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<document id="F4D52C39AF89F241743D89597A61CC3B" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7821FFBAFEECFB369DF7FC71" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra pitilla Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="54" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="53" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="3CB7A9E6FDF53B97F98CFCDC87639424" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E37BA37612F203D8DE0941322C6514D8">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="68226ADFBFCE848B8EA7918AB7E0152B" type="personal">
<mods:role id="420E6363654EA1DE731882ABD8C28E53">
<mods:roleTerm id="ADF6D13D9DE6AF384EE130EF25F016E5">Author</mods:roleTerm>
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<mods:namePart id="F79EF36324BC057204B1EBEB1C5C2CDA">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="3374C47CA39E7AC51DDCB6DAE3B68089">text</mods:typeOfResource>
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<mods:title id="B547E5C1937E9724FA3117827FE7F660">Zootaxa</mods:title>
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<mods:part id="028ADF4638DB4F81564EB31F4A8981D2">
<mods:date id="CC470632FC8E77675E98957112A09645">2002</mods:date>
<mods:detail id="9D636CFA48C285D569719C2A038C967D" type="volume">
<mods:number id="A898E330AEDB1151AC0CBB28DF60CF90">119</mods:number>
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<mods:classification id="CA800ADD0896D2CEE9023EA50A8D933D">journal article</mods:classification>
<mods:identifier id="D7CEF305C9C1951A932344ABABB2B0B5" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="54238D0177F2266E65C823B60C44B212" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="2CE0692ABDEEB42FAFD8A9C83EF1C5B7" type="ISSN">1175­5326</mods:identifier>
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<treatment id="03A287CA7821FFBAFEECFB369DF7FC71" ID-DOI="http://doi.org/10.5281/zenodo.6278023" ID-GBIF-Taxon="121065209" ID-Zenodo-Dep="6278023" LSID="urn:lsid:plazi:treatment:03A287CA7821FFBAFEECFB369DF7FC71" httpUri="http://treatment.plazi.org/id/03A287CA7821FFBAFEECFB369DF7FC71" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection id="C31165577821FFBBFEECFB369E01FB09" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BB436DC7821FFBBFEECFB369EB2FB32" blockId="52.[264,646,1203,1270]" box="[264,646,1203,1230]" pageId="52" pageNumber="53">
<heading id="D0FC81B07821FFBBFEECFB369EB2FB32" bold="true" box="[264,646,1203,1230]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<emphasis id="B97FEACE7821FFBBFEECFB369EB2FB32" bold="true" box="[264,646,1203,1230]" pageId="52" pageNumber="53">
<taxonomicName id="4C0B4D5F7821FFBBFEECFB369DDDFB32" ID-CoL="664LW" authority="Erwin" authorityName="Erwin" box="[264,489,1203,1230]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pitilla" status="sp. nov.">
<emphasis id="B97FEACE7821FFBBFEECFB369DA6FB31" bold="true" box="[264,402,1204,1230]" italics="true" pageId="52" pageNumber="53">Agra pitilla</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B57821FFBBFE12FB319EB2FB32" box="[502,646,1203,1229]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC7821FFBBFEECFB5E9E01FB09" blockId="52.[264,646,1203,1270]" box="[264,565,1244,1270]" pageId="52" pageNumber="53">
(
<figureCitation id="13302A597821FFBBFEF5FB5E9D9FFB09" box="[273,427,1244,1270]" captionStart="FIGURE 24. A" captionStartId="53.[264,368,1780,1804]" captionTargetBox="[420,1176,994,1728]" captionTargetId="figure@53.[402,1185,969,1754]" captionTargetPageId="53" captionText="FIGURE 24. A. pitilla, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.3; WE = 2.9; male SW = 4.2, female SW = 4.4; AL = 3.5." httpUri="https://zenodo.org/record/162838/files/figure.png" pageId="52" pageNumber="53">Figs. 24a, 24</figureCitation>
b, 24c, 24d)
</paragraph>
</subSubSection>
<subSubSection id="C31165577821FFBBFEECFAA99E5AFA69" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8BB436DC7821FFBBFEECFAA99E5AFA69" blockId="52.[264,1324,1323,1870]" pageId="52" pageNumber="53">
<emphasis id="B97FEACE7821FFBBFEECFAA99DB5FABA" bold="true" box="[264,385,1323,1349]" pageId="52" pageNumber="53">
<typeStatus id="54B0887E7821FFBBFEECFAA99D48FABA" box="[264,380,1323,1349]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<collectingCountry id="F31C764C7821FFBBFE32FAAE9EBCFAB9" box="[470,648,1324,1350]" name="Costa Rica" pageId="52" pageNumber="53">COSTA RICA</collectingCountry>
: Guanacaste, P.N. Guanacaste, Estacíon
<taxonomicName id="4C0B4D5F7821FFBBFB8DFAAE9886FAB9" box="[1129,1202,1324,1350]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, 9.0 km S Santa Cecilia,
<quantity id="4CF39B397821FFBBFE2AFAD69E20FA91" box="[462,532,1364,1390]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="52" pageNumber="53" unit="m" value="700.0">700m</quantity>
,
<geoCoordinate id="EE3F501B7821FFBBFDCAFAD69E8FFA91" box="[558,699,1364,1390]" direction="north" orientation="latitude" pageId="52" pageNumber="53" precision="15" value="10.990556">10°59'26”N</geoCoordinate>
,
<geoCoordinate id="EE3F501B7821FFBBFD31FAD69F53FA91" box="[725,871,1364,1390]" direction="west" orientation="longitude" pageId="52" pageNumber="53" precision="15" value="-85.42777">85°25'40”W</geoCoordinate>
, January, (GNP Biodiversity Survey)(
<taxonomicName id="4C0B4D5F7821FFBBFEA1FAFE9DA7FA69" box="[325,403,1404,1430]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197821FFBBFEA1FAFE9DA7FA69" box="[325,403,1404,1430]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="52" pageNumber="53">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197821FFBBFE7AFAFE9DE5FA69" LSID="urn:lsid:biocol.org:col:14323" box="[414,465,1404,1430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="52" pageNumber="53">CRI</collectionCode>
001­024216).
</paragraph>
</subSubSection>
<subSubSection id="C31165577821FFBBFEDCFA219821FA19" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BB436DC7821FFBBFEDCFA219821FA19" blockId="52.[264,1324,1323,1870]" pageId="52" pageNumber="53">
<emphasis id="B97FEACE7821FFBBFEDCFA219D83FA42" bold="true" box="[312,439,1443,1469]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
Black fore body with rufinistic elytral apex, the apical margin infuscated and markedly lobed at middle.
<collectingCountry id="F31C764C7821FFBBFD90FA4E9E9BFA19" box="[628,687,1484,1510]" name="India" pageId="52" pageNumber="53">Hind</collectingCountry>
trochanter sparsely setiferous.
</paragraph>
</subSubSection>
<subSubSection id="C31165577821FFBAFEDCFA719DDCFEA1" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="8BB436DC7821FFBAFEDCFA719DDCFEA1" blockId="52.[264,1324,1323,1870]" lastBlockId="53.[264,1326,284,910]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="B97FEACE7821FFBBFEDCFA719DE4F9F2" bold="true" box="[312,464,1523,1549]" pageId="52" pageNumber="53">Description.</emphasis>
Color and luster: Black. Apical 2/3 of elytra rufinistic, margin of elytral apex, abdominal sternum VI, and tergum VI infuscated; tarsi and abdomen rufotestaceous. Form: Labrum flat. Head (
<figureCitation id="13302A597821FFBBFDA3F9C69E98F9A1" box="[583,684,1604,1630]" captionStart="FIGURE 24. A" captionStartId="53.[264,368,1780,1804]" captionTargetBox="[420,1176,994,1728]" captionTargetId="figure@53.[402,1185,969,1754]" captionTargetPageId="53" captionText="FIGURE 24. A. pitilla, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.3; WE = 2.9; male SW = 4.2, female SW = 4.4; AL = 3.5." httpUri="https://zenodo.org/record/162838/files/figure.png" pageId="52" pageNumber="53">Fig. 24a</figureCitation>
) markedly short behind eyes, quadrate in both sexes. Elytra with markedly and evenly punctulate interneurs, apex (
<figureCitation id="13302A597821FFBBFBE3F9EE9855F979" box="[1031,1121,1644,1670]" captionStart="FIGURE 24. A" captionStartId="53.[264,368,1780,1804]" captionTargetBox="[420,1176,994,1728]" captionTargetId="figure@53.[402,1185,969,1754]" captionTargetPageId="53" captionText="FIGURE 24. A. pitilla, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.3; WE = 2.9; male SW = 4.2, female SW = 4.4; AL = 3.5." httpUri="https://zenodo.org/record/162838/files/figure.png" pageId="52" pageNumber="53">Fig. 24</figureCitation>
b) slightly lobed between lateral and sutural angles. Metasternum of male densely pubescent; abdominal sterna each with long scattered setae. Apex of sternum VI (
<figureCitation id="13302A597821FFBBFC34F93E9813F929" box="[976,1063,1724,1750]" captionStart="FIGURE 24. A" captionStartId="53.[264,368,1780,1804]" captionTargetBox="[420,1176,994,1728]" captionTargetId="figure@53.[402,1185,969,1754]" captionTargetPageId="53" captionText="FIGURE 24. A. pitilla, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.3; WE = 2.9; male SW = 4.2, female SW = 4.4; AL = 3.5." httpUri="https://zenodo.org/record/162838/files/figure.png" pageId="52" pageNumber="53">Fig. 24</figureCitation>
c) in male and female acutely angulate apicolaterally, caudal margin emarginate, slightly u­notched in male, vnotched in female. Male middle tibia with small apicomedial brush. Aedeagus (
<figureCitation id="13302A597821FFBBFB59F88E9920F8D9" box="[1213,1300,1804,1830]" captionStart="FIGURE 24. A" captionStartId="53.[264,368,1780,1804]" captionTargetBox="[420,1176,994,1728]" captionTargetId="figure@53.[402,1185,969,1754]" captionTargetPageId="53" captionText="FIGURE 24. A. pitilla, male, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.3; WE = 2.9; male SW = 4.2, female SW = 4.4; AL = 3.5." httpUri="https://zenodo.org/record/162838/files/figure.png" pageId="52" pageNumber="53">Fig. 24</figureCitation>
d) ostium short, slightly left pleuropic; apex of phallus a simple diamond­shaped lobe; left paramere unisetose, right paramere glabrous. Size: medium, 13.0 to
<quantity id="4CF39B397820FFBAFBC7FE9E98BFFEC9" box="[1059,1163,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="53" pageNumber="54" unit="mm" value="13.5">13.5 mm</quantity>
in length, 3.1 o
<quantity id="4CF39B397820FFBAFEFAFEC69D4CFEA1" box="[286,376,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="53" pageNumber="54" unit="mm" value="3.7">3.7 mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C31165577820FFBAFEDCFEE9980EFCC1" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BB436DC7820FFBAFEDCFEE99DB4FD11" blockId="53.[264,1326,284,910]" pageId="53" pageNumber="54">
<emphasis id="B97FEACE7820FFBAFEDCFEE99EA2FE7A" bold="true" box="[312,662,363,389]" pageId="53" pageNumber="54">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C7820FFBAFD44FEEE9F13FE79" box="[672,807,364,390]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
: 7
<typeStatus id="54B0887E7820FFBAFCAAFEEE9FF5FE79" box="[846,961,364,390]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D0DFD557820FFBAFC34FEEE9804FE79" box="[976,1072,364,390]" pageId="53" pageNumber="54" type="male">5 males</specimenCount>
,
<specimenCount id="9D0DFD557820FFBAFBDBFEEE9881FE79" box="[1087,1205,364,390]" pageId="53" pageNumber="54" type="female">2 females</specimenCount>
, from the Guanacaste Conservation Area.
<collectingCountry id="F31C764C7820FFBAFD61FE169F3DFE51" box="[645,777,404,430]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
, Alajuela, 1f, Dos Rios, 2.0 km W, N slope of Volcán de Rincon,
<quantity id="4CF39B397820FFBAFE0AFE3E9E00FE29" box="[494,564,444,470]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="53" pageNumber="54" unit="m" value="550.0">550m</quantity>
,
<geoCoordinate id="EE3F501B7820FFBAFDA7FE3E9E8EFE29" box="[579,698,444,470]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="10.833333">10° 50' N</geoCoordinate>
,
<geoCoordinate id="EE3F501B7820FFBAFD2DFE3E9F67FE29" box="[713,851,444,470]" direction="west" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="-85.36667">085° 22' W</geoCoordinate>
, May (J.T. Doyen &amp; P.A. Opler)(
<collectionCode id="ED1AAE197820FFBAFB0FFE3E991EFE29" box="[1259,1322,444,470]" httpUri="http://grbio.org/cool/wq7v-amgw" name="University of California at Berkeley" pageId="53" pageNumber="54">UCB</collectionCode>
: ADP80686); Guanacaste, 1f, Santa Cecilia, 9.0 km S,
<quantity id="4CF39B397820FFBAFC68FE669FE6FE01" box="[908,978,484,510]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="53" pageNumber="54" unit="m" value="700.0">700m</quantity>
, 10° 59' 26 N, 85° 25' 40' W, LN 330200,380200, June (C. Moraga)(
<taxonomicName id="4C0B4D5F7820FFBAFD33FD8E9F17FDD9" box="[727,803,524,550]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFD33FD8E9F17FDD9" box="[727,803,524,550]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFCCAFD8E9F54FDD9" LSID="urn:lsid:biocol.org:col:14323" box="[814,864,524,550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
002047455),
<specimenCount id="9D0DFD557820FFBAFC18FD8E986EFDD9" box="[1020,1114,524,550]" pageId="53" pageNumber="54" type="male">6 males</specimenCount>
, P.N. Guanacaste, Estacíon
<taxonomicName id="4C0B4D5F7820FFBAFE91FDB69D8AFDB1" box="[373,446,564,590]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, April (P. Rios)(
<taxonomicName id="4C0B4D5F7820FFBAFD65FDB69EFBFDB1" box="[641,719,564,590]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFD65FDB69EFBFDB1" box="[641,719,564,590]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFD3FFDB69F39FDB1" LSID="urn:lsid:biocol.org:col:14323" box="[731,781,564,590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
001794228), March (P. Rios, C. Moraga &amp; R. Blanco)(
<taxonomicName id="4C0B4D5F7820FFBAFE8BFDDE9D88FD89" box="[367,444,604,630]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFE8BFDDE9D88FD89" box="[367,444,604,630]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFE2EFDDE9DC8FD89" LSID="urn:lsid:biocol.org:col:14323" box="[458,508,604,630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
000­211428), December (C. Moraga)(
<taxonomicName id="4C0B4D5F7820FFBAFC27FDDE983BFD89" box="[963,1039,604,630]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFC27FDDE983BFD89" box="[963,1039,604,630]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFBF9FDDE9864FD89" LSID="urn:lsid:biocol.org:col:14323" box="[1053,1104,604,630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
000­637527), May (P. Rios &amp; M.
<taxonomicName id="4C0B4D5F7820FFBAFE28FD069E0AFD61" box="[460,574,644,670]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="64" phylum="Arthropoda" rank="species" species="zumbado">Zumbado</taxonomicName>
) (
<collectionCode id="ED1AAE197820FFBAFDB8FD069EBBFD61" LSID="urn:lsid:biocol.org:col:14323" box="[604,655,644,670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
000­269888), LN 330200, 380200 #3262, October (P. Rios)(
<taxonomicName id="4C0B4D5F7820FFBAFEB5FD2E9DABFD39" box="[337,415,684,710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFEB5FD2E9DABFD39" box="[337,415,684,710]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFE53FD2E9DDEFD39" LSID="urn:lsid:biocol.org:col:14323" box="[439,490,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
001­997986), January (GNP Biodiversity Survey)(
<taxonomicName id="4C0B4D5F7820FFBAFB8BFD2E9888FD39" box="[1135,1212,684,710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFB8BFD2E9888FD39" box="[1135,1212,684,710]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE197820FFBAFB37FD2E9932FD39" LSID="urn:lsid:biocol.org:col:14323" box="[1235,1286,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="53" pageNumber="54">CRI</collectionCode>
00­ 1024216).
</paragraph>
<paragraph id="8BB436DC7820FFBAFEDCFD79980EFCC1" blockId="53.[264,1326,284,910]" pageId="53" pageNumber="54">
<emphasis id="B97FEACE7820FFBAFEDCFD799E37FCEA" bold="true" box="[312,515,763,789]" pageId="53" pageNumber="54">Specific epithet.</emphasis>
The specific epithet,
<emphasis id="B97FEACE7820FFBAFCF5FD7E9F69FCEA" box="[785,861,764,789]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C0B4D5F7820FFBAFCF5FD7E9F6EFCEA" box="[785,858,764,789]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">pitilla</taxonomicName>
,
</emphasis>
is a noun in apposition based on the name of the
<taxonomicName id="4C0B4D5F7820FFBAFE7EFCA69DD7FCC1" box="[410,483,804,830]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE197820FFBAFE7EFCA69DD7FCC1" box="[410,483,804,830]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="53" pageNumber="54">INBio</collectionCode>
</taxonomicName>
Field Station at which the
<typeStatus id="54B0887E7820FFBAFCFBFCA69FB8FCC1" box="[799,908,804,830]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C31165577820FFBAFEDCFCC99DF7FC71" pageId="53" pageNumber="54" type="description">
<paragraph id="8BB436DC7820FFBAFEDCFCC99DF7FC71" blockId="53.[264,1326,284,910]" pageId="53" pageNumber="54">
<emphasis id="B97FEACE7820FFBAFEDCFCC99DB1FC9A" bold="true" box="[312,389,843,869]" pageId="53" pageNumber="54">Notes.</emphasis>
All specimens are from the Guanacaste Conservation Area, 7 of them from Estacíon
<taxonomicName id="4C0B4D5F7820FFBAFE90FCF69D8AFC71" box="[372,446,884,910]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>