162 lines
18 KiB
XML
162 lines
18 KiB
XML
<document id="C1527329A9BFE824A2F80766D92D61DE" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7832FFA7FEECFD2E9922FBD1" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra santarosa Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="41" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="40" updateTime="1698190273211" updateUser="plazi">
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<mods:titleInfo id="949A0C3BBEB46A9A9B26B25243015416">
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<mods:title id="4B2064CB12DDCD630F9597C5120579B3">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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<mods:name id="3B34284FFE148643CE47EA5D4D4B99A3" type="personal">
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<mods:roleTerm id="89B0E9785D24BAC722BAA6D0D9B6A5C8">Author</mods:roleTerm>
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<mods:namePart id="8CE3C6A3C62B1500174700A4B9356B47">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="EE9D20AA643C3C83E7ACC4C8CE863FBC">text</mods:typeOfResource>
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<mods:title id="998CF1D143D5C4CB7A5002A5CB32E505">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C9F8F960ACCC93CC94B1E846F6973F39">
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<mods:date id="14821F761C4303D98D1ECF08C3EAAFAB">2002</mods:date>
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<mods:detail id="951E6ACF6AAE25D7938BF8B58287F083" type="volume">
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<mods:number id="7A77DED17C5D576DBE0AA9477671051C">119</mods:number>
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<mods:classification id="7AFFC87DE379A1DEB2A79CB6609247CE">journal article</mods:classification>
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<mods:identifier id="787C4D94D986939CD6C1DF73E3C1C81C" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="E8708E326D8C9D70E6F8044A926CABD2" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="C762EDDAAE83660C7B5842ED9ED9040F" type="ISSN">11755326</mods:identifier>
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<treatment id="03A287CA7832FFA7FEECFD2E9922FBD1" ID-DOI="http://doi.org/10.5281/zenodo.6277996" ID-GBIF-Taxon="119330628" ID-Zenodo-Dep="6277996" LSID="urn:lsid:plazi:treatment:03A287CA7832FFA7FEECFD2E9922FBD1" httpUri="http://treatment.plazi.org/id/03A287CA7832FFA7FEECFD2E9922FBD1" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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<subSubSection id="C31165577832FFA8FEECFD2E9E02FD11" pageId="39" pageNumber="40" type="nomenclature">
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<paragraph id="8BB436DC7832FFA8FEECFD2E9E81FD3A" blockId="39.[264,693,683,750]" box="[264,693,683,710]" pageId="39" pageNumber="40">
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<heading id="D0FC81B07832FFA8FEECFD2E9E81FD3A" bold="true" box="[264,693,683,710]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
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<emphasis id="B97FEACE7832FFA8FEECFD2E9E81FD3A" bold="true" box="[264,693,683,710]" pageId="39" pageNumber="40">
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<taxonomicName id="4C0B4D5F7832FFA8FEECFD2E9E2CFD3A" ID-CoL="B2X3" authority="Erwin" authorityName="Erwin" box="[264,536,683,710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="santarosa" status="sp. nov.">
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<emphasis id="B97FEACE7832FFA8FEECFD2E9DF5FD39" bold="true" box="[264,449,684,710]" italics="true" pageId="39" pageNumber="40">Agra santarosa</emphasis>
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Erwin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24C57B57832FFA8FDC2FD299E81FD3A" box="[550,693,683,709]" pageId="39" pageNumber="40" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC7832FFA8FEECFD569E02FD11" blockId="39.[264,693,683,750]" box="[264,566,724,750]" pageId="39" pageNumber="40">
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(
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<figureCitation id="13302A597832FFA8FEF5FD569D9EFD11" box="[273,426,724,750]" captionStart="FIGURE 16. A" captionStartId="39.[264,368,1797,1821]" captionTargetBox="[406,1180,972,1763]" captionTargetId="figure@39.[405,1181,969,1770]" captionTargetPageId="39" captionText="FIGURE 16. A. santarosa, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0." httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="39" pageNumber="40">Figs. 16a, 16</figureCitation>
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b, 16c, 16d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577832FFA8FEECFCA198B0FC99" pageId="39" pageNumber="40" type="materials_examined">
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<paragraph id="8BB436DC7832FFA8FEECFCA198B0FC99" blockId="39.[264,1324,803,870]" pageId="39" pageNumber="40">
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<bibRef id="52ABA22A7832FFA8FEECFCA198B0FC99" pageId="39" pageNumber="40" type="book chapter">
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<emphasis id="B97FEACE7832FFA8FEECFCA19DB5FCC2" bold="true" box="[264,385,803,829]" pageId="39" pageNumber="40">
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<typeStatus id="54B0887E7832FFA8FEECFCA19D48FCC2" box="[264,380,803,829]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<author id="53B6DE397832FFA8FE6EFCA69EA6FCC2" box="[394,658,803,830]" fns="I" npo="LF" pageId="39" pageNumber="40">
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Male,
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<collectingCountry id="F31C764C7832FFA8FE33FCA19EA6FCC2" box="[471,658,803,829]" name="Costa Rica" pageId="39" pageNumber="40">
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<emphasis id="B97FEACE7832FFA8FE33FCA19EA6FCC2" bold="true" box="[471,658,803,829]" pageId="39" pageNumber="40">COSTA RICA</emphasis>
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</collectingCountry>
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</author>
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: Guanacaste, Santa Rosa National Park,
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<quantity id="4CF39B397832FFA8FB9EFCA698F4FCC1" box="[1146,1216,804,830]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="39" pageNumber="40" unit="m" value="280.0">280m</quantity>
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,
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<geoCoordinate id="EE3F501B7832FFA8FB29FCA69918FCC1" box="[1229,1324,804,830]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="10.833333">10º50'N</geoCoordinate>
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||
<geoCoordinate id="EE3F501B7832FFA8FEECFCCE9D59FC99" box="[264,365,844,870]" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="-85.61667">85º37'W</geoCoordinate>
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, May [
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<year id="FFA3778F7832FFA8FE59FCCE9DCBFC99" box="[445,511,844,870]" pageId="39" pageNumber="40">1980</year>
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],
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<title id="F90FDDEA7832FFA8FDF7FCCE9FB6FC99" box="[531,898,844,870]" jopScore="0.40974104" pageId="39" pageNumber="40" type="_" typeScore="32">(D.H. Janzen & W. Hallwachs)</title>
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(
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<journalOrPublisher id="51CA3C107832FFA8FC6EFCCE981FFC99" box="[906,1067,844,870]" jopScore="0.42491663" pageId="39" pageNumber="40">
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<collectionCode id="ED1AAE197832FFA8FC6EFCCE9FD7FC99" LSID="urn:lsid:biocol.org:col:34871" box="[906,995,844,870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40">USNM</collectionCode>
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: ADP
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</journalOrPublisher>
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<pagination id="B2C2C8287832FFA8FBC8FCCE984CFC99" box="[1068,1144,844,870]" pageId="39" pageNumber="40" type="pagination">55010</pagination>
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).
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</bibRef>
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF7466547832FFA8FEECF887993AF89E" httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="39" pageNumber="40" targetBox="[406,1180,972,1763]" targetPageId="39">
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<paragraph id="8BB436DC7832FFA8FEECF887993AF89E" blockId="39.[264,1323,1797,1889]" pageId="39" pageNumber="40">
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<emphasis id="B97FEACE7832FFA8FEECF8879DADF8E2" bold="true" box="[264,409,1797,1821]" pageId="39" pageNumber="40">FIGURE 16.</emphasis>
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<taxonomicName id="4C0B4D5F7832FFA8FE46F8879E1EF8E3" box="[418,554,1797,1820]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="santarosa">
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<emphasis id="B97FEACE7832FFA8FE46F8879E1EF8E3" box="[418,554,1797,1820]" italics="true" pageId="39" pageNumber="40">A. santarosa</emphasis>
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</taxonomicName>
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, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0.
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</paragraph>
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</caption>
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<subSubSection id="C3116557783DFFA7FEDCFE999E2FFE51" pageId="40" pageNumber="41" type="diagnosis">
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||
<paragraph id="8BB436DC783DFFA7FEDCFE999E2FFE51" blockId="40.[264,1324,283,1070]" pageId="40" pageNumber="41">
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<emphasis id="B97FEACE783DFFA7FEDCFE999D84FECA" bold="true" box="[312,432,283,309]" pageId="40" pageNumber="41">Diagnosis</emphasis>
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. Black. Femora rufous, tibiae rufopiceous, knees and tarsi piceous. Elytral interneurs with regularly spaced and slightly cribriform punctulae; apex with more or less similarsized lateral and sutural dentiform projections. Eyes very large, hemispherical, head short behind eyes.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3116557783DFFA7FEDCFE399E9DFCE9" pageId="40" pageNumber="41" type="description">
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||
<paragraph id="8BB436DC783DFFA7FEDCFE399E9DFCE9" blockId="40.[264,1324,283,1070]" pageId="40" pageNumber="41">
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<emphasis id="B97FEACE783DFFA7FEDCFE399DFDFE2A" bold="true" box="[312,457,443,469]" pageId="40" pageNumber="41">Description</emphasis>
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. Color and luster: Black. Antennomeres 411 bicolored, red and black, tibiae rufous. surface Shiny. Form: Labrum entire. Head (
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<figureCitation id="13302A59783DFFA7FC49FE66983BFE01" box="[941,1039,484,510]" captionStart="FIGURE 16. A" captionStartId="39.[264,368,1797,1821]" captionTargetBox="[406,1180,972,1763]" captionTargetId="figure@39.[405,1181,969,1770]" captionTargetPageId="39" captionText="FIGURE 16. A. santarosa, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0." httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="40" pageNumber="41">Fig. 16a</figureCitation>
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) behind eye moderately short, tapered in both sexes, more so in male. Prothorax "inflated" at basal third. Prothorax subtubular, short, robust, sutures effaced, dorsal surface flattened and densely punctate. Elytral apex (
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<figureCitation id="13302A59783DFFA7FE48FDDE9E36FD89" box="[428,514,604,630]" captionStart="FIGURE 16. A" captionStartId="39.[264,368,1797,1821]" captionTargetBox="[406,1180,972,1763]" captionTargetId="figure@39.[405,1181,969,1770]" captionTargetPageId="39" captionText="FIGURE 16. A. santarosa, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0." httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="40" pageNumber="41">Fig. 16</figureCitation>
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b) with more or less similar acute dentiform projections; interneurs with regularly spaced and slightly cribriform punctulae. Sternum VI (
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<figureCitation id="13302A59783DFFA7FBA2FD0698A9FD61" box="[1094,1181,644,670]" captionStart="FIGURE 16. A" captionStartId="39.[264,368,1797,1821]" captionTargetBox="[406,1180,972,1763]" captionTargetId="figure@39.[405,1181,969,1770]" captionTargetPageId="39" captionText="FIGURE 16. A. santarosa, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0." httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="40" pageNumber="41">Fig. 16</figureCitation>
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c) deeply vnotched in male, shallowly vnotched in female. Aedeagus (
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<figureCitation id="13302A59783DFFA7FC3AFD2E9800FD39" box="[990,1076,684,710]" captionStart="FIGURE 16. A" captionStartId="39.[264,368,1797,1821]" captionTargetBox="[406,1180,972,1763]" captionTargetId="figure@39.[405,1181,969,1770]" captionTargetPageId="39" captionText="FIGURE 16. A. santarosa, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.4; WE = 2.4; male SW = 4.6, female SW = 4.8; AL = 3.0." httpUri="https://zenodo.org/record/162826/files/figure.png" pageId="40" pageNumber="41">Fig. 16</figureCitation>
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d) with ostium elongate, apex a double scimitar, parameres glabrous. Size: moderately large, 17.5 to 20.0mm in length,
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<quantity id="4CF39B39783DFFA7FE9AFD7E9E0CFCE9" box="[382,568,764,790]" metricMagnitude="-3" metricUnit="m" metricValue="5.01" metricValueMax="5.16" metricValueMin="4.86" pageId="40" pageNumber="41" unit="mm" value="5.01" valueMax="5.16" valueMin="4.86">4.86 to 5.16mm</quantity>
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in width.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3116557783DFFA7FEDCFCA19EEFFBF9" pageId="40" pageNumber="41" type="materials_examined">
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||
<paragraph id="8BB436DC783DFFA7FEDCFCA1988AFC49" blockId="40.[264,1324,283,1070]" pageId="40" pageNumber="41">
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||
<emphasis id="B97FEACE783DFFA7FEDCFCA19EA6FCC2" bold="true" box="[312,658,803,829]" pageId="40" pageNumber="41">Other specimens examined.</emphasis>
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||
<collectingCountry id="F31C764C783DFFA7FD7DFCA69F2BFCC1" box="[665,799,804,830]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
|
||
: 2
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||
<typeStatus id="54B0887E783DFFA7FCA5FCA69F81FCC1" box="[833,949,804,830]" pageId="40" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
, both females, from the following Guanacaste Conservation Area.
