129 lines
15 KiB
XML
129 lines
15 KiB
XML
<document id="92B8F8851935FF6D3656404CFB8AB421" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="11755326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7826FFBBFEECFEEE9DDDFD61" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra pia Liebke" docType="treatment" docVersion="11" lastPageNumber="53" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="52" updateTime="1698190273211" updateUser="plazi">
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<mods:title id="CC1AD828F88F8E2E2E2BF0A2770FF0F1">The Beetle Family Carabidae of Costa Rica: Twentynine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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<mods:name id="2DF1F4EC2828C866DFA9110DCA63DD31" type="personal">
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<mods:roleTerm id="9AF0E3F6D4433841EA3DC01306EB1CC5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6D5FED8A2A451D92D81F4F4F9FE8450">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="2E2861E33F6F9801705BC75E7D5AD8A9">text</mods:typeOfResource>
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<mods:title id="230E2E4B5DC700AA5B3530415B3289C1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A6A986A2291CEEE6EA8853588006FBC8">
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<mods:date id="D6F1BD4CAA0843B219B3756810533BB7">2002</mods:date>
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<mods:detail id="8F68E9349B175362E632E40E31B07319" type="volume">
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<mods:number id="D821E87C3364F93DC038512DB01BFEA1">119</mods:number>
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</mods:detail>
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<mods:start id="E13DA147660132C0187366E48A95F7C7">1</mods:start>
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<mods:end id="D3694578C8B8847A05EDBD49E1EB7119">68</mods:end>
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<mods:classification id="DC66AAB19B1329B9CFA3CB6CB664DBD7">journal article</mods:classification>
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<mods:identifier id="0CDCFF454655A6762E62A960623C2A5E" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<mods:identifier id="D2D575296B764F5B930889116C02EADA" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<mods:identifier id="6C0028CCE7F40E73FF207E2CF9B62817" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="38CCC21028C4F3B079F42375FABF15D1" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA7826FFBBFEECFEEE9DDDFD61" ID-DOI="http://doi.org/10.5281/zenodo.6278021" ID-GBIF-Taxon="121065205" ID-Zenodo-Dep="6278021" LSID="urn:lsid:plazi:treatment:03A287CA7826FFBBFEECFEEE9DDDFD61" httpUri="http://treatment.plazi.org/id/03A287CA7826FFBBFEECFEEE9DDDFD61" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<subSubSection id="C31165577826FFBCFEECFEEE9E01FE51" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8BB436DC7826FFBCFEECFEEE9DFBFE7A" blockId="51.[264,565,363,430]" box="[264,463,363,390]" pageId="51" pageNumber="52">
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<heading id="D0FC81B07826FFBCFEECFEEE9DFBFE7A" bold="true" box="[264,463,363,390]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
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<taxonomicName id="4C0B4D5F7826FFBCFEECFEEE9DFBFE7A" ID-CoL="664M7" authority="Liebke" authorityName="Liebke" box="[264,463,363,390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pia">
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<emphasis id="B97FEACE7826FFBCFEECFEEE9DFBFE7A" bold="true" box="[264,463,363,390]" pageId="51" pageNumber="52">
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<emphasis id="B97FEACE7826FFBCFEECFEEE9D45FE79" bold="true" box="[264,369,364,390]" italics="true" pageId="51" pageNumber="52">Agra pia</emphasis>
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Liebke
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BB436DC7826FFBCFEECFE169E01FE51" blockId="51.[264,565,363,430]" box="[264,565,404,430]" pageId="51" pageNumber="52">
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(
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<figureCitation id="13302A597826FFBCFEF5FE169D9FFE51" box="[273,427,404,430]" captionStart="FIGURE 23. A" captionStartId="51.[264,367,1630,1654]" captionTargetBox="[459,1134,841,1596]" captionTargetId="figure@51.[449,1137,833,1603]" captionTargetPageId="51" captionText="FIGURE 23. A. pia, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1." httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="51" pageNumber="52">Figs. 23a, 23</figureCitation>
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b, 23c, 23d)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577826FFBCFEECFE669889FDE4" pageId="51" pageNumber="52" type="reference_group">
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<paragraph id="8BB436DC7826FFBCFEECFE669889FDE4" blockId="51.[264,1322,483,539]" pageId="51" pageNumber="52">
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<taxonomicName id="4C0B4D5F7826FFBCFEECFE669E1FFE04" ID-CoL="664M7" authority="Liebke 1940:236" authorityName="Liebke" authorityPageNumber="236" authorityYear="1940" box="[264,555,484,507]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pia">
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<emphasis id="B97FEACE7826FFBCFEECFE669D5CFE04" box="[264,360,484,507]" italics="true" pageId="51" pageNumber="52">Agra pia</emphasis>
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Liebke 1940:236
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</taxonomicName>
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.
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<typeStatus id="54B0887E7826FFBCFDD2FE669EAAFE04" box="[566,670,484,507]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
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: Presumed lost.
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<typeStatus id="54B0887E7826FFBCFCABFE619F98FE04" box="[847,940,483,507]" pageId="51" pageNumber="52" type="neotype">
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<emphasis id="B97FEACE7826FFBCFCABFE619F98FE04" bold="true" box="[847,940,483,507]" pageId="51" pageNumber="52">Neotype</emphasis>
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</typeStatus>
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, here designated,
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<collectingCountry id="F31C764C7826FFBCFB96FE6698D9FE04" box="[1138,1261,484,507]" name="Costa Rica" pageId="51" pageNumber="52">Costa Rica</collectingCountry>
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: Puntarenas, Las Cruces,
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<geoCoordinate id="EE3F501B7826FFBCFDF1FD869E5FFDE4" box="[533,619,516,539]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="8.766666">08º46'N</geoCoordinate>
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<geoCoordinate id="EE3F501B7826FFBCFD96FD869EE7FDE4" box="[626,723,516,539]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="-82.95">82º 57'W</geoCoordinate>
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, February (F.T. Hovore)(
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<taxonomicName id="4C0B4D5F7826FFBCFC04FD869812FDE4" ID-CoL="B2P8" box="[992,1062,516,539]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">INBio</taxonomicName>
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: ADP
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<date id="FFB5101C7826FFBCFB83FD869883FDE4" box="[1127,1207,516,539]" pageId="51" pageNumber="52" value="15-10-05">100515</date>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577826FFBCFEECFDC99D7BFCF9" pageId="51" pageNumber="52" type="diagnosis">
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<paragraph id="8BB436DC7826FFBCFEECFDC99D7BFCF9" blockId="51.[264,1323,587,774]" pageId="51" pageNumber="52">
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<emphasis id="B97FEACE7826FFBCFEECFDC99DBCFD9A" bold="true" box="[264,392,587,613]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
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<taxonomicName id="4C0B4D5F7826FFBCFE77FDCE9DC9FD9A" box="[403,509,588,613]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pia">
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<emphasis id="B97FEACE7826FFBCFE77FDCE9DC9FD9A" box="[403,509,588,613]" italics="true" pageId="51" pageNumber="52">Agra pia</emphasis>
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</taxonomicName>
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is the only species in
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<collectingCountry id="F31C764C7826FFBCFCF1FDCE9FACFD99" box="[789,920,588,614]" name="Costa Rica" pageId="51" pageNumber="52">Costa Rica</collectingCountry>
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whose adults have the following attributes: Metallic purple elytra with slightly prominent sutural tooth. Pronotal disc with raised median keel on the anterior portion. Black, pronotum with light greenish brasslike luster, elytra purple, with light green brasslike luster. Antennae, legs and ventral side black.
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</paragraph>
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</subSubSection>
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<caption id="DF7466547826FFBCFEECF9DC9915F945" httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="51" pageNumber="52" targetBox="[459,1134,841,1596]" targetPageId="51">
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<paragraph id="8BB436DC7826FFBCFEECF9DC9915F945" blockId="51.[264,1323,1630,1722]" pageId="51" pageNumber="52">
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<emphasis id="B97FEACE7826FFBCFEECF9DC9DAEF989" bold="true" box="[264,410,1630,1654]" pageId="51" pageNumber="52">FIGURE 23.</emphasis>
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<taxonomicName id="4C0B4D5F7826FFBCFE47F9DC9DD0F98A" box="[419,484,1630,1653]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pia">
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<emphasis id="B97FEACE7826FFBCFE47F9DC9DD0F98A" box="[419,484,1630,1653]" italics="true" pageId="51" pageNumber="52">A. pia</emphasis>
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</taxonomicName>
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, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1.
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</paragraph>
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</caption>
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<subSubSection id="C31165577826FFBBFEDCF96F9D67FDD9" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
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<paragraph id="8BB436DC7826FFBBFEDCF96F9D67FDD9" blockId="51.[264,1323,1773,1880]" lastBlockId="52.[264,1324,284,670]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<emphasis id="B97FEACE7826FFBCFEDCF96F9DE4F8F8" bold="true" box="[312,464,1773,1799]" pageId="51" pageNumber="52">Description.</emphasis>
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Color and luster: Black. Antennae, legs and ventral side black. Shiny, pronotum with light greenish brasslike luster, elytra purple, with light green brasslike luster. Form: Female (see Notes). Head (
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<figureCitation id="13302A597826FFBCFD37F8BC9F0DF8A7" box="[723,825,1854,1880]" captionStart="FIGURE 23. A" captionStartId="51.[264,367,1630,1654]" captionTargetBox="[459,1134,841,1596]" captionTargetId="figure@51.[449,1137,833,1603]" captionTargetPageId="51" captionText="FIGURE 23. A. pia, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1." httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="51" pageNumber="52">Fig. 23a</figureCitation>
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) behind eyes short and quadrate, slightly tapered in male, vertex dimpled dorsally. Elytron (
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<figureCitation id="13302A597821FFBBFCB7FE9E9F93FEC9" box="[851,935,284,310]" captionStart="FIGURE 23. A" captionStartId="51.[264,367,1630,1654]" captionTargetBox="[459,1134,841,1596]" captionTargetId="figure@51.[449,1137,833,1603]" captionTargetPageId="51" captionText="FIGURE 23. A. pia, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1." httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="52" pageNumber="53">Fig. 23</figureCitation>
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b) with apex slightly oblique, lateral tooth small, sutural tooth produced, acute; interneurs of very fine nearly contiguous punctulae; intervals slightly convex on disc. Male venter without accessory patches of setae. Sternum VI (
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<figureCitation id="13302A597821FFBBFE11FE169E79FE51" box="[501,589,404,430]" captionStart="FIGURE 23. A" captionStartId="51.[264,367,1630,1654]" captionTargetBox="[459,1134,841,1596]" captionTargetId="figure@51.[449,1137,833,1603]" captionTargetPageId="51" captionText="FIGURE 23. A. pia, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1." httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="52" pageNumber="53">Fig. 23</figureCitation>
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c) deeply emarginate in male, narrowly vnotched in female. Aedeagus (
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<figureCitation id="13302A597821FFBBFE74FE3E9DDCFE29" box="[400,488,444,470]" captionStart="FIGURE 23. A" captionStartId="51.[264,367,1630,1654]" captionTargetBox="[459,1134,841,1596]" captionTargetId="figure@51.[449,1137,833,1603]" captionTargetPageId="51" captionText="FIGURE 23. A. pia, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom scanned from Liebke, 1940), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.1; WE = 1.5; male SW = 2.8, AL = 3.1." httpUri="https://zenodo.org/record/162837/files/figure.png" pageId="52" pageNumber="53">Fig. 23</figureCitation>
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d) with ostium elongate, slightly constricted medially, apex a simple cordiform lobe, parameres glabrous. Size: small,
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<quantity id="4CF39B397821FFBBFCBBFE669FF4FE01" box="[863,960,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="52" pageNumber="53" unit="mm" value="12.9">12.9mm</quantity>
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in length,
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<quantity id="4CF39B397821FFBBFBACFE6698AFFE01" box="[1096,1179,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="52" pageNumber="53" unit="mm" value="3.2">3.2mm</quantity>
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in width in width.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31165577821FFBBFEDCFDB19DDDFD61" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="8BB436DC7821FFBBFEDCFDB19FD6FDB1" blockId="52.[264,1324,284,670]" box="[312,994,563,590]" pageId="52" pageNumber="53">
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<emphasis id="B97FEACE7821FFBBFEDCFDB19EA4FDB2" bold="true" box="[312,656,563,589]" pageId="52" pageNumber="53">Other specimens examined.</emphasis>
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Only the
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<typeStatus id="54B0887E7821FFBBFCE0FDB69F53FDB1" box="[772,871,564,590]" pageId="52" pageNumber="53" type="neotype">Neotype</typeStatus>
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is known.
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</paragraph>
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<paragraph id="8BB436DC7821FFBBFEDCFDD99DDDFD61" blockId="52.[264,1324,284,670]" pageId="52" pageNumber="53">
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<emphasis id="B97FEACE7821FFBBFEDCFDD99DB1FD8A" bold="true" box="[312,389,603,629]" pageId="52" pageNumber="53">Notes.</emphasis>
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Liebke’s description was adequate to allow recognition of this species without access to the
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<typeStatus id="54B0887E7821FFBBFE47FD069DD1FD61" box="[419,485,644,670]" pageId="52" pageNumber="53">types</typeStatus>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |