9619E37F902" bold="true" box="[312,515,1763,1789]" pageId="42" pageNumber="43">Specific epithet.</emphasis>
|
||
The specific epithet,
|
||
<emphasis id="B97FEACE783FFFA5FCF5F9669F54F902" box="[785,864,1764,1789]" italics="true" pageId="42" pageNumber="43">
|
||
<taxonomicName id="4C0B4D5F783FFFA5FCF5F9669F68F902" box="[785,860,1764,1789]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="ubicki">ubicki</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is the Latinized genitive form of the family name of the collector of the
|
||
<typeStatus id="54B0887E783FFFA5FD40F88E9F20F8D9" box="[676,788,1804,1830]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
|
||
, Darryl Ubick of the Department of Entomology, California Academy of Sciences.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116557783EFFA4FEDCFE9998A1F89F" pageId="43" pageNumber="44" type="description">
|
||
<paragraph id="8BB436DC783EFFA4FEDCFE999E8AFEA1" blockId="43.[264,1321,283,350]" pageId="43" pageNumber="44">
|
||
<emphasis id="B97FEACE783EFFA4FEDCFE999DB1FECA" bold="true" box="[312,389,283,309]" pageId="43" pageNumber="44">Notes.</emphasis>
|
||
Adults of this species group are commonly found in suspended dry leaves in the understory of Neotropical forests.
|
||
</paragraph>
|
||
<caption id="DF746654783EFFA4FEECFB0998E2FB18" httpUri="https://zenodo.org/record/162830/files/figure.png" pageId="43" pageNumber="44" targetBox="[419,1166,412,1134]" targetPageId="43">
|
||
<paragraph id="8BB436DC783EFFA4FEECFB0998E2FB18" blockId="43.[264,1323,1163,1255]" pageId="43" pageNumber="44">
|
||
<emphasis id="B97FEACE783EFFA4FEECFB099DACFB5C" bold="true" box="[264,408,1163,1187]" pageId="43" pageNumber="44">FIGURE 18.</emphasis>
|
||
<taxonomicName id="4C0B4D5F783EFFA4FE7BFB099DCAFB5D" box="[415,510,1163,1186]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ubicki">
|
||
<emphasis id="B97FEACE783EFFA4FE7BFB099DCAFB5D" box="[415,510,1163,1186]" italics="true" pageId="43" pageNumber="44">A. ubicki</emphasis>
|
||
</taxonomicName>
|
||
, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.5; WE = 1.5; male SW = 2.2, female SW = 3.0; AL = 2.0.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB436DC783EFFA4FEECFABC9E7AFAA7" blockId="43.[264,590,1341,1368]" box="[264,590,1341,1368]" pageId="43" pageNumber="44">
|
||
The
|
||
<emphasis id="B97FEACE783EFFA4FED9FABF9D97FAA8" bold="true" box="[317,419,1341,1367]" pageId="43" pageNumber="44">palmata</emphasis>
|
||
speciesgroup
|
||
</paragraph>
|
||
<paragraph id="8BB436DC783EFFA4FEECFA0C98F7F98F" blockId="43.[264,1323,1422,1888]" pageId="43" pageNumber="44">
|
||
Diagnostic combination of the
|
||
<emphasis id="B97FEACE783EFFA4FD96FA0C9EE6FA58" box="[626,722,1422,1447]" italics="true" pageId="43" pageNumber="44">palmata</emphasis>
|
||
speciesgroup includes the following: Labrum flat; mentum with lateral lobes acute,
|
||
<taxonomicName id="4C0B4D5F783EFFA4FD6AFA349E87FA2F" box="[654,691,1462,1488]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
|
||
spiniform. Head elongate or
|
||
<taxonomicName id="4C0B4D5F783EFFA4FBEFFA349807FA2F" box="[1035,1075,1462,1488]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
|
||
, broader across middle than occiput, eyes large; head posterior to eyes normally elongate,
|
||
<taxonomicName id="4C0B4D5F783EFFA4FBAAFA5C9847FA07" box="[1102,1139,1502,1528]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
|
||
ballooned (lateral aspect), eyes less than 1.5 times their own diameter from cervical constriction. Tibia moderately or slightly compressed, laterally barely expanded, apex never prolonged; all tibiae lateroapically oblique. Elytral apex bidentate, dentiform projections small.
|
||
</paragraph>
|
||
<paragraph id="8BB436DC783EFFA4FEDCF9FC9EA8F8EF" blockId="43.[264,1323,1422,1888]" pageId="43" pageNumber="44">
|
||
The
|
||
<emphasis id="B97FEACE783EFFA4FE89F9FC9DF9F968" box="[365,461,1662,1687]" italics="true" pageId="43" pageNumber="44">palmata</emphasis>
|
||
speciesgroup was revised by Erwin (1984). At that time, it contained 39 species whose composite range extends from southern
|
||
<collectingCountry id="F31C764C783EFFA4FC6FF9249FD1F93F" box="[907,997,1702,1728]" name="Mexico" pageId="43" pageNumber="44">México</collectingCountry>
|
||
to
|
||
<collectingCountry id="F31C764C783EFFA4FBEEF924984DF93F" box="[1034,1145,1702,1728]" name="Paraguay" pageId="43" pageNumber="44">Paraguay</collectingCountry>
|
||
. Since then, 13 new species have been discovered from throughout the overall range, plus another one from
|
||
<collectingCountry id="F31C764C783EFFA4FEACF9749DFFF8EF" box="[328,459,1782,1808]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
|
||
, described below.
|
||
</paragraph>
|
||
<paragraph id="8BB436DC783EFFA4FEDCF89C98A1F89F" blockId="43.[264,1323,1422,1888]" pageId="43" pageNumber="44">
|
||
Four other species of the
|
||
<emphasis id="B97FEACE783EFFA4FD86F89C9EF6F8C8" box="[610,706,1822,1847]" italics="true" pageId="43" pageNumber="44">palmata</emphasis>
|
||
speciesgroup are known from
|
||
<collectingCountry id="F31C764C783EFFA4FBD3F89C988FF8C7" box="[1079,1211,1822,1848]" name="Costa Rica" pageId="43" pageNumber="44">Costa Rica</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C0B4D5F783EFFA4FB23F89C9DB9F89F" authority="Erwin" authorityName="Erwin" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atriperna">
|
||
<emphasis id="B97FEACE783EFFA4FB23F89C9D0AF8A0" italics="true" pageId="43" pageNumber="44">A. atriperna</emphasis>
|
||
Erwin
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B4D5F783EFFA4FE78F8C49E99F89F" authority="Liebke" authorityName="Liebke" box="[412,685,1862,1888]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="B97FEACE783EFFA4FE78F8C49D9AF8A0" box="[412,430,1862,1887]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="B97FEACE783EFFA4FE58F8C49E61F8A0" box="[444,597,1862,1887]" italics="true" pageId="43" pageNumber="44">costaricensis</emphasis>
|
||
Liebke
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0B4D5F783EFFA4FD59F8C49FACF89F" authority="Straneo" authorityName="Straneo" box="[701,920,1862,1888]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="flavipes">
|
||
<emphasis id="B97FEACE783EFFA4FD59F8C49F0CF8A0" box="[701,824,1862,1887]" italics="true" pageId="43" pageNumber="44">A. flavipes</emphasis>
|
||
Straneo
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0B4D5F783EFFA4FC3DF8C498A1F89F" authority="Erwin." authorityName="Erwin." box="[985,1173,1862,1888]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="kayae">
|
||
<emphasis id="B97FEACE783EFFA4FC3DF8C4980BF8A0" box="[985,1087,1862,1887]" italics="true" pageId="43" pageNumber="44">A. kayae</emphasis>
|
||
Erwin.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |