treatments-xml/data/03/A2/87/03A287CA782BFFB0FEECFC6E9F22FDD9.xml
2024-06-21 12:22:17 +02:00

172 lines
20 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="AA468C73A619FA8504D24533200E4FE6" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA782BFFB0FEECFC6E9F22FDD9" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra zumbado Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="64" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="63" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="EDAC514E7631E25BE546E3F10D88173E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3E3BBF8A066A1010A28452822F4F4F49">
<mods:title id="A83ACA5E7BED5E7412A9B85E8ED30F3C">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="79000F0D19BFF38E49898E08E85AF048" type="personal">
<mods:role id="91FAC3D05EC7DB927FFB19DC253A848C">
<mods:roleTerm id="81D233EC08D447800BE4640B3C27EA55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE6C095B48F03B84FBE49C6259660814">Erwin, Terry L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4E48B10F5BD4CD6E2BD4D8A4B792D0DB">text</mods:typeOfResource>
<mods:relatedItem id="C56D94D0592C61B7DDD645AA3C3943EB" type="host">
<mods:titleInfo id="2F888D5D16778764B30132A7482F198B">
<mods:title id="6F7C31EF1A314F77DAFEAF21F66DA0C0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2A50B5F28A397071E9AEFD9E2B59AB87">
<mods:date id="4139D352EC15D10B316E91155F33747F">2002</mods:date>
<mods:detail id="92382FD6AA777554C9FB0D409FDC5181" type="volume">
<mods:number id="4E56CCF4D1AB491DD1C2287F899EB103">119</mods:number>
</mods:detail>
<mods:extent id="E8C40AEBB050339785EFE420EBA108D2" unit="page">
<mods:start id="09CFF58AE2C3C347C1974751D7D76EE9">1</mods:start>
<mods:end id="29BDE71DA1759DB0556232D881677BD0">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B962BFAD45F8C747F35021D6612B1FEB">journal article</mods:classification>
<mods:identifier id="9172F85870CB1703C4581D353B9F99D0" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="C66C7486BF668B8A7733536F019E188D" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="27A7E393EEA86056B8E28C73A5FB4CEC" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="DEE11A824AA0B624D44FA7F36C2594A1" type="Zenodo-Dep">162803</mods:identifier>
</mods:mods>
<treatment id="03A287CA782BFFB0FEECFC6E9F22FDD9" ID-DOI="http://doi.org/10.5281/zenodo.6278036" ID-GBIF-Taxon="121065204" ID-Zenodo-Dep="6278036" LSID="urn:lsid:plazi:treatment:03A287CA782BFFB0FEECFC6E9F22FDD9" httpUri="http://treatment.plazi.org/id/03A287CA782BFFB0FEECFC6E9F22FDD9" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<subSubSection id="C3116557782BFFB1FEECFC6E9E02FBD1" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8BB436DC782BFFB1FEECFC6E9E99FBFA" blockId="62.[264,685,1003,1070]" box="[264,685,1003,1030]" pageId="62" pageNumber="63">
<heading id="D0FC81B0782BFFB1FEECFC6E9E99FBFA" bold="true" box="[264,685,1003,1030]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B97FEACE782BFFB1FEECFC6E9E99FBFA" bold="true" box="[264,685,1003,1030]" pageId="62" pageNumber="63">
<taxonomicName id="4C0B4D5F782BFFB1FEECFC6E9E24FBFA" ID-CoL="B339" authority="Erwin" authorityName="Erwin" box="[264,528,1003,1030]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="zumbado" status="sp. nov.">
<emphasis id="B97FEACE782BFFB1FEECFC6E9D8DFBF9" bold="true" box="[264,441,1004,1030]" italics="true" pageId="62" pageNumber="63">Agra zumbado</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B5782BFFB1FDF9FC699E99FBFA" box="[541,685,1003,1029]" pageId="62" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC782BFFB1FEECFB969E02FBD1" blockId="62.[264,685,1003,1070]" box="[264,566,1044,1070]" pageId="62" pageNumber="63">
(
<figureCitation id="13302A59782BFFB1FEF5FB969D9FFBD1" box="[273,427,1044,1070]" captionStart="FIGURE 30. A" captionStartId="63.[264,367,1550,1574]" captionTargetBox="[417,1175,620,1523]" captionTargetId="figure@63.[412,1175,609,1524]" captionTargetPageId="63" captionText="FIGURE 30. A. zumbado, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.7; WE = 1.8; male SW = 3.2, female SW = 3.4; AL = 3.0." httpUri="https://zenodo.org/record/162844/files/figure.png" pageId="62" pageNumber="63">Figs. 30a, 30</figureCitation>
b, 30c, 30d)
</paragraph>
</subSubSection>
<subSubSection id="C3116557782BFFB1FEECFBE19E4FFB31" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8BB436DC782BFFB1FEECFBE19E4FFB31" blockId="62.[264,1325,1123,1870]" pageId="62" pageNumber="63">
<emphasis id="B97FEACE782BFFB1FEECFBE19DB5FB82" bold="true" box="[264,385,1123,1149]" pageId="62" pageNumber="63">
<typeStatus id="54B0887E782BFFB1FEECFBE19D48FB82" box="[264,380,1123,1149]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingCountry id="F31C764C782BFFB1FE33FBE69EBDFB81" box="[471,649,1124,1150]" name="Costa Rica" pageId="62" pageNumber="63">COSTA RICA</collectingCountry>
: Guanacaste, Guanacaste National Park, Estacíon
<taxonomicName id="4C0B4D5F782BFFB1FB3AFBE69913FB81" box="[1246,1319,1124,1150]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, 9.0 km S Santa Cecilia,
<quantity id="4CF39B39782BFFB1FDB2FB0E9EA8FB59" box="[598,668,1164,1190]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="62" pageNumber="63" unit="m" value="700.0">700m</quantity>
,
<geoCoordinate id="EE3F501B782BFFB1FD56FB0E9F08FB59" box="[690,828,1164,1190]" direction="north" orientation="latitude" pageId="62" pageNumber="63" precision="15" value="10.990556">10°59'26&quot;N</geoCoordinate>
<geoCoordinate id="EE3F501B782BFFB1FCAAFB0E9FEAFB59" box="[846,990,1164,1190]" direction="west" orientation="longitude" pageId="62" pageNumber="63" precision="15" value="-85.42777">85°25'40&quot;W</geoCoordinate>
, April, (C. Moraga &amp; P. Rios)(
<taxonomicName id="4C0B4D5F782BFFB1FEB5FB369DABFB31" box="[337,415,1204,1230]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19782BFFB1FEB5FB369DABFB31" box="[337,415,1204,1230]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="62" pageNumber="63">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19782BFFB1FE4EFB369DEAFB31" LSID="urn:lsid:biocol.org:col:14323" box="[426,478,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="62" pageNumber="63">CRI</collectionCode>
000­166003).
</paragraph>
</subSubSection>
<subSubSection id="C3116557782BFFB1FEDCFB599FC2FAE1" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8BB436DC782BFFB1FEDCFB599FC2FAE1" blockId="62.[264,1325,1123,1870]" pageId="62" pageNumber="63">
<emphasis id="B97FEACE782BFFB1FEDCFB599D8CFB0A" bold="true" box="[312,440,1243,1269]" pageId="62" pageNumber="63">Diagnosis.</emphasis>
This is the only species of Costa Rican
<taxonomicName id="4C0B4D5F782BFFB1FC7AFB5E9FEDFB0A" box="[926,985,1244,1269]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEACE782BFFB1FC7AFB5E9FEDFB0A" box="[926,985,1244,1269]" italics="true" pageId="62" pageNumber="63">Agra</emphasis>
</taxonomicName>
with ferrugineus venter and appendages and bright brassy elytra and tri­dentate elytral apex.
</paragraph>
</subSubSection>
<subSubSection id="C3116557782BFFB1FEDCFAA99FCBF951" pageId="62" pageNumber="63" type="description">
<paragraph id="8BB436DC782BFFB1FEDCFAA99FCBF951" blockId="62.[264,1325,1123,1870]" pageId="62" pageNumber="63">
<emphasis id="B97FEACE782BFFB1FEDCFAA99DE4FABA" bold="true" box="[312,464,1323,1349]" pageId="62" pageNumber="63">Description.</emphasis>
Color and luster: With ferruginous venter and appendages and bright brassy elytra. Form: Head (
<figureCitation id="13302A59782BFFB1FDB3FAD69E89FA91" box="[599,701,1364,1390]" captionStart="FIGURE 30. A" captionStartId="63.[264,367,1550,1574]" captionTargetBox="[417,1175,620,1523]" captionTargetId="figure@63.[412,1175,609,1524]" captionTargetPageId="63" captionText="FIGURE 30. A. zumbado, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.7; WE = 1.8; male SW = 3.2, female SW = 3.4; AL = 3.0." httpUri="https://zenodo.org/record/162844/files/figure.png" pageId="62" pageNumber="63">Fig. 30a</figureCitation>
) behind eyes short, quadrate with broadly rounded hind angles. Elytron with striatio­punctate interneurs, 2, 4, and 6 with setigerous foveae; apex (
<figureCitation id="13302A59782BFFB1FEB6FA269D9EFA41" box="[338,426,1444,1470]" captionStart="FIGURE 30. A" captionStartId="63.[264,367,1550,1574]" captionTargetBox="[417,1175,620,1523]" captionTargetId="figure@63.[412,1175,609,1524]" captionTargetPageId="63" captionText="FIGURE 30. A. zumbado, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.7; WE = 1.8; male SW = 3.2, female SW = 3.4; AL = 3.0." httpUri="https://zenodo.org/record/162844/files/figure.png" pageId="62" pageNumber="63">Fig. 30</figureCitation>
b) slightly oblique, lateral tooth small, sutural apex rounded,
<taxonomicName id="4C0B4D5F782BFFB1FB6AFA269887FA41" box="[1166,1203,1444,1470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
projected acutely, margin slightly lobed at middle. Male metasternum, abdominal sterna 2­6, and trochanters densely pubescent. Middle tibia of male with sparse medial brush. Sternum VI (
<figureCitation id="13302A59782BFFB1FEF5F99E9D5DF9C9" box="[273,361,1564,1590]" captionStart="FIGURE 30. A" captionStartId="63.[264,367,1550,1574]" captionTargetBox="[417,1175,620,1523]" captionTargetId="figure@63.[412,1175,609,1524]" captionTargetPageId="63" captionText="FIGURE 30. A. zumbado, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.7; WE = 1.8; male SW = 3.2, female SW = 3.4; AL = 3.0." httpUri="https://zenodo.org/record/162844/files/figure.png" pageId="62" pageNumber="63">Fig. 30</figureCitation>
c) of male shallowly V­shaped with acute posterio­lateral angles, that of female deeply notched medially with obtuse posterio­lateral angles. Aedeagus (
<figureCitation id="13302A59782BFFB1FB9EF9C698E7F9A1" box="[1146,1235,1604,1630]" captionStart="FIGURE 30. A" captionStartId="63.[264,367,1550,1574]" captionTargetBox="[417,1175,620,1523]" captionTargetId="figure@63.[412,1175,609,1524]" captionTargetPageId="63" captionText="FIGURE 30. A. zumbado, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 1.7; WE = 1.8; male SW = 3.2, female SW = 3.4; AL = 3.0." httpUri="https://zenodo.org/record/162844/files/figure.png" pageId="62" pageNumber="63">Fig. 30</figureCitation>
d) with ostium elongate; apex of aedeagus a broad medieval axe­shaped lobe, parameres glabrous. Size: Small: 11.0 to 12.0mm in length,
<quantity id="4CF39B39782BFFB1FD31F9169FBBF951" box="[725,911,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.62" metricValueMin="3.48" pageId="62" pageNumber="63" unit="mm" value="3.55" valueMax="3.62" valueMin="3.48">3.48 to 3.62mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C3116557782BFFB0FEDCF9399DCFFE51" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8BB436DC782BFFB1FEDCF9399FA4F8B1" blockId="62.[264,1325,1123,1870]" pageId="62" pageNumber="63">
<emphasis id="B97FEACE782BFFB1FEDCF9399EACF92A" bold="true" box="[312,664,1723,1749]" pageId="62" pageNumber="63">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C782BFFB1FD47F93E9F1FF929" box="[675,811,1724,1750]" name="Costa Rica" pageId="62" pageNumber="63">Costa Rica</collectingCountry>
: 3
<typeStatus id="54B0887E782BFFB1FCB7F93E9FF3F929" box="[851,967,1724,1750]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D0DFD55782BFFB1FC33F93E9877F929" box="[983,1091,1724,1750]" pageId="62" pageNumber="63" type="female">1 female</specimenCount>
,
<specimenCount id="9D0DFD55782BFFB1FBB6F93E9887F929" box="[1106,1203,1724,1750]" pageId="62" pageNumber="63" type="male">2 males</specimenCount>
, from the Guanacaste Conservation Area.
<collectingCountry id="F31C764C782BFFB1FD6BF9669F22F901" box="[655,790,1764,1790]" name="Costa Rica" pageId="62" pageNumber="63">Costa Rica</collectingCountry>
, Guanacaste,
<quantity id="4CF39B39782BFFB1FC24F9669FDCF901" box="[960,1000,1764,1790]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="m" value="1.0">1m</quantity>
, March, (C. Moraga &amp; R. Blanco)(
<taxonomicName id="4C0B4D5F782BFFB1FE8BF88E9DF1F8D9" box="[367,453,1804,1830]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19782BFFB1FE8BF88E9DF1F8D9" box="[367,453,1804,1830]" pageId="62" pageNumber="63">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19782BFFB1FE3DF88E9E3FF8D9" LSID="urn:lsid:biocol.org:col:14323" box="[473,523,1804,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="62" pageNumber="63">CRI</collectionCode>
000­211532), 1f, March ­ April, (C. Moraga)(
<taxonomicName id="4C0B4D5F782BFFB1FBB9F88E9886F8D9" box="[1117,1202,1804,1830]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19782BFFB1FBB9F88E9886F8D9" box="[1117,1202,1804,1830]" pageId="62" pageNumber="63">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19782BFFB1FB22F88E98CCF8D9" LSID="urn:lsid:biocol.org:col:14323" box="[1222,1272,1804,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="62" pageNumber="63">CRI</collectionCode>
000­ 748607),
<quantity id="4CF39B39782BFFB1FE9CF8B69D94F8B1" box="[376,416,1844,1870]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="m" value="1.0">1m</quantity>
, May, (P. Rios)(
<taxonomicName id="4C0B4D5F782BFFB1FDBAF8B69E81F8B1" box="[606,693,1844,1870]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19782BFFB1FDBAF8B69E81F8B1" box="[606,693,1844,1870]" pageId="62" pageNumber="63">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19782BFFB1FD5BF8B69EC7F8B1" LSID="urn:lsid:biocol.org:col:14323" box="[703,755,1844,1870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="62" pageNumber="63">CRI</collectionCode>
001­877475).
</paragraph>
<paragraph id="8BB436DC782AFFB0FEDCFE999DCFFE51" blockId="63.[264,1324,283,550]" pageId="63" pageNumber="64">
<emphasis id="B97FEACE782AFFB0FEDCFE999DCBFECA" bold="true" box="[312,511,283,309]" pageId="63" pageNumber="64">Specific epithet.</emphasis>
The specific epithet,
<taxonomicName id="4C0B4D5F782AFFB0FD1AFE9E9F5CFECA" box="[766,872,284,309]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zumbado">
<emphasis id="B97FEACE782AFFB0FD1AFE9E9F5CFECA" box="[766,872,284,309]" italics="true" pageId="63" pageNumber="64">zumbado</emphasis>
</taxonomicName>
, is used as a noun in apposition based on the family name of the collector of many interesting
<taxonomicName id="4C0B4D5F782AFFB0FC4FFEC69FD2FEA2" box="[939,998,324,349]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEACE782AFFB0FC4FFEC69FD2FEA2" box="[939,998,324,349]" italics="true" pageId="63" pageNumber="64">Agra</emphasis>
</taxonomicName>
specimens, ex­parataxonomist, and now
<taxonomicName id="4C0B4D5F782AFFB0FE27FEEE9E38FE79" box="[451,524,364,390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19782AFFB0FE27FEEE9E38FE79" box="[451,524,364,390]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="63" pageNumber="64">INBio</collectionCode>
</taxonomicName>
Inventory chief, Manuel
<taxonomicName id="4C0B4D5F782AFFB0FCADFEEE9FD6FE79" box="[841,994,364,390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="2" phylum="Arthropoda" rank="species" species="zumbado" status="sp. nov.">A. Zumbado</taxonomicName>
of San Antonio de Belén, Heredia,
<collectingCountry id="F31C764C782AFFB0FE97FE169DC2FE51" box="[371,502,404,430]" name="Costa Rica" pageId="63" pageNumber="64">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3116557782AFFB0FEDCFE399F22FDD9" pageId="63" pageNumber="64" type="description">
<paragraph id="8BB436DC782AFFB0FEDCFE399F22FDD9" blockId="63.[264,1324,283,550]" pageId="63" pageNumber="64">
<emphasis id="B97FEACE782AFFB0FEDCFE399DB1FE2A" bold="true" box="[312,389,443,469]" pageId="63" pageNumber="64">Notes.</emphasis>
This species features all the external attributes of typical
<taxonomicName id="4C0B4D5F782AFFB0FBCDFE3E98A5FE29" box="[1065,1169,444,470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="purpurea">purpurea</taxonomicName>
group members, however, I have classified it in its own subgroup because of the shorter stylomere 2 and the broad­ax shape of the aedeagal apex.
</paragraph>
</subSubSection>
</treatment>
</document>