155 lines
19 KiB
XML
155 lines
19 KiB
XML
<document id="DF3A0611844E12DF5BFA2AE64CF5A6BB" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7838FFA1FEECFCCE9EDBFE51" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra schwarzeneggeri Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="47" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="46" updateTime="1698190273211" updateUser="plazi">
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<mods:mods id="17E5AAE81768DB083BB6CE604E05BE12" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4C2EB41649E6F1E82726D5B1BB4D9182">
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<mods:title id="4DB48ACF522F7F37374CC454430E8819">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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</mods:titleInfo>
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<mods:name id="57241336B4809A0C234EDB9974C9C048" type="personal">
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<mods:roleTerm id="EBFBED12B76673622CDAC76A6ADF7B98">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2F63F308D0B067E9BFE409E0BB43938E">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="3CEB23A1F5CE7775C5240EAD98B95337">text</mods:typeOfResource>
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<mods:titleInfo id="9594371911B2F4CCD1E6C925E6333370">
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<mods:title id="5FBC86F2AB6A6327A507E69F7974C854">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="586549929F1E8FDE583D7E4A028EAA08">
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<mods:date id="811C7441F6DB13053296BAEF18ED0134">2002</mods:date>
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<mods:detail id="63599971C0EBA266404491D48BD18BD2" type="volume">
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<mods:number id="17DEC13735E364DA58DED1A129BF2A81">119</mods:number>
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<mods:start id="30916A06E2B7FBF8E432A47BD530675C">1</mods:start>
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<mods:end id="8AF59E4BCDC437D8BE5B1C52F453959C">68</mods:end>
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<mods:classification id="9B01D46167E3E24FE735CD0C2EAB3DCA">journal article</mods:classification>
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<mods:identifier id="82725E09B3A1D003AFFD9EB7B1E479CC" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="24F7CE74C180D201EED9F09870FBCC66" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="6479B4DECF2DED8ECDB1D88BD0B2C13C" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="46CF7D24166249D9716FCB923EB9153E" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA7838FFA1FEECFCCE9EDBFE51" ID-DOI="http://doi.org/10.5281/zenodo.6278005" ID-GBIF-Taxon="121065199" ID-Zenodo-Dep="6278005" LSID="urn:lsid:plazi:treatment:03A287CA7838FFA1FEECFCCE9EDBFE51" httpUri="http://treatment.plazi.org/id/03A287CA7838FFA1FEECFCCE9EDBFE51" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="C31165577838FFA2FEECFCCE9E02FC71" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="8BB436DC7838FFA2FEECFCCE9F3EFC9A" blockId="45.[264,778,843,910]" box="[264,778,843,870]" pageId="45" pageNumber="46">
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<heading id="D0FC81B07838FFA2FEECFCCE9F3EFC9A" bold="true" box="[264,778,843,870]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<emphasis id="B97FEACE7838FFA2FEECFCCE9F3EFC9A" bold="true" box="[264,778,843,870]" pageId="45" pageNumber="46">
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<taxonomicName id="4C0B4D5F7838FFA2FEECFCCE9E59FC9A" ID-CoL="B2X8" authority="Erwin" authorityName="Erwin" box="[264,621,843,870]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schwarzeneggeri" status="sp. nov.">
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<emphasis id="B97FEACE7838FFA2FEECFCCE9E22FC99" bold="true" box="[264,534,844,870]" italics="true" pageId="45" pageNumber="46">Agra schwarzeneggeri</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B57838FFA2FD9EFCC99F3EFC9A" box="[634,778,843,869]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC7838FFA2FEECFCF69E02FC71" blockId="45.[264,778,843,910]" box="[264,566,884,910]" pageId="45" pageNumber="46">
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(
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<figureCitation id="13302A597838FFA2FEF5FCF69D9FFC71" box="[273,427,884,910]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, postdorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Figs. 20a, 20</figureCitation>
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b, 20c, 20d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577838FFA2FEECFC439E8EFBFC" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="8BB436DC7838FFA2FEECFC439E8EFBFC" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
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<emphasis id="B97FEACE7838FFA2FEECFC439DB5FC24" bold="true" box="[264,385,961,987]" pageId="45" pageNumber="46">
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<typeStatus id="54B0887E7838FFA2FEECFC439D48FC24" box="[264,380,961,987]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male,
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<collectingCountry id="F31C764C7838FFA2FE37FC439EB6FC24" box="[467,642,961,987]" name="Costa Rica" pageId="45" pageNumber="46">COSTA RICA</collectingCountry>
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: Cartago,
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<taxonomicName id="4C0B4D5F7838FFA2FD13FC439F52FC24" box="[759,870,961,987]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
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,
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<quantity id="4CF39B397838FFA2FC95FC439F83FC24" box="[881,951,961,987]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="45" pageNumber="46" unit="m" value="650.0">650m</quantity>
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,
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<geoCoordinate id="EE3F501B7838FFA2FC26FC43982CFC24" box="[962,1048,961,987]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.883333">9º 53'N</geoCoordinate>
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<geoCoordinate id="EE3F501B7838FFA2FBFBFC4398B7FC24" box="[1055,1155,961,987]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.63333">83º38'W</geoCoordinate>
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, February, (H. & A. Howden)(
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<collectionCode id="ED1AAE197838FFA2FE26FC6B9E2FFBFC" LSID="urn:lsid:biocol.org:col:34871" box="[450,539,1001,1027]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46">USNM</collectionCode>
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: ADP64794).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577838FFA2FEDCFB939D77FB84" pageId="45" pageNumber="46" type="diagnosis">
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<paragraph id="8BB436DC7838FFA2FEDCFB939D77FB84" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
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<emphasis id="B97FEACE7838FFA2FEDCFB939D83FBD4" bold="true" box="[312,439,1041,1067]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
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The male of this species has an enormously enlarged middle femur (
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<figureCitation id="13302A597838FFA2FB1EFB939D13FBAC" captionStart="FIGURE 19. A" captionStartId="44.[264,368,1791,1815]" captionTargetBox="[406,1178,1107,1755]" captionTargetId="figure@44.[405,1181,1089,1765]" captionTargetPageId="44" captionText="FIGURE 19. A. granodeoro, male, dorsal aspect: a) head; b) apex of elytron; c) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.3; WE = 2.2; AL = 5.0." httpUri="https://zenodo.org/record/162832/files/figure.png" pageId="45" pageNumber="46">Fig. 19</figureCitation>
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c); females have elytra with brassygreen tint, black forebody and rufous antennomeres 311.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577838FFA2FEDCFB0B985EFA1C" pageId="45" pageNumber="46" type="description">
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<paragraph id="8BB436DC7838FFA2FEDCFB0B985EFA1C" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
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<emphasis id="B97FEACE7838FFA2FEDCFB0B9DE4FB5C" bold="true" box="[312,464,1161,1187]" pageId="45" pageNumber="46">Description.</emphasis>
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Color and luster: Black. Elytra with brassy green reflections. Antennomeres 34 testaceous, infuscated apically, 511 testaceous. Form: Head (
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<figureCitation id="13302A597838FFA2FB92FB3398E9FB34" box="[1142,1245,1201,1227]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, postdorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20a</figureCitation>
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) short behind eyes, that of male tapered and rounded behind, that of female quadrate, vertex dimpled. Prothorax widest at middle, markedly tapered anteriorly in male, somewhat "swollen" at middle in female. Middle leg with markedly enlarged femur (
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<figureCitation id="13302A597838FFA2FBBBFAAB988FFABC" box="[1119,1211,1321,1347]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, postdorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
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d). Elytra broad,
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<taxonomicName id="4C0B4D5F7838FFA2FEBAFAD39DB7FA94" box="[350,387,1361,1387]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
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very convex, apex (
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<figureCitation id="13302A597838FFA2FD67FAD39EE9FA94" box="[643,733,1361,1387]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, postdorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
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b) medially dentate, interneurs 2,4,6,8 with very large setigerous foveae, interneurs of fine closespaced punctures. Sternum VI (
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<figureCitation id="13302A597838FFA2FB5AFAFB9922FA6C" box="[1214,1302,1401,1427]" captionStart="FIGURE 20. A" captionStartId="46.[264,367,1318,1342]" captionTargetBox="[434,1149,491,1285]" captionTargetId="figure@46.[433,1153,489,1291]" captionTargetPageId="46" captionText="FIGURE 20. A. schwarzeneggeri, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) middle leg, postdorsal aspect. Measures (mm): HL = 2.7; WE = 2.3; male femur diameter = 1.1; male SW = 4.0, female SW = 4.6." httpUri="https://zenodo.org/record/162834/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
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c) with deeply circular emargination in female, deeply vnotched in female. Aedeagus (see Notes). Size: medium, 14.0 to 17.0mm in length,
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<quantity id="4CF39B397838FFA2FCAAFA4B9FCDFA1C" box="[846,1017,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="4.88" metricValueMax="5.16" metricValueMin="4.6" pageId="45" pageNumber="46" unit="mm" value="4.88" valueMax="5.16" valueMin="4.6">4.6 to 5.16mm</quantity>
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in width.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577838FFA2FEA5FA739D87F8B4" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="8BB436DC7838FFA2FEA5FA73989AF954" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
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<emphasis id="B97FEACE7838FFA2FEA5FA739EABF9F4" bold="true" box="[321,671,1521,1547]" pageId="45" pageNumber="46">Other specimens examined.</emphasis>
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<collectingCountry id="F31C764C7838FFA2FD4DFA739F04F9F4" box="[681,816,1521,1547]" name="Costa Rica" pageId="45" pageNumber="46">Costa Rica</collectingCountry>
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: 3
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<typeStatus id="54B0887E7838FFA2FCB2FA739FFDF9F4" box="[854,969,1521,1547]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
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, females, from the Cordillera Volcánica Central Conservation Area.
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<collectingCountry id="F31C764C7838FFA2FD35F99B9F61F9CC" box="[721,853,1561,1587]" name="Costa Rica" pageId="45" pageNumber="46">Costa Rica</collectingCountry>
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, 1f, Cartago, Pavones
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<geoCoordinate id="EE3F501B7838FFA2FB86F99B98FDF9CC" box="[1122,1225,1561,1587]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.61667">83°37'W</geoCoordinate>
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<geoCoordinate id="EE3F501B7838FFA2FB35F99B9912F9CC" box="[1233,1318,1561,1587]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.95">9°57'N</geoCoordinate>
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, May, (F.T. Hovore)(
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<collectionCode id="ED1AAE197838FFA2FE1DF9C39E73F9A4" box="[505,583,1601,1627]" pageId="45" pageNumber="46">FTHC</collectionCode>
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||
: ADP70530), 1f,
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<taxonomicName id="4C0B4D5F7838FFA2FCFAF9C39FB9F9A4" box="[798,909,1601,1627]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
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, 3.0 km SEIICA/
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<taxonomicName id="4C0B4D5F7838FFA2FB85F9C39883F9A4" box="[1121,1207,1601,1627]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="51" phylum="Arthropoda" rank="species" species="catie">
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||
<collectionCode id="ED1AAE197838FFA2FB85F9C39883F9A4" LSID="urn:lsid:biocol.org:col:15110" box="[1121,1207,1601,1627]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15110" name="Tropical Agricultural Research and Training Center (CATIE)" pageId="45" pageNumber="46">CATIE</collectionCode>
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||
</taxonomicName>
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,
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<geoCoordinate id="EE3F501B7838FFA2FB22F9C39919F9A4" box="[1222,1325,1601,1627]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-83.63333">83°38'W</geoCoordinate>
|
||
<geoCoordinate id="EE3F501B7838FFA2FEECF9EB9D69F97C" box="[264,349,1641,1667]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="9.883333">9°53'N</geoCoordinate>
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, April, (E. Giesbert)(
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<collectionCode id="ED1AAE197838FFA2FD8DF9EB9E82F97C" box="[617,694,1641,1667]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="45" pageNumber="46">FSCA</collectionCode>
|
||
: ADP4420), 1f, August, (D.R. Whitehead)(
|
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<collectionCode id="ED1AAE197838FFA2FB36F9EB991DF97C" LSID="urn:lsid:biocol.org:col:34871" box="[1234,1321,1641,1667]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46">USNM</collectionCode>
|
||
: ADP61717),
|
||
<quantity id="4CF39B397838FFA2FE40F9139DF8F954" box="[420,460,1681,1707]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="m" value="1.0">1m</quantity>
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, February, (H.F. Howden & A. Howden)(
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<collectionCode id="ED1AAE197838FFA2FC52F913983BF954" LSID="urn:lsid:biocol.org:col:34404" box="[950,1039,1681,1707]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34404" name="University of Alberta, E.H. Strickland Entomological Museum" pageId="45" pageNumber="46">UASM</collectionCode>
|
||
: ADP54794).
|
||
</paragraph>
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<paragraph id="8BB436DC7838FFA2FEDCF93B9D87F8B4" blockId="45.[264,1326,961,1867]" pageId="45" pageNumber="46">
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<emphasis id="B97FEACE7838FFA2FEDCF93B9DCBF92C" bold="true" box="[312,511,1721,1747]" pageId="45" pageNumber="46">Specific epithet.</emphasis>
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The specific epithet,
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<taxonomicName id="4C0B4D5F7838FFA2FD1BF9389FF2F92C" box="[767,966,1722,1747]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schwarzeneggeri">
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<emphasis id="B97FEACE7838FFA2FD1BF9389FF2F92C" box="[767,966,1722,1747]" italics="true" pageId="45" pageNumber="46">schwarzeneggeri</emphasis>
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</taxonomicName>
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is the Latinized genitive form of the family name of the actor, Arnold Schwarzenegger, in reference to the markedly developed (bicepslike) middle femora of the males of this species reminiscent of the actor’s physic.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3116557783BFFA1FEDCFE999EDBFE51" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BB436DC783BFFA1FEDCFE999EDBFE51" blockId="46.[264,1323,283,430]" pageId="46" pageNumber="47">
|
||
<emphasis id="B97FEACE783BFFA1FEDCFE999DB1FECA" bold="true" box="[312,389,283,309]" pageId="46" pageNumber="47">Notes.</emphasis>
|
||
The genitalia of the dissected male is missing from the vial pinned beneath the specimen, thus has
|
||
<taxonomicName id="4C0B4D5F783BFFA1FE0FFEC69E24FEA1" box="[491,528,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
|
||
been illustrated. The aedeagus of
|
||
<taxonomicName id="4C0B4D5F783BFFA1FC44FEC698C9FEA1" authority="Straneo 1982" authorityName="Straneo" authorityYear="1982" box="[928,1277,324,350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="perrinae">
|
||
<emphasis id="B97FEACE783BFFA1FC44FEC69873FEA2" box="[928,1095,324,349]" italics="true" pageId="46" pageNumber="47">Agra perrinae</emphasis>
|
||
(Straneo 1982)
|
||
</taxonomicName>
|
||
has a moderately long ostium, apex of phallus a simple diamondshaped lobe which is markedly arched dorsally, parameres glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |