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<document id="788771B55FC23C2ADB0C50118658A578" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA781AFF9FFEECFB549FBAFEA1" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra jimwappes Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="17" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="16" updateTime="1698190273211" updateUser="plazi">
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<mods:titleInfo id="9EFE7B6557E50A2AC98C2D64B88CBA1E">
<mods:title id="0F905920C1A22B821C2840E832020B0E">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="D44F25B6B8ADB742400CE5E3FB6DE7A1" type="personal">
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<mods:namePart id="7C7C4D965D050E3F6F23DE375ABAAEF8">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="92EBEBA359A540E7F3FC7E113B90DF76">text</mods:typeOfResource>
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<mods:title id="1553BC6265DB9EAD2B725050CE96E40E">Zootaxa</mods:title>
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<mods:part id="BF1386C4250D40593513C6C8A17BE91E">
<mods:date id="50E739FDA22602CEAC5210E51AC2081E">2002</mods:date>
<mods:detail id="2C18D07B853A0ACF690D1E8961D70F81" type="volume">
<mods:number id="89760DC25AC1A71EA4A6B703D0220135">119</mods:number>
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<mods:classification id="F6BEB856599B1235A0E9487115AF4B55">journal article</mods:classification>
<mods:identifier id="9DFCAB0361D8A4C1B8BA5A6FF5F7DB32" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="D5C3BBFC6143CF5B6089A8C215A534B0" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="E45B56A003DF9F2BDEC8A7835F8A8743" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="489A1B2E253710514FB97A433F96CEDD" type="Zenodo-Dep">162803</mods:identifier>
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<treatment id="03A287CA781AFF9FFEECFB549FBAFEA1" ID-DOI="http://doi.org/10.5281/zenodo.6277966" ID-GBIF-Taxon="119330638" ID-Zenodo-Dep="6277966" LSID="urn:lsid:plazi:treatment:03A287CA781AFF9FFEECFB549FBAFEA1" httpUri="http://treatment.plazi.org/id/03A287CA781AFF9FFEECFB549FBAFEA1" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C3116557781AFF80FEECFB549DFBFAE6" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BB436DC781AFF80FEECFB549E8BFB0F" blockId="15.[264,703,1238,1305]" box="[264,703,1238,1264]" pageId="15" pageNumber="16">
<heading id="D0FC81B0781AFF80FEECFB549E8BFB0F" bold="true" box="[264,703,1238,1264]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B97FEACE781AFF80FEECFB549E8BFB0F" bold="true" box="[264,703,1238,1264]" pageId="15" pageNumber="16">
<taxonomicName id="4C0B4D5F781AFF80FEECFB549E16FB0F" ID-CoL="B2PQ" authority="Erwin" authorityName="Erwin" box="[264,546,1238,1264]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jimwappes" status="sp. nov.">
<emphasis id="B97FEACE781AFF80FEECFB549DFFFB0F" bold="true" box="[264,459,1238,1264]" italics="true" pageId="15" pageNumber="16">Agra jimwappes</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B5781AFF80FDCBFB549E8BFB0F" box="[559,703,1238,1264]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC781AFF80FEECFB7D9DFBFAE6" blockId="15.[264,703,1238,1305]" box="[264,463,1279,1305]" pageId="15" pageNumber="16">
(
<figureCitation id="13302A59781AFF80FEF5FB7D9DBEFAE6" box="[273,394,1279,1305]" captionStart="FIGURE 1. A" captionStartId="16.[264,367,823,847]" captionTargetBox="[428,1157,464,786]" captionTargetId="figure@16.[423,1164,449,800]" captionTargetPageId="16" captionText="FIGURE 1. A. jimwappesi, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect. Measures (mm): HL = 2.6; WE = 1.9; SW = 3.6." httpUri="https://zenodo.org/record/162804/files/figure.png" pageId="15" pageNumber="16">Figs. 1a, 1</figureCitation>
b, 1c)
</paragraph>
</subSubSection>
<subSubSection id="C3116557781AFF80FEECFACC9E45FA6E" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BB436DC781AFF80FEECFACC9E45FA6E" blockId="15.[264,1324,1358,1904]" pageId="15" pageNumber="16">
<emphasis id="B97FEACE781AFF80FEECFACC9DB5FA97" bold="true" box="[264,385,1358,1384]" pageId="15" pageNumber="16">
<typeStatus id="54B0887E781AFF80FEECFACC9D48FA97" box="[264,380,1358,1384]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female,
<collectingCountry id="F31C764C781AFF80FE0AFACD9EAAFA96" box="[494,670,1359,1385]" name="Costa Rica" pageId="15" pageNumber="16">COSTA RICA</collectingCountry>
: Guanacaste, La Pacifica,
<geoCoordinate id="EE3F501B781AFF80FC37FACD9805FA96" box="[979,1073,1359,1385]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="10.466666">10º28'N</geoCoordinate>
<geoCoordinate id="EE3F501B781AFF80FBDCFACD989FFA96" box="[1080,1195,1359,1385]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="-85.11667">085º07'W</geoCoordinate>
, June (J.E. Wappes)(
<collectionCode id="ED1AAE19781AFF80FE9DFAF59DE4FA6E" LSID="urn:lsid:biocol.org:col:34871" box="[377,464,1399,1425]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
: ADP85171).
</paragraph>
</subSubSection>
<subSubSection id="C3116557781AFF80FEDCFA1C9F3BFA1E" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BB436DC781AFF80FEDCFA1C9F3BFA1E" blockId="15.[264,1324,1358,1904]" pageId="15" pageNumber="16">
<emphasis id="B97FEACE781AFF80FEDCFA1C9D83FA47" bold="true" box="[312,439,1438,1464]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
With character states of the arrowi group, and elytral apex clearly obliquely truncate with a small lateral tooth.
</paragraph>
</subSubSection>
<subSubSection id="C3116557781AFF80FEDCFA6C9F2DF92F" pageId="15" pageNumber="16" type="description">
<paragraph id="8BB436DC781AFF80FEDCFA6C9F2DF92F" blockId="15.[264,1324,1358,1904]" pageId="15" pageNumber="16">
<emphasis id="B97FEACE781AFF80FEDCFA6C9DE4F9F7" bold="true" box="[312,464,1518,1544]" pageId="15" pageNumber="16">Description.</emphasis>
Color and luster: Castaneous, head and knees piceous; head and prontoum very shiny, elytra somewhat matte in appearance. Form: Male unknown. Head (
<figureCitation id="13302A59781AFF80FB18F9959D17F9A6" captionStart="FIGURE 1. A" captionStartId="16.[264,367,823,847]" captionTargetBox="[428,1157,464,786]" captionTargetId="figure@16.[423,1164,449,800]" captionTargetPageId="16" captionText="FIGURE 1. A. jimwappesi, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect. Measures (mm): HL = 2.6; WE = 1.9; SW = 3.6." httpUri="https://zenodo.org/record/162804/files/figure.png" pageId="15" pageNumber="16">Fig. 1a</figureCitation>
) behind eyes somewhat rounded in female. Elytron (
<figureCitation id="13302A59781AFF80FC74F9BD9FE6F9A6" box="[912,978,1599,1625]" captionStart="FIGURE 1. A" captionStartId="16.[264,367,823,847]" captionTargetBox="[428,1157,464,786]" captionTargetId="figure@16.[423,1164,449,800]" captionTargetPageId="16" captionText="FIGURE 1. A. jimwappesi, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect. Measures (mm): HL = 2.6; WE = 1.9; SW = 3.6." httpUri="https://zenodo.org/record/162804/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
b) with apex moderately produced medially, lateral tooth small, sutural angle obtuse,
<taxonomicName id="4C0B4D5F781AFF80FC49F9E59FE6F97E" box="[941,978,1639,1665]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="not">not</taxonomicName>
projected acutely; interneurs of biserite rows of punctulae. Sternum VI (
<figureCitation id="13302A59781AFF80FCF0F90C9F6CF957" box="[788,856,1678,1704]" captionStart="FIGURE 1. A" captionStartId="16.[264,367,823,847]" captionTargetBox="[428,1157,464,786]" captionTargetId="figure@16.[423,1164,449,800]" captionTargetPageId="16" captionText="FIGURE 1. A. jimwappesi, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect. Measures (mm): HL = 2.6; WE = 1.9; SW = 3.6." httpUri="https://zenodo.org/record/162804/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
c) of female shallowly v­shaped. Size: medium,
<quantity id="4CF39B39781AFF80FE93F9349DECF92F" box="[375,472,1718,1744]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="15" pageNumber="16" unit="mm" value="15.1">15.1mm</quantity>
in length,
<quantity id="4CF39B39781AFF80FDB2F9349E9CF92F" box="[598,680,1718,1744]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="16" unit="mm" value="3.8">3.8mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C3116557781AFF80FEDCF95C9D91F88F" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BB436DC781AFF80FEDCF95C9916F907" blockId="15.[264,1324,1358,1904]" box="[312,1314,1758,1784]" pageId="15" pageNumber="16">
<emphasis id="B97FEACE781AFF80FEDCF95C9EA4F907" bold="true" box="[312,656,1758,1784]" pageId="15" pageNumber="16">Other specimens examined.</emphasis>
None, the
<typeStatus id="54B0887E781AFF80FCF5F95C9F4AF907" box="[785,894,1758,1784]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
is only specimen known at present.
</paragraph>
<paragraph id="8BB436DC781AFF80FEDCF8849D91F88F" blockId="15.[264,1324,1358,1904]" pageId="15" pageNumber="16">
<emphasis id="B97FEACE781AFF80FEDCF8849DCAF8DF" bold="true" box="[312,510,1798,1824]" pageId="15" pageNumber="16">Specific epithet.</emphasis>
The specific epithet,
<taxonomicName id="4C0B4D5F781AFF80FD1FF8859F4CF8DF" box="[763,888,1799,1824]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jimwappes">
<emphasis id="B97FEACE781AFF80FD1FF8859F4CF8DF" box="[763,888,1799,1824]" italics="true" pageId="15" pageNumber="16">jimwappes</emphasis>
</taxonomicName>
, is a noun in apposition based on the name of James E. Wappes, the collector of the
<typeStatus id="54B0887E781AFF80FCCEF8AC9FAEF8B7" box="[810,922,1838,1864]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
, and many other excellent records of this genus.
</paragraph>
</subSubSection>
<subSubSection id="C31165577805FF9FFEDCFE999FBAFEA1" pageId="16" pageNumber="17" type="description">
<paragraph id="8BB436DC7805FF9FFEDCFE999FBAFEA1" blockId="16.[264,1323,283,350]" pageId="16" pageNumber="17">
<emphasis id="B97FEACE7805FF9FFEDCFE999DB1FECA" bold="true" box="[312,389,283,309]" pageId="16" pageNumber="17">Notes.</emphasis>
Adults of this species are similar to adults of
<taxonomicName id="4C0B4D5F7805FF9FFC4BFE9E9874FECA" box="[943,1088,284,309]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="winnie">
<emphasis id="B97FEACE7805FF9FFC4BFE9E9874FECA" box="[943,1088,284,309]" italics="true" pageId="16" pageNumber="17">Agra winnie</emphasis>
</taxonomicName>
, however, the truncated elytral apex of
<taxonomicName id="4C0B4D5F7805FF9FFE1AFEC69E8AFEA2" box="[510,702,324,349]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jimwappes">
<emphasis id="B97FEACE7805FF9FFE1AFEC69E8AFEA2" box="[510,702,324,349]" italics="true" pageId="16" pageNumber="17">Agra jimwappes</emphasis>
</taxonomicName>
is clearly unique.
</paragraph>
</subSubSection>
</treatment>
</document>