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<document id="B56738B9EC6E38D14C5B6AE86C826EC3" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA780EFF93FEECFDC49D4BFC49" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra fugax Erwin, new species" docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="28" updateTime="1698190273211" updateUser="plazi">
<mods:mods id="C1AA214CAC458052EF44477E836F2C95" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D1A3E0D7E89778FA4854A90FF1351425">
<mods:title id="F149874BF3DE1655C5F3EA34E936A6B0">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="0DFA2BD7C9A0575D94AABBA8FDC03C4D" type="personal">
<mods:role id="C4CF3CF879BA6C45287384162DBB3737">
<mods:roleTerm id="9F6F64ECB55182A89EF15F54A9726AE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B574279472272092F83DE127A3CE8389">Erwin, Terry L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="91C12F1BC438E0FA61495410247D8899">text</mods:typeOfResource>
<mods:relatedItem id="0C64D4FC60CA5690701408D716E16B8D" type="host">
<mods:titleInfo id="82D2D527B953D4033F206FB8C0C2F1EC">
<mods:title id="79AC41663FEADB25CAF0207263123043">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3C57BB4E3227FE81D181AA9D065339C0">
<mods:date id="7FFEB3523A9D135BB8F8040D5B609E74">2002</mods:date>
<mods:detail id="68EBEBC15453AC26B000AF1728A7D07B" type="volume">
<mods:number id="923FC6CD9EF5F5DC4F973F04F31E8E90">119</mods:number>
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<mods:start id="FFF600EDED1F904661A34CE9DF944AA6">1</mods:start>
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<mods:classification id="0D858496286FD9716C924C0A26D2AB85">journal article</mods:classification>
<mods:identifier id="234E11890DF6D007192C2B502EAFA8D1" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="D53DC187CF6CD18B87CD8A6E50BEFE00" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
<mods:identifier id="169C8BB6BF92408EF29F24103C0D1F7A" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="CE4036A01FC33127030C965D2F540156" type="Zenodo-Dep">162803</mods:identifier>
</mods:mods>
<treatment id="03A287CA780EFF93FEECFDC49D4BFC49" ID-DOI="http://doi.org/10.5281/zenodo.6277980" ID-GBIF-Taxon="119330640" ID-Zenodo-Dep="6277980" LSID="urn:lsid:plazi:treatment:03A287CA780EFF93FEECFDC49D4BFC49" httpUri="http://treatment.plazi.org/id/03A287CA780EFF93FEECFDC49D4BFC49" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C3116557780EFF94FEECFDC49DCFFD76" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BB436DC780EFF94FEECFDC49EB2FD9F" blockId="27.[264,646,582,649]" box="[264,646,582,608]" pageId="27" pageNumber="28">
<heading id="D0FC81B0780EFF94FEECFDC49EB2FD9F" bold="true" box="[264,646,582,608]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B97FEACE780EFF94FEECFDC49EB2FD9F" bold="true" box="[264,646,582,608]" pageId="27" pageNumber="28">
<taxonomicName id="4C0B4D5F780EFF94FEECFDC49DDDFD9F" ID-CoL="B2MX" authority="Erwin" authorityName="Erwin" box="[264,489,582,608]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fugax" status="sp. nov.">
<emphasis id="B97FEACE780EFF94FEECFDC49DA6FD9F" bold="true" box="[264,402,582,608]" italics="true" pageId="27" pageNumber="28">Agra fugax</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B5780EFF94FE12FDC49EB2FD9F" box="[502,646,582,608]" pageId="27" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC780EFF94FEECFDED9DCFFD76" blockId="27.[264,646,582,649]" box="[264,507,623,649]" pageId="27" pageNumber="28">
(
<figureCitation id="13302A59780EFF94FEF5FDED9DB8FD76" box="[273,396,623,649]" captionStart="FIGURE 8. A" captionStartId="27.[264,368,1809,1833]" captionTargetBox="[437,1145,810,1795]" captionTargetId="figure@27.[434,1152,802,1798]" captionTargetPageId="27" captionText="FIGURE 8. A. fugax, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects .. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4." httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="27" pageNumber="28">Figs. 8a, 8</figureCitation>
b, 8c, 8d)
</paragraph>
</subSubSection>
<subSubSection id="C3116557780EFF94FEECFD31992DFD09" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BB436DC780EFF94FEECFD31992DFD09" blockId="27.[264,1324,691,758]" pageId="27" pageNumber="28">
<emphasis id="B97FEACE780EFF94FEECFD319DB5FD32" bold="true" box="[264,385,691,717]" pageId="27" pageNumber="28">
<typeStatus id="54B0887E780EFF94FEECFD319D48FD32" box="[264,380,691,717]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male.
<collectingCountry id="F31C764C780EFF94FE3FFD369EBBFD31" box="[475,655,692,718]" name="Costa Rica" pageId="27" pageNumber="28">COSTA RICA</collectingCountry>
: Puntarenas, Carara Biological Reserve, Estacíon Quebrada Bonita,
<quantity id="4CF39B39780EFF94FE4AFD5E9DD0FD09" box="[430,484,732,758]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="28" unit="m" value="50.0">50m</quantity>
,
<geoCoordinate id="EE3F501B780EFF94FE15FD5E9E48FD09" box="[497,636,732,758]" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="9.776388">09°46'35”N</geoCoordinate>
<geoCoordinate id="EE3F501B780EFF94FD67FD5E9F20FD09" box="[643,788,732,758]" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="15" value="-84.60305">84°36'11”W</geoCoordinate>
, June (R.
<taxonomicName id="4C0B4D5F780EFF94FC6DFD5E9FD5FD09" box="[905,993,732,758]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="66" phylum="Arthropoda" rank="species" species="zuniga">Zuñiga</taxonomicName>
)(
<taxonomicName id="4C0B4D5F780EFF94FC15FD5E980AFD09" box="[1009,1086,732,758]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780EFF94FC15FD5E980AFD09" box="[1009,1086,732,758]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="27" pageNumber="28">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19780EFF94FBADFD5E9848FD09" LSID="urn:lsid:biocol.org:col:14323" box="[1097,1148,732,758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="27" pageNumber="28">CRI</collectionCode>
000­349227).
</paragraph>
</subSubSection>
<caption id="DF746654780EFF94FEECF893984DF895" httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="27" pageNumber="28" targetBox="[437,1145,810,1795]" targetPageId="27">
<paragraph id="8BB436DC780EFF94FEECF893984DF895" blockId="27.[264,1325,1809,1898]" pageId="27" pageNumber="28">
<emphasis id="B97FEACE780EFF94FEECF8939DBAF8D6" bold="true" box="[264,398,1809,1833]" pageId="27" pageNumber="28">FIGURE 8.</emphasis>
<taxonomicName id="4C0B4D5F780EFF94FE72F8939DC5F8D7" box="[406,497,1809,1832]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fugax">
<emphasis id="B97FEACE780EFF94FE72F8939DC5F8D7" box="[406,497,1809,1832]" italics="true" pageId="27" pageNumber="28">A. fugax</emphasis>
</taxonomicName>
, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects.. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4.
</paragraph>
</caption>
<subSubSection id="C31165577809FF93FEDCFE99980FFE79" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BB436DC7809FF93FEDCFE99980FFE79" blockId="28.[264,1324,283,950]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDCFE999D8CFECA" bold="true" box="[312,440,283,309]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
Femur concolorous; elytron with sutural and lateral apices modestly dentate, margin between them slightly arcuate. Head markedly convex and nearly quadrate basally. Elytral interneurs each a series of large shiny (brassy) foveae.
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF93FEDCFE1198DAFD61" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB436DC7809FF93FEDCFE1198DAFD61" blockId="28.[264,1324,283,950]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDCFE119DE4FE52" bold="true" box="[312,464,403,429]" pageId="28" pageNumber="29">Description.</emphasis>
Color and luster: Piceous with testaceous appendages, femur concolorous; elytral interneurs each a series of large shiny (brassy) foveae. Form: Head (
<figureCitation id="13302A597809FF93FB28FE3E9915FE29" box="[1228,1313,444,470]" captionStart="FIGURE 8. A" captionStartId="27.[264,368,1809,1833]" captionTargetBox="[437,1145,810,1795]" captionTargetId="figure@27.[434,1152,802,1798]" captionTargetPageId="27" captionText="FIGURE 8. A. fugax, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects .. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4." httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="28" pageNumber="29">Fig. 8a</figureCitation>
) behind eyes markedly convex and nearly quadrate posteriorly. Elytron (
<figureCitation id="13302A597809FF93FBB6FE6698A3FE01" box="[1106,1175,484,510]" captionStart="FIGURE 8. A" captionStartId="27.[264,368,1809,1833]" captionTargetBox="[437,1145,810,1795]" captionTargetId="figure@27.[434,1152,802,1798]" captionTargetPageId="27" captionText="FIGURE 8. A. fugax, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects .. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4." httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
b) with apex moderately oblique, lateral and sutural dentiform projections moderately large, acute; interneurs each a series of large shiny (brassy) foveae. Abdominal sternum VI (
<figureCitation id="13302A597809FF93FB28FDB69920FDB1" box="[1228,1300,564,590]" captionStart="FIGURE 8. A" captionStartId="27.[264,368,1809,1833]" captionTargetBox="[437,1145,810,1795]" captionTargetId="figure@27.[434,1152,802,1798]" captionTargetPageId="27" captionText="FIGURE 8. A. fugax, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects .. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4." httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
c) shallowly U­notched in female. Aedeagus (
<figureCitation id="13302A597809FF93FCF0FDDE9F68FD89" box="[788,860,604,630]" captionStart="FIGURE 8. A" captionStartId="27.[264,368,1809,1833]" captionTargetBox="[437,1145,810,1795]" captionTargetId="figure@27.[434,1152,802,1798]" captionTargetPageId="27" captionText="FIGURE 8. A. fugax, female, dorsal aspect: a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) aedeagus, dorsal (middle), ventral (left), left lateral (right) aspects .. Measures (mm): HL = 2.4; PL = 3.0; WE = 1.6; male SW = 3.0, female SW = 3.6; AL = 2.4." httpUri="https://zenodo.org/record/162812/files/figure.png" pageId="28" pageNumber="29">Fig. 8</figureCitation>
d) with ostium moderately short, apex simple, cordiform. Size: medium,
<quantity id="4CF39B397809FF93FD7EFD069F68FD61" box="[666,860,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.3800000000000001" metricValueMin="1.3599999999999999" pageId="28" pageNumber="29" unit="mm" value="13.7" valueMax="13.8" valueMin="13.6">13.6 to 13.8 mm</quantity>
in length,
<quantity id="4CF39B397809FF93FC3DFD06984AFD61" box="[985,1150,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="28" pageNumber="29" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2 to 3.8 mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C31165577809FF93FEDBFD299D4BFC49" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB436DC7809FF93FEDBFD299DBBFCE9" blockId="28.[264,1324,283,950]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDBFD299EA1FD3A" bold="true" box="[319,661,683,709]" pageId="28" pageNumber="29">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C7809FF93FD78FD2E9F14FD39" box="[668,800,684,710]" name="Costa Rica" pageId="28" pageNumber="29">Costa Rica</collectingCountry>
: 1
<typeStatus id="54B0887E7809FF93FCA4FD2E9F9CFD39" box="[832,936,684,710]" pageId="28" pageNumber="29" type="paratype">paratype</typeStatus>
, female, from the Cordillera Volcánica Central Conservation Area: 1f, Cartago,
<taxonomicName id="4C0B4D5F7809FF93FC95FD569FD4FD11" box="[881,992,724,750]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
,
<geoCoordinate id="EE3F501B7809FF93FC11FD569868FD11" box="[1013,1116,724,750]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B7809FF93FB89FD5698F7FD11" box="[1133,1219,724,750]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="9.883333">9°53'N</geoCoordinate>
, (WAR: ADP4214).
</paragraph>
<paragraph id="8BB436DC7809FF93FEDCFCA19D4BFC49" blockId="28.[264,1324,283,950]" pageId="28" pageNumber="29">
<emphasis id="B97FEACE7809FF93FEDCFCA19E37FCC2" bold="true" box="[312,515,803,829]" pageId="28" pageNumber="29">Specific epithet.</emphasis>
The specific epithet,
<emphasis id="B97FEACE7809FF93FCF6FCA69F6EFCC2" box="[786,858,804,829]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="4C0B4D5F7809FF93FCF6FCA69F62FCC2" box="[786,854,804,829]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fugax">fugax</taxonomicName>
,
</emphasis>
meaning swift, is a manuscript name from Max Liebke's collection in WAR that I am retaining, however, I have chosen a different
<typeStatus id="54B0887E7809FF93FED0FCF69D95FC71" box="[308,417,884,910]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
specimen than he labeled with his manuscript name, a male that is in better condition.
</paragraph>
</subSubSection>
</treatment>
</document>