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||
<collectingCountry id="F31C764C783DFFA7FD57FCCE9F03FC99" box="[691,823,844,870]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
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||
, Guanacaste, 1f, June, (D.H. Janzen & W. Hallwachs)(
|
||
<taxonomicName id="4C0B4D5F783DFFA7FE7CFCF69DD9FC71" box="[408,493,884,910]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
|
||
<collectionCode id="ED1AAE19783DFFA7FE7CFCF69DD9FC71" box="[408,493,884,910]" pageId="40" pageNumber="41">INBIO</collectionCode>
|
||
</taxonomicName>
|
||
: ADP55016), 1f, Estacíon Maritza, Volcán de Orosi, East slope,
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||
<quantity id="4CF39B39783DFFA7FB05FCF69913FC71" box="[1249,1319,884,910]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="40" pageNumber="41" unit="m" value="600.0">600m</quantity>
|
||
, LN 326900,373000, June, (GNP Biodiversity Survey)(
|
||
<taxonomicName id="4C0B4D5F783DFFA7FC69FC1E9FD7FC49" box="[909,995,924,950]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
|
||
<collectionCode id="ED1AAE19783DFFA7FC69FC1E9FD7FC49" box="[909,995,924,950]" pageId="40" pageNumber="41">INBIO</collectionCode>
|
||
</taxonomicName>
|
||
:
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||
<collectionCode id="ED1AAE19783DFFA7FC09FC1E9814FC49" LSID="urn:lsid:biocol.org:col:14323" box="[1005,1056,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="40" pageNumber="41">CRI</collectionCode>
|
||
000134050).
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||
</paragraph>
|
||
<paragraph id="8BB436DC783DFFA7FEDCFC419EEFFBF9" blockId="40.[264,1324,283,1070]" pageId="40" pageNumber="41">
|
||
<emphasis id="B97FEACE783DFFA7FEDCFC419DCBFC22" bold="true" box="[312,511,963,989]" pageId="40" pageNumber="41">Specific epithet.</emphasis>
|
||
The specific epithet,
|
||
<emphasis id="B97FEACE783DFFA7FD1BFC469F4DFC22" box="[767,889,964,989]" italics="true" pageId="40" pageNumber="41">
|
||
<taxonomicName id="4C0B4D5F783DFFA7FD1BFC469F41FC22" box="[767,885,964,989]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="santarosa">santarosa</taxonomicName>
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||
,
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||
</emphasis>
|
||
is a noun in apposition based on the
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<typeStatus id="54B0887E783DFFA7FEECFC6E9D0EFBF9" box="[264,314,1004,1030]" pageId="40" pageNumber="41">type</typeStatus>
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locality, Santa Rosa National Park.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116557783DFFA7FEDCFB919922FBD1" box="[312,1302,1043,1070]" pageId="40" pageNumber="41" type="description">
|
||
<paragraph id="8BB436DC783DFFA7FEDCFB919922FBD1" blockId="40.[264,1324,283,1070]" box="[312,1302,1043,1070]" pageId="40" pageNumber="41">
|
||
<emphasis id="B97FEACE783DFFA7FEDCFB919DB1FBD2" bold="true" box="[312,389,1043,1069]" pageId="40" pageNumber="41">Notes.</emphasis>
|
||
An illustration of famulagroup stylomere 2 is on the
|
||
<taxonomicName id="4C0B4D5F783DFFA7FBE4FB9698ACFBD2" box="[1024,1176,1044,1069]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="fortuna">
|
||
<emphasis id="B97FEACE783DFFA7FBE4FB9698ACFBD2" box="[1024,1176,1044,1069]" italics="true" pageId="40" pageNumber="41">Agra fortuna</emphasis>
|
||
</taxonomicName>
|
||
web page.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |