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<document id="2D980EE4A950CDC5780F3A348C4CA23E" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA780BFF90FEECFE33984DFB59" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra quesada Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="32" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="31" updateTime="1698190273211" updateUser="plazi">
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<mods:title id="F5CE8ED04CB8A6F83117DDB0CDC8ACAB">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
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<mods:name id="DF42342C3AB730B794D175DAA754C866" type="personal">
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<mods:roleTerm id="826812E39E239B257262BB57E761DDD4">Author</mods:roleTerm>
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<mods:namePart id="1D01992786C3A386B373B071DBAFE211">Erwin, Terry L.</mods:namePart>
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<mods:date id="DF0C80FE6B1C0FB1A813A11F23DB3F09">2002</mods:date>
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<mods:identifier id="3B52FA92ABD242A52B2994F76725C37C" type="DOI">10.5281/zenodo.162803</mods:identifier>
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<treatment id="03A287CA780BFF90FEECFE33984DFB59" ID-DOI="http://doi.org/10.5281/zenodo.6277984" ID-GBIF-Taxon="121065210" ID-Zenodo-Dep="6277984" LSID="urn:lsid:plazi:treatment:03A287CA780BFF90FEECFE33984DFB59" httpUri="http://treatment.plazi.org/id/03A287CA780BFF90FEECFE33984DFB59" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="C3116557780BFF91FEECFE339E5BFE0C" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BB436DC780BFF91FEECFE339E97FE34" blockId="30.[264,675,433,499]" box="[264,675,433,459]" pageId="30" pageNumber="31">
<heading id="D0FC81B0780BFF91FEECFE339E97FE34" bold="true" box="[264,675,433,459]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="B97FEACE780BFF91FEECFE339E97FE34" bold="true" box="[264,675,433,459]" pageId="30" pageNumber="31">
<taxonomicName id="4C0B4D5F780BFF91FEECFE339E33FE34" ID-CoL="664KY" authority="Erwin" authorityName="Erwin" box="[264,519,433,459]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="quesada" status="sp. nov.">
<emphasis id="B97FEACE780BFF91FEECFE339D9BFE34" bold="true" box="[264,431,433,459]" italics="true" pageId="30" pageNumber="31">Agra quesada</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B5780BFF91FDF0FE339E97FE34" box="[532,675,433,459]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC780BFF91FEECFE5B9E5BFE0C" blockId="30.[264,675,433,499]" box="[264,623,473,499]" pageId="30" pageNumber="31">
(
<figureCitation id="13302A59780BFF91FEF5FE5B9D9EFE0C" box="[273,426,473,499]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="30" pageNumber="31">Figs. 10a, 10</figureCitation>
b, 10c, 10d, 10e)
</paragraph>
</subSubSection>
<subSubSection id="C3116557780BFF91FEECFDA19E5AFD71" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BB436DC780BFF91FEECFDA19E5AFD71" blockId="30.[264,1325,547,654]" pageId="30" pageNumber="31">
<emphasis id="B97FEACE780BFF91FEECFDA19DB5FDC2" bold="true" box="[264,385,547,573]" pageId="30" pageNumber="31">
<typeStatus id="54B0887E780BFF91FEECFDA19D48FDC2" box="[264,380,547,573]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<emphasis id="B97FEACE780BFF91FE30FDA19EA7FDC2" bold="true" box="[468,659,547,573]" pageId="30" pageNumber="31">
<collectingCountry id="F31C764C780BFF91FE30FDA19EBAFDC2" box="[468,654,547,573]" name="Costa Rica" pageId="30" pageNumber="31">COSTA RICA</collectingCountry>
:
</emphasis>
Guanacaste, P.N. Guanacaste, Estacíon
<taxonomicName id="4C0B4D5F780BFF91FB89FDA69882FDC1" box="[1133,1206,548,574]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, 9.0 km S Santa Cecilia,
<quantity id="4CF39B39780BFF91FE53FDCE9DC9FD99" box="[439,509,588,614]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="m" value="700.0">700m</quantity>
,
<geoCoordinate id="EE3F501B780BFF91FDEEFDCE9EA8FD99" box="[522,668,588,614]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-85.42777">85°25'40”W</geoCoordinate>
<geoCoordinate id="EE3F501B780BFF91FD42FDCE9F07FD99" box="[678,819,588,614]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="15" value="10.990556">10°59'26&quot;N</geoCoordinate>
, LN 330200,380200, December, (C. Moraga)(
<taxonomicName id="4C0B4D5F780BFF91FEA7FDF69DAEFD71" box="[323,410,628,654]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780BFF91FEA7FDF69DAEFD71" box="[323,410,628,654]" pageId="30" pageNumber="31">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19780BFF91FE41FDF69DE3FD71" LSID="urn:lsid:biocol.org:col:14323" box="[421,471,628,654]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="30" pageNumber="31">CRI</collectionCode>
000­959644)
</paragraph>
</subSubSection>
<caption id="DF746654780BFF91FEECF88C9867F894" httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="30" pageNumber="31" targetBox="[378,1210,719,1773]" targetPageId="30">
<paragraph id="8BB436DC780BFF91FEECF88C9867F894" blockId="30.[264,1324,1806,1899]" pageId="30" pageNumber="31">
<emphasis id="B97FEACE780BFF91FEECF88C9DA4F8D9" bold="true" box="[264,400,1806,1830]" pageId="30" pageNumber="31">FIGURE 10</emphasis>
.
<taxonomicName id="4C0B4D5F780BFF91FE7AF88C9E21F8DA" box="[414,533,1806,1829]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="quesada">
<emphasis id="B97FEACE780BFF91FE7AF88C9E21F8DA" box="[414,533,1806,1829]" italics="true" pageId="30" pageNumber="31">A. quesada</emphasis>
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, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S2L = 0.9; AL = 5.0.
</paragraph>
</caption>
<subSubSection id="C3116557780AFF90FEDCFE999E74FEA1" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BB436DC780AFF90FEDCFE999E74FEA1" blockId="31.[264,1325,283,1190]" pageId="31" pageNumber="32">
<emphasis id="B97FEACE780AFF90FEDCFE999D84FECA" bold="true" box="[312,432,283,309]" pageId="31" pageNumber="32">Diagnosis</emphasis>
. Femur infuscated in basal two­thirds; elytron with sutural apex prolonged, acutely pointed (
<figureCitation id="13302A59780AFF90FE28FEC69E16FEA1" box="[460,546,324,350]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="C3116557780AFF90FEDCFEE99DFFFD39" pageId="31" pageNumber="32" type="description">
<paragraph id="8BB436DC780AFF90FEDCFEE99DFFFD39" blockId="31.[264,1325,283,1190]" pageId="31" pageNumber="32">
<emphasis id="B97FEACE780AFF90FEDCFEE99DFDFE7A" bold="true" box="[312,457,363,389]" pageId="31" pageNumber="32">Description</emphasis>
. Color and luster: Piceous with rufous appendages, femur infuscated in basal two­thirds. Surface shiny. Form: Head (
<figureCitation id="13302A59780AFF90FCCCFE169FB8FE51" box="[808,908,404,430]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10a</figureCitation>
) moderately elongate behind eyes, rounded behind in female. Elytron with apex (
<figureCitation id="13302A59780AFF90FCD9FE3E9FA0FE29" box="[829,916,444,470]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
b) markedly oblique, lateral tooth small, sutural apex prolonged, acutely pointed; interneurs of large deep foveae in regular rows. Abdominal sternum VI (
<figureCitation id="13302A59780AFF90FD62FD8E9ED5FDD9" box="[646,737,524,550]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
c) shallowly U­notched in female, shallowly vnotched in male. Stylomere 2 (
<figureCitation id="13302A59780AFF90FD62FDB69ED4FDB1" box="[646,736,564,590]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
d). Aedeagus (
<figureCitation id="13302A59780AFF90FC76FDB69FDEFDB1" box="[914,1002,564,590]" captionStart="FIGURE 10. A" captionStartId="30.[264,367,1806,1830]" captionTargetBox="[378,1210,719,1773]" captionTargetId="figure@30.[374,1212,713,1778]" captionTargetPageId="30" captionText="FIGURE 10. A. quesada, female, dorsal aspect; a) head; b) apex of elytron; c) Sternum VI, female, ventral aspect; d) stylomere 2; e) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 3.8; WE = 2.4; SW = 4.6; S 2 L = 0.9; AL = 5.0." httpUri="https://zenodo.org/record/162814/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
e) with ostium of medium length, apex simple lobate, shaft of aedeagus with distinct swelling in basal half, markedly narrowed apically; parameres glabrous. Size: very large,
<quantity id="4CF39B39780AFF90FC4FFD069853FD61" box="[939,1127,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" metricValueMax="2.35" metricValueMin="2.23" pageId="31" pageNumber="32" unit="mm" value="22.9" valueMax="23.5" valueMin="22.3">22.3 to 23.5mm</quantity>
in length,
<quantity id="4CF39B39780AFF90FB03FD069D6FFD39" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="31" pageNumber="32" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.8 to 5.8mm</quantity>
in width.
</paragraph>
</subSubSection>
<subSubSection id="C3116557780AFF90FEA6FD519F11FB81" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BB436DC780AFF90FEA6FD51991EFBF9" blockId="31.[264,1325,283,1190]" pageId="31" pageNumber="32">
<emphasis id="B97FEACE780AFF90FEA6FD519E94FD12" bold="true" box="[322,672,723,749]" pageId="31" pageNumber="32">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C780AFF90FD4EFD569F07FD11" box="[682,819,724,750]" name="Costa Rica" pageId="31" pageNumber="32">Costa Rica</collectingCountry>
: 4
<typeStatus id="54B0887E780AFF90FCBDFD569FF8FD11" box="[857,972,724,750]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
, females, from the following Conservation Area: Guanacaste, La Amistad Caribe, Arenal Tilarán.
<collectingCountry id="F31C764C780AFF90FBD6FD7E9881FCE9" box="[1074,1205,764,790]" name="Costa Rica" pageId="31" pageNumber="32">Costa Rica</collectingCountry>
, Alajuela, 1f, P.N. Volcán Tenorio, Upala, Bijagua, Albergue Heliconias,
<quantity id="4CF39B39780AFF90FC09FCA69807FCC1" box="[1005,1075,804,830]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="m" value="700.0">700m</quantity>
, LN 422600,299100, May, (G. Rodriguez)(
<taxonomicName id="4C0B4D5F780AFF90FDC0FCCE9E4CFC99" box="[548,632,844,870]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780AFF90FDC0FCCE9E4CFC99" box="[548,632,844,870]" pageId="31" pageNumber="32">INBIO</collectionCode>
</taxonomicName>
: INB0003074640); Guanacaste, 1f, P.N. Guanacaste, Estacíon
<taxonomicName id="4C0B4D5F780AFF90FE60FCF69DF9FC71" box="[388,461,884,910]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, Santa Cecilia, 9.0 km S,
<quantity id="4CF39B39780AFF90FCBDFCF69FABFC71" box="[857,927,884,910]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="m" value="700.0">700m</quantity>
, 85°25'40'W
<geoCoordinate id="EE3F501B780AFF90FBB8FCF698DCFC71" box="[1116,1256,884,910]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="10.990556">10°59'26&quot;N</geoCoordinate>
, LN 330200,380200, November, (GNP Biodiversity Survey)(
<taxonomicName id="4C0B4D5F780AFF90FC20FC1E982EFC49" box="[964,1050,924,950]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780AFF90FC20FC1E982EFC49" box="[964,1050,924,950]" pageId="31" pageNumber="32">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19780AFF90FBC9FC1E9855FC49" LSID="urn:lsid:biocol.org:col:14323" box="[1069,1121,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="31" pageNumber="32">CRI</collectionCode>
001­024201), 1f, May ­ June, (P. Rios)(
<taxonomicName id="4C0B4D5F780AFF90FDEEFC469E54FC21" box="[522,608,964,990]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780AFF90FDEEFC469E54FC21" box="[522,608,964,990]" pageId="31" pageNumber="32">INBIO</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19780AFF90FD88FC469EABFC21" LSID="urn:lsid:biocol.org:col:14323" box="[620,671,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="31" pageNumber="32">CRI</collectionCode>
001­355554); Limón, 1f, Manzanillo,
<collectionCode id="ED1AAE19780AFF90FBBAFC46989CFC21" box="[1118,1192,964,990]" pageId="31" pageNumber="32">RNFS</collectionCode>
Gandoca y Manzanillo,
<quantity id="4CF39B39780AFF90FE74FC6E9DEEFBF9" box="[400,474,1004,1030]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="31" pageNumber="32" unit="m" value="5.0" valueMax="10.0" valueMin="0.0">0­10m</quantity>
,
<geoCoordinate id="EE3F501B780AFF90FE3BFC6E9E67FBF9" box="[479,595,1004,1030]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="9.597777">9º35'52”N</geoCoordinate>
<geoCoordinate id="EE3F501B780AFF90FDB0FC6E9EEAFBF9" box="[596,734,1004,1030]" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="-82.605835">82º36'21”W</geoCoordinate>
,November,(F.
<taxonomicName id="4C0B4D5F780AFF90FC98FC6E9FC8FBF9" box="[892,1020,1004,1030]" pageId="31" pageNumber="32">A.Quesada</taxonomicName>
)(
<taxonomicName id="4C0B4D5F780AFF90FBE8FC6E9862FBF9" box="[1036,1110,1004,1030]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED1AAE19780AFF90FBE8FC6E9862FBF9" box="[1036,1110,1004,1030]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="31" pageNumber="32">INBio</collectionCode>
</taxonomicName>
:
<collectionCode id="ED1AAE19780AFF90FBBEFC6E98B8FBF9" LSID="urn:lsid:biocol.org:col:14323" box="[1114,1164,1004,1030]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="31" pageNumber="32">CRI</collectionCode>
000­784200).
</paragraph>
<paragraph id="8BB436DC780AFF90FEDCFB919F11FB81" blockId="31.[264,1325,283,1190]" pageId="31" pageNumber="32">
<emphasis id="B97FEACE780AFF90FEDCFB919DCBFBD2" bold="true" box="[312,511,1043,1069]" pageId="31" pageNumber="32">Specific epithet.</emphasis>
The specific epithet,
<emphasis id="B97FEACE780AFF90FCE5FB969F5EFBD2" box="[769,874,1044,1069]" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="4C0B4D5F780AFF90FCE5FB969F52FBD2" box="[769,870,1044,1069]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="quesada">quesada</taxonomicName>
,
</emphasis>
is used as a noun in apposition based on the family name of the collector of the
<typeStatus id="54B0887E780AFF90FD18FBBE9F5DFBA9" box="[764,873,1084,1110]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
specimen, parataxonomist, Freddy
<taxonomicName id="4C0B4D5F780AFF90FAEBFBBE9D59FB81" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1" phylum="Arthropoda" rank="species" species="quesada" status="sp. nov.">A. Quesada</taxonomicName>
of San Ramón, Alajuela,
<collectingCountry id="F31C764C780AFF90FD79FBE69F15FB81" box="[669,801,1124,1150]" name="Costa Rica" pageId="31" pageNumber="32">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3116557780AFF90FEDCFB09984DFB59" box="[312,1145,1163,1190]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BB436DC780AFF90FEDCFB09984DFB59" blockId="31.[264,1325,283,1190]" box="[312,1145,1163,1190]" pageId="31" pageNumber="32">
<emphasis id="B97FEACE780AFF90FEDCFB099D4AFB5A" bold="true" box="[312,382,1163,1189]" pageId="31" pageNumber="32">Notes</emphasis>
. Adults of this species are among the largest
<taxonomicName id="4C0B4D5F780AFF90FC75FB0E9FF8FB5A" box="[913,972,1164,1189]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEACE780AFF90FC75FB0E9FF8FB5A" box="[913,972,1164,1189]" italics="true" pageId="31" pageNumber="32">Agra</emphasis>
</taxonomicName>
in
<collectingCountry id="F31C764C780AFF90FC15FB0E9842FB59" box="[1009,1142,1164,1190]" name="Costa Rica" pageId="31" pageNumber="32">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>