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<document id="4BA89A6B687C231AD82A94766CF7C56F" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA783AFFBFFEECFD63987DFC49" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra solanoi Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="49" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="48" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="C9E56F5D8878D2EA48A9C1745DD1DCF0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="32C83F5E5E3BCAFCE81717B1A00DD20C">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="46F097A35FD22FA21B4FF7333EF5A707" type="personal">
<mods:role id="F429FFC9C40BBD1FFF3F4E752A5FFD56">
<mods:roleTerm id="AD1E603C78A5BBCD70C34AFB634F4A92">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB337A14ED4A5CA5D3F603D9F22D4B5C">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="0D1A0417F2710BBC49816D5C3C1C27B6">text</mods:typeOfResource>
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<mods:title id="30A6B826ADFF30CFFCFCAB885046EAA4">Zootaxa</mods:title>
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<mods:part id="EF315D3FE2DCB72EAD53CF26363681FF">
<mods:date id="D89E997268C12C60824F50B4EC81B957">2002</mods:date>
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<mods:number id="695488FCF1881A597CE79FC4D362F154">119</mods:number>
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<mods:classification id="00319D61CC3F064FC43B76B170607DBF">journal article</mods:classification>
<mods:identifier id="B44CD72E28E3C75895BA489F0F480146" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="23B713814A633571C7342B52D453FCB8" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="994FFDDAE295778DCA3CD506697FDEDC" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA783AFFBFFEECFD63987DFC49" ID-DOI="http://doi.org/10.5281/zenodo.6278013" ID-GBIF-Taxon="121065198" ID-Zenodo-Dep="6278013" LSID="urn:lsid:plazi:treatment:03A287CA783AFFBFFEECFD63987DFC49" httpUri="http://treatment.plazi.org/id/03A287CA783AFFBFFEECFD63987DFC49" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<subSubSection id="C3116557783AFFA0FEECFD639E02FCDC" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BB436DC783AFFA0FEECFD639EACFD04" blockId="47.[264,664,737,803]" box="[264,664,737,763]" pageId="47" pageNumber="48">
<heading id="D0FC81B0783AFFA0FEECFD639EACFD04" bold="true" box="[264,664,737,763]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="B97FEACE783AFFA0FEECFD639EACFD04" bold="true" box="[264,664,737,763]" pageId="47" pageNumber="48">
<taxonomicName id="4C0B4D5F783AFFA0FEECFD639DCFFD04" ID-CoL="B2Y2" authority="Erwin" authorityName="Erwin" box="[264,507,737,763]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="solanoi" status="sp. nov.">
<emphasis id="B97FEACE783AFFA0FEECFD639D90FD04" bold="true" box="[264,420,737,763]" italics="true" pageId="47" pageNumber="48">Agra solanoi</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B5783AFFA0FDECFD639EACFD04" box="[520,664,737,763]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC783AFFA0FEECFC8B9E02FCDC" blockId="47.[264,664,737,803]" box="[264,566,777,803]" pageId="47" pageNumber="48">
(
<figureCitation id="13302A59783AFFA0FEF5FC8B9D9EFCDC" box="[273,426,777,803]" captionStart="FIGURE 21. A" captionStartId="48.[264,367,1751,1775]" captionTargetBox="[381,1202,1013,1731]" captionTargetId="figure@48.[381,1206,1013,1739]" captionTargetPageId="48" captionText="FIGURE 21. A. solanoi, dorsal aspect: a) head, male, female; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.0; WE = 2.8; male SW = 4.4, female SW = 4.6; AL = 4.5." httpUri="https://zenodo.org/record/162835/files/figure.png" pageId="47" pageNumber="48">Figs. 21a, 21</figureCitation>
b, 21c, 21d)
</paragraph>
</subSubSection>
<subSubSection id="C3116557783AFFA0FEECFCDB981DFC64" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BB436DC783AFFA0FEECFCDB981DFC64" blockId="47.[264,1325,857,1883]" pageId="47" pageNumber="48">
<emphasis id="B97FEACE783AFFA0FEECFCDB9DB5FC8C" bold="true" box="[264,385,857,883]" pageId="47" pageNumber="48">
<typeStatus id="54B0887E783AFFA0FEECFCDB9D48FC8C" box="[264,380,857,883]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<emphasis id="B97FEACE783AFFA0FE38FCDB9EAAFC8C" bold="true" box="[476,670,857,883]" pageId="47" pageNumber="48">
<collectingCountry id="F31C764C783AFFA0FE38FCDB9EADFC8C" box="[476,665,857,883]" name="Costa Rica" pageId="47" pageNumber="48">COSTA RICA</collectingCountry>
:
</emphasis>
Limón, Tortuguero National Park, Cerro Tortuguero,
<quantity id="4CF39B39783AFFA0FEECFC039D79FC64" box="[264,333,897,923]" metricMagnitude="2" metricUnit="m" metricValue="1.19" pageId="47" pageNumber="48" unit="m" value="119.0">119m</quantity>
,
<geoCoordinate id="EE3F501B783AFFA0FEBDFC039D8EFC64" box="[345,442,897,923]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="10.566667">10°34'N</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FE25FC039E1CFC64" box="[449,552,897,923]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-83.53333">83°32'W</geoCoordinate>
, May, (J. Solano)(
<taxonomicName id="4C0B4D5F783AFFA0FCE4FC039F7AFC64" box="[768,846,897,923]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19783AFFA0FCE4FC039F7AFC64" box="[768,846,897,923]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="47" pageNumber="48">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19783AFFA0FCBDFC039FB8FC64" LSID="urn:lsid:biocol.org:col:14323" box="[857,908,897,923]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="47" pageNumber="48">CRI</collectionCode>
000­242786).
</paragraph>
</subSubSection>
<subSubSection id="C3116557783AFFA0FEDCFC2B9804FBC4" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BB436DC783AFFA0FEDCFC2B9804FBC4" blockId="47.[264,1325,857,1883]" pageId="47" pageNumber="48">
<emphasis id="B97FEACE783AFFA0FEDCFC2B9D84FC3C" bold="true" box="[312,432,937,963]" pageId="47" pageNumber="48">Diagnosis</emphasis>
. Males with dense, short vestiture on metasternum and long dense vestiture on abdominal sterna III­V. Elytral apex spinose medially, markedly projected acutely laterally, with slight lobe between; interneurs striatio­punctate with regular unisetose foveae in intervals 2, 4, 6, and 8. Sternum VI of male with lateral angles acute.
</paragraph>
</subSubSection>
<subSubSection id="C3116557783AFFA0FEDCFBCB993AFA5C" pageId="47" pageNumber="48" type="description">
<paragraph id="8BB436DC783AFFA0FEDCFBCB993AFA5C" blockId="47.[264,1325,857,1883]" pageId="47" pageNumber="48">
<emphasis id="B97FEACE783AFFA0FEDCFBCB9DFDFB9C" bold="true" box="[312,457,1097,1123]" pageId="47" pageNumber="48">Description</emphasis>
. Color and luster: Brunneous. Antennae rufotestaceous; slightly infuscated femora contrast with testaceous tibiae and tarsi. Form: Head (
<figureCitation id="13302A59783AFFA0FBCEFBF398BAFB74" box="[1066,1166,1137,1163]" captionStart="FIGURE 21. A" captionStartId="48.[264,367,1751,1775]" captionTargetBox="[381,1202,1013,1731]" captionTargetId="figure@48.[381,1206,1013,1739]" captionTargetPageId="48" captionText="FIGURE 21. A. solanoi, dorsal aspect: a) head, male, female; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.0; WE = 2.8; male SW = 4.4, female SW = 4.6; AL = 4.5." httpUri="https://zenodo.org/record/162835/files/figure.png" pageId="47" pageNumber="48">Fig. 21a</figureCitation>
) behind eyes short, markedly angulate posteriorly in male, slightly rounded in female. Prothorax robust, wider than head. Elytron (
<figureCitation id="13302A59783AFFA0FDA4FB439EA1FB24" box="[576,661,1217,1243]" captionStart="FIGURE 21. A" captionStartId="48.[264,367,1751,1775]" captionTargetBox="[381,1202,1013,1731]" captionTargetId="figure@48.[381,1206,1013,1739]" captionTargetPageId="48" captionText="FIGURE 21. A. solanoi, dorsal aspect: a) head, male, female; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.0; WE = 2.8; male SW = 4.4, female SW = 4.6; AL = 4.5." httpUri="https://zenodo.org/record/162835/files/figure.png" pageId="47" pageNumber="48">Fig. 21</figureCitation>
b) with apex slightly lobed medially, lateral tooth large, acute, sutural apex spinose. Males with dense, short vestiture on metasternum, long dense vestiture on abdominal sterna III­V; middle tibia slightly clavate apico­medially. Sternum VI (
<figureCitation id="13302A59783AFFA0FEDFFABB9DA0FAAC" box="[315,404,1337,1363]" captionStart="FIGURE 21. A" captionStartId="48.[264,367,1751,1775]" captionTargetBox="[381,1202,1013,1731]" captionTargetId="figure@48.[381,1206,1013,1739]" captionTargetPageId="48" captionText="FIGURE 21. A. solanoi, dorsal aspect: a) head, male, female; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.0; WE = 2.8; male SW = 4.4, female SW = 4.6; AL = 4.5." httpUri="https://zenodo.org/record/162835/files/figure.png" pageId="47" pageNumber="48">Fig. 21</figureCitation>
c) deeply V­notched in male, narrowly and shallowly V­notched in female. Aedeagus (
<figureCitation id="13302A59783AFFA0FE6BFAE39DD3FA84" box="[399,487,1377,1403]" captionStart="FIGURE 21. A" captionStartId="48.[264,367,1751,1775]" captionTargetBox="[381,1202,1013,1731]" captionTargetId="figure@48.[381,1206,1013,1739]" captionTargetPageId="48" captionText="FIGURE 21. A. solanoi, dorsal aspect: a) head, male, female; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.0; WE = 2.8; male SW = 4.4, female SW = 4.6; AL = 4.5." httpUri="https://zenodo.org/record/162835/files/figure.png" pageId="47" pageNumber="48">Fig. 21</figureCitation>
c) with ostium moderately long, left pleuropic, apex narrowly lobate, parameres setiferous. Size: medium, 14.6 to 17.0 mm in length,
<quantity id="4CF39B39783AFFA0FC1DFA0B98A9FA5C" box="[1017,1181,1417,1443]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="47" pageNumber="48" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4 to 2.8 mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C3116557783AFFBFFEDCFA339DD5FC99" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BB436DC783AFFBFFEDCFA339DAAFD11" blockId="47.[264,1325,857,1883]" lastBlockId="48.[264,1323,284,950]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B97FEACE783AFFA0FEDCFA339F2BFA34" bold="true" box="[312,799,1457,1483]" pageId="47" pageNumber="48">
Other specimens examined.
<collectingCountry id="F31C764C783AFFA0FD73FA339F2BFA34" box="[663,799,1457,1483]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
</emphasis>
: 17
<typeStatus id="54B0887E783AFFA0FCB6FA339FF2FA34" box="[850,966,1457,1483]" pageId="47" pageNumber="48" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D0DFD55783AFFA0FC35FA339808FA34" box="[977,1084,1457,1483]" pageId="47" pageNumber="48" type="male">10 males</specimenCount>
,
<specimenCount id="9D0DFD55783AFFA0FBACFA339888FA34" box="[1096,1212,1457,1483]" pageId="47" pageNumber="48" type="female">7 females</specimenCount>
, from the following Conservation Areas: Arenal Tilarán, Cordillera Volcanica Central, Guanacaste, Osa, Pacifico Central, and Tortuguero.
<collectingCountry id="F31C764C783AFFA0FD1CF9839FB6F9E4" box="[760,898,1537,1563]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
, Cartago,
<quantity id="4CF39B39783AFFA0FBECF9839804F9E4" box="[1032,1072,1537,1563]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
,
<taxonomicName id="4C0B4D5F783AFFA0FBA7F9839887F9E4" box="[1091,1203,1537,1563]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
,
<geoCoordinate id="EE3F501B783AFFA0FB22F9839919F9E4" box="[1222,1325,1537,1563]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FEECF9AB9D68F9BC" box="[264,348,1577,1603]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="9.883333">9°53'N</geoCoordinate>
, May, (H.F. Howden &amp; A. Howden)(
<collectionCode id="ED1AAE19783AFFA0FCD0F9AB9FB9F9BC" LSID="urn:lsid:biocol.org:col:34404" box="[820,909,1577,1603]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="47" pageNumber="48">UASM</collectionCode>
: ADP85301),
<quantity id="4CF39B39783AFFA0FBDBF9AB9853F9BC" box="[1087,1127,1577,1603]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
, January, (V.M. Kirk)(
<collectionCode id="ED1AAE19783AFFA0FEB6F9D39D9FF994" LSID="urn:lsid:biocol.org:col:34404" box="[338,427,1617,1643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="47" pageNumber="48">UASM</collectionCode>
: ADP21812),
<quantity id="4CF39B39783AFFA0FDB6F9D39E4DF994" box="[594,633,1617,1643]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
, January, (V.M. Kirk)(
<collectionCode id="ED1AAE19783AFFA0FC67F9D39FE8F994" LSID="urn:lsid:biocol.org:col:34404" box="[899,988,1617,1643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="47" pageNumber="48">UASM</collectionCode>
: ADP21823),
<quantity id="4CF39B39783AFFA0FB67F9D3989FF994" box="[1155,1195,1617,1643]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
,
<taxonomicName id="4C0B4D5F783AFFA0FB5CF9D39D68F96C" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">
Turrialba,
<quantity id="4CF39B39783AFFA0FEECF9FB9D68F96C" box="[264,348,1657,1683]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="47" pageNumber="48" unit="m" value="1200.0">1200m</quantity>
</taxonomicName>
,
<geoCoordinate id="EE3F501B783AFFA0FE8CF9FB9DFBF96C" box="[360,463,1657,1683]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FE32F9FB9E1FF96C" box="[470,555,1657,1683]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="9.883333">9°53'N</geoCoordinate>
, March, (W.R. Enns)(UMoC: ADP
<date id="FFB5101C783AFFA0FC2FF9FB9810F96C" box="[971,1060,1657,1683]" pageId="47" pageNumber="48" value="29-10-05">100529</date>
), 1f,
<taxonomicName id="4C0B4D5F783AFFA0FBBBF9FB98FAF96C" box="[1119,1230,1657,1683]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
, 3.0 km
<collectionCode id="ED1AAE19783AFFA0FEECF9239D19F944" box="[264,301,1697,1723]" pageId="47" pageNumber="48">SE</collectionCode>
,
<taxonomicName id="4C0B4D5F783AFFA0FEDDF9239DBBF944" box="[313,399,1697,1723]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="51" phylum="Arthropoda" rank="species" species="catie">
<collectionCode id="ED1AAE19783AFFA0FEDDF9239DBBF944" LSID="urn:lsid:biocol.org:col:15110" box="[313,399,1697,1723]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15110" name="Tropical Agricultural Research and Training Center (CATIE)" pageId="47" pageNumber="48">CATIE</collectionCode>
</taxonomicName>
,
<geoCoordinate id="EE3F501B783AFFA0FE7FF9239E36F944" box="[411,514,1697,1723]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FDEEF9239E6AF944" box="[522,606,1697,1723]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="9.883333">9°53'N</geoCoordinate>
, June, (J.A. Chemsak, H. Katasura &amp; A. &amp; E. Michelbacher) (
<collectionCode id="ED1AAE19783AFFA0FEF5F94B9D67F91C" box="[273,339,1737,1763]" httpUri="http://grbio.org/cool/wq7v-amgw" name="University of California at Berkeley" pageId="47" pageNumber="48">UCB</collectionCode>
: ADP70518); Guanacaste,
<quantity id="4CF39B39783AFFA0FD4BF94B9EE3F91C" box="[687,727,1737,1763]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
, Tilarán,
<quantity id="4CF39B39783AFFA0FCBEF94B9FABF91C" box="[858,927,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="48" unit="m" value="800.0">800m</quantity>
,
<geoCoordinate id="EE3F501B783AFFA0FC52F94B9829F91C" box="[950,1053,1737,1763]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-84.98333">84°59'W</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FBCBF94B98A6F91C" box="[1071,1170,1737,1763]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="10.466666">10°28'N</geoCoordinate>
, (F. Nevermann)(WAR: ADP4224); Heredia, 1f, Estacíon La Selva, Puerto Viejo, 3.0 km S,
<quantity id="4CF39B39783AFFA0FAE6F9739D7AF8CC" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="47" pageNumber="48" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50­ 150m</quantity>
,
<geoCoordinate id="EE3F501B783AFFA0FEBFF89B9DF7F8CC" box="[347,451,1817,1843]" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="-83.98333">83°59'W</geoCoordinate>
<geoCoordinate id="EE3F501B783AFFA0FE28F89B9E1BF8CC" box="[460,559,1817,1843]" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="10.45">10°27'N</geoCoordinate>
, January, (D.L. Wagner)(
<taxonomicName id="4C0B4D5F783AFFA0FC84F89B9F87F8CC" box="[864,947,1817,1843]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
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</taxonomicName>
:
<collectionCode id="ED1AAE19783AFFA0FC24F89B9FC6F8CC" LSID="urn:lsid:biocol.org:col:14323" box="[960,1010,1817,1843]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="47" pageNumber="48">CRI</collectionCode>
002­724432), at light,
<quantity id="4CF39B39783AFFA0FB1AF89B9911F8CC" box="[1278,1317,1817,1843]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
, June, (F.T. Hovore)(
<collectionCode id="ED1AAE19783AFFA0FE10F8C39E76F8A4" box="[500,578,1857,1883]" pageId="47" pageNumber="48">FTHC</collectionCode>
: ADP87130), 1f, June, (F.T. Hovore)(
<collectionCode id="ED1AAE19783AFFA0FC1FF8C3987DF8A4" box="[1019,1097,1857,1883]" pageId="47" pageNumber="48">FTHC</collectionCode>
: ADP
<date id="FFB5101C783AFFA0FB76F8C398DEF8A4" box="[1170,1258,1857,1883]" pageId="47" pageNumber="48" value="31-10-05">100531</date>
),
<quantity id="4CF39B39783AFFA0FB1AF8C39912F8A4" box="[1278,1318,1857,1883]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="48" unit="m" value="1.0">1m</quantity>
, Puerto Viejo, 3.0 km S, Finca La Selva,
<geoCoordinate id="EE3F501B7825FFBFFCC9FE9E9FA0FEC9" box="[813,916,284,310]" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-84.01667">84°01'W</geoCoordinate>
<geoCoordinate id="EE3F501B7825FFBFFC41FE9E983DFEC9" box="[933,1033,284,310]" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="10.433333">10°26'N</geoCoordinate>
, April, (H.
<taxonomicName id="4C0B4D5F7825FFBFFB7FFE9E9D78FEA1" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="32" phylum="Arthropoda" rank="species" species="hespenheide">A. Hespenheide</taxonomicName>
)(
<collectionCode id="ED1AAE197825FFBFFEB8FEC69DABFEA1" box="[348,415,324,350]" httpUri="http://grbio.org/cool/g8qa-33h5" name="Hoyt Arboretum" pageId="48" pageNumber="49">HAH</collectionCode>
: ADP5094); Limón, 1f, P.N. Tortuguero, Estacíon Cuatro Esquinas, 0m,
<geoCoordinate id="EE3F501B7825FFBFFEECFEEE9D5BFE79" box="[264,367,364,390]" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-83.53333">83°32'W</geoCoordinate>
<geoCoordinate id="EE3F501B7825FFBFFE99FEEE9DD4FE79" box="[381,480,364,390]" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="10.566667">10°34'N</geoCoordinate>
, LN 280000,590500, October, (J. Solano)(
<taxonomicName id="4C0B4D5F7825FFBFFC10FEEE987EFE79" box="[1012,1098,364,390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFC10FEEE987EFE79" box="[1012,1098,364,390]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFBBFFEEE98B9FE79" LSID="urn:lsid:biocol.org:col:14323" box="[1115,1165,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
000­296760),
<quantity id="4CF39B397825FFBFFEECFE169D04FE51" box="[264,304,404,430]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1.0">1m</quantity>
, Tortuguero, Cerro Tortuguero,
<quantity id="4CF39B397825FFBFFD48FE169F3EFE51" box="[684,778,404,430]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="48" pageNumber="49" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0­100m</quantity>
,
<geoCoordinate id="EE3F501B7825FFBFFCF3FE169F4BFE51" box="[791,895,404,430]" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-83.53333">83°32'W</geoCoordinate>
<geoCoordinate id="EE3F501B7825FFBFFC63FE169FDFFE51" box="[903,1003,404,430]" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="10.566667">10°34'N</geoCoordinate>
, LN 285000,588000, May, (J. Solano)(
<taxonomicName id="4C0B4D5F7825FFBFFE76FE3E9DDCFE29" box="[402,488,444,470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFE76FE3E9DDCFE29" box="[402,488,444,470]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFE10FE3E9E13FE29" LSID="urn:lsid:biocol.org:col:14323" box="[500,551,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
000­242793),
<quantity id="4CF39B397825FFBFFD28FE3E9EC0FE29" box="[716,756,444,470]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1.0">1m</quantity>
, October, (J. Solano)(
<taxonomicName id="4C0B4D5F7825FFBFFC1CFE3E987AFE29" box="[1016,1102,444,470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFC1CFE3E987AFE29" box="[1016,1102,444,470]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFBBEFE3E98B9FE29" LSID="urn:lsid:biocol.org:col:14323" box="[1114,1165,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
000­
<date id="FFB5101C7825FFBFFB25FE3E9928FE29" box="[1217,1308,444,470]" pageId="48" pageNumber="49" value="05-03-22">032205</date>
), 1f, June, (R. Delgado)(
<taxonomicName id="4C0B4D5F7825FFBFFDFAFE669E47FE01" box="[542,627,484,510]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFDFAFE669E47FE01" box="[542,627,484,510]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFD64FE669E87FE01" LSID="urn:lsid:biocol.org:col:14323" box="[640,691,484,510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
001­363770), 1f, Puntarenas, A.C. Osa, Peninsula de Osa, Bosque Esquinas,
<quantity id="4CF39B397825FFBFFDC8FD8E9E46FDD9" box="[556,626,524,550]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="m" value="200.0">200m</quantity>
, LS 301400,542200 #2816, May, (J.F.
<taxonomicName id="4C0B4D5F7825FFBFFBBDFD8E98F7FDD9" box="[1113,1219,524,550]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="32" phylum="Arthropoda" rank="species" species="quesada">Quesada</taxonomicName>
)(
<taxonomicName id="4C0B4D5F7825FFBFFB37FD8E991CFDD9" box="[1235,1320,524,550]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFB37FD8E991CFDD9" box="[1235,1320,524,550]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFEECFDB69D0FFDB1" LSID="urn:lsid:biocol.org:col:14323" box="[264,315,564,590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
001­862232),
<quantity id="4CF39B397825FFBFFE08FDB69E27FDB1" box="[492,531,564,590]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1.0">1m</quantity>
, P.N. Manuel Antonio, Quepos,
<quantity id="4CF39B397825FFBFFC34FDB69833FDB1" box="[976,1031,564,590]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="48" pageNumber="49" unit="m" value="80.0">80m</quantity>
,
<geoCoordinate id="EE3F501B7825FFBFFBFBFDB698B2FDB1" box="[1055,1158,564,590]" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-84.15">84°09'W</geoCoordinate>
<geoCoordinate id="EE3F501B7825FFBFFB7FFDB698DBFDB1" box="[1179,1263,564,590]" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="9.4">9°24'N</geoCoordinate>
, LS 370900,448800 #1181, April, (C. Cano)(
<taxonomicName id="4C0B4D5F7825FFBFFD08FDDE9F76FD89" box="[748,834,604,630]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFD08FDDE9F76FD89" box="[748,834,604,630]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFCABFDDE9FB6FD89" LSID="urn:lsid:biocol.org:col:14323" box="[847,898,604,630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
001­718849), 1f, Peninsula de Osa., Rancho Quemado,
<quantity id="4CF39B397825FFBFFDE1FD069E7FFD61" box="[517,587,644,670]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="m" value="200.0">200m</quantity>
, LS 292500,511000, September, (F.
<taxonomicName id="4C0B4D5F7825FFBFFBC3FD0698F7FD61" box="[1063,1219,644,670]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1" phylum="Arthropoda" rank="species" species="quesada" status="sp. nov.">A. Quesada</taxonomicName>
)(
<taxonomicName id="4C0B4D5F7825FFBFFB37FD06991CFD61" box="[1235,1320,644,670]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197825FFBFFB37FD06991CFD61" box="[1235,1320,644,670]" pageId="48" pageNumber="49">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197825FFBFFEECFD2E9D08FD39" LSID="urn:lsid:biocol.org:col:14323" box="[264,316,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="48" pageNumber="49">CRI</collectionCode>
001­
<date id="FFB5101C7825FFBFFE95FD2E9DFFFD39" box="[369,459,684,710]" pageId="48" pageNumber="49" value="51-04-19">190451</date>
),
<quantity id="4CF39B397825FFBFFE09FD2E9E21FD39" box="[493,533,684,710]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="m" value="1.0">1m</quantity>
,
<taxonomicName id="4C0B4D5F7825FFBFFDCBFD2E9EF4FD39" box="[559,704,684,710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="62" phylum="Arthropoda" rank="species" species="monteverde">Monteverde</taxonomicName>
,
<geoCoordinate id="EE3F501B7825FFBFFD3DFD2E9F74FD39" box="[729,832,684,710]" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-84.8">84°48'W</geoCoordinate>
<geoCoordinate id="EE3F501B7825FFBFFCB2FD2E9F8DFD39" box="[854,953,684,710]" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="10.3">10°18'N</geoCoordinate>
, May, (F.T. Hovore)(
<collectionCode id="ED1AAE197825FFBFFB3DFD2E9913FD39" box="[1241,1319,684,710]" pageId="48" pageNumber="49">FTHC</collectionCode>
: ADP70527).
</paragraph>
<paragraph id="8BB436DC7825FFBFFEDCFD799DD5FC99" blockId="48.[264,1323,284,950]" pageId="48" pageNumber="49">
<emphasis id="B97FEACE7825FFBFFEDCFD799E35FCEA" bold="true" box="[312,513,763,789]" pageId="48" pageNumber="49">Specific epithet.</emphasis>
The specific epithet,
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<emphasis id="B97FEACE7825FFBFFCEFFD7E9F55FCEA" box="[779,865,764,789]" italics="true" pageId="48" pageNumber="49">solanoi</emphasis>
</taxonomicName>
, is the Latinized genitive form of the family name of collector of the
<typeStatus id="54B0887E7825FFBFFD6FFCA69ECFFCC1" box="[651,763,804,830]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
, parataxonomist José Solano of San Isidro del General, San José.
</paragraph>
</subSubSection>
<subSubSection id="C31165577825FFBFFEDCFCF1987DFC49" pageId="48" pageNumber="49" type="description">
<paragraph id="8BB436DC7825FFBFFEDCFCF1987DFC49" blockId="48.[264,1323,284,950]" pageId="48" pageNumber="49">
<emphasis id="B97FEACE7825FFBFFEDCFCF19D4AFC72" bold="true" box="[312,382,883,909]" pageId="48" pageNumber="49">Notes</emphasis>
. This widespread species ranges from sea level to
<quantity id="4CF39B397825FFBFFC3BFCF69805FC71" box="[991,1073,884,910]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="48" pageNumber="49" unit="m" value="1200.0">1200m</quantity>
altitude, in both wet and dry forests. Adults of this species are commonly attracted to lights.
</paragraph>
</subSubSection>
</treatment>
</document>