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<document id="BA09DF73FAB9C87955FD00754445D481" ID-DOI="10.5281/zenodo.162803" ID-GBIF-Dataset="f514e964-2196-461c-84ff-64243643bc81" ID-ISSN="1175­5326" ID-Zenodo-Dep="162803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459445184067" checkinUser="plazi" docAuthor="Erwin, Terry L." docDate="2002" docId="03A287CA7802FF97FEECFCA69E0AFC71" docLanguage="en" docName="zt00119.pdf" docOrigin="Zootaxa 119" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agra turrialba Erwin, new species" docType="treatment" docVersion="11" lastPageNumber="25" masterDocId="FF9BFFB27815FF8FFFE4FF829C34FFFF" masterDocTitle="The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="24" updateTime="1698190273211" updateUser="plazi">
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<mods:title id="AA7AE573EBCB286A39FCBD2C75F9ABD7">The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina)</mods:title>
</mods:titleInfo>
<mods:name id="A469B5B8D0AC93CC82E55FCEE9F9FDF8" type="personal">
<mods:role id="7BC97005BF35B684C3140AF2066C87A5">
<mods:roleTerm id="A6C08C437E314D18387FEBD39E334BAA">Author</mods:roleTerm>
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<mods:namePart id="C5CD856A9471809591D02C3B72091AD5">Erwin, Terry L.</mods:namePart>
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<mods:typeOfResource id="8BC202B58A35DB41BD93B5F5A5BE5600">text</mods:typeOfResource>
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<mods:title id="CB03163302F08585315C6224714FF831">Zootaxa</mods:title>
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<mods:part id="EC057E140E31833F65ED2BD3FA3832FD">
<mods:date id="BB463D36D9A5DB70C4E2337179E75D25">2002</mods:date>
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<mods:classification id="FE011755E10286E592D2BAA100751B0B">journal article</mods:classification>
<mods:identifier id="1A63A8722D5C0270D019E6A091EAE1D1" type="DOI">10.5281/zenodo.162803</mods:identifier>
<mods:identifier id="F7229738C3978B18223F7F04028A151C" type="GBIF-Dataset">f514e964-2196-461c-84ff-64243643bc81</mods:identifier>
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<treatment id="03A287CA7802FF97FEECFCA69E0AFC71" ID-DOI="http://doi.org/10.5281/zenodo.6277976" ID-GBIF-Taxon="121065214" ID-Zenodo-Dep="6277976" LSID="urn:lsid:plazi:treatment:03A287CA7802FF97FEECFCA69E0AFC71" httpUri="http://treatment.plazi.org/id/03A287CA7802FF97FEECFCA69E0AFC71" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<subSubSection id="C31165577802FF98FEECFCA69DC8FC99" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BB436DC7802FF98FEECFCA69E9FFCC2" blockId="23.[264,683,803,870]" box="[264,683,803,830]" pageId="23" pageNumber="24">
<heading id="D0FC81B07802FF98FEECFCA69E9FFCC2" bold="true" box="[264,683,803,830]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B97FEACE7802FF98FEECFCA69E9FFCC2" bold="true" box="[264,683,803,830]" pageId="23" pageNumber="24">
<taxonomicName id="4C0B4D5F7802FF98FEECFCA69E3AFCC2" ID-CoL="B2ZY" authority="Erwin" authorityName="Erwin" box="[264,526,803,830]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="turrialba" status="sp. nov.">
<emphasis id="B97FEACE7802FF98FEECFCA69D8CFCC1" bold="true" box="[264,440,804,830]" italics="true" pageId="23" pageNumber="24">Agra turrialba</emphasis>
Erwin
</taxonomicName>
,
<taxonomicNameLabel id="A24C57B57802FF98FDFFFCA19E9FFCC2" box="[539,683,803,829]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436DC7802FF98FEECFCCE9DC8FC99" blockId="23.[264,683,803,870]" box="[264,508,844,870]" pageId="23" pageNumber="24">
(
<figureCitation id="13302A597802FF98FEF5FCCE9DB9FC99" box="[273,397,844,870]" captionStart="FIGURE 6. A" captionStartId="24.[264,368,1771,1795]" captionTargetBox="[458,1128,988,1737]" captionTargetId="figure@24.[450,1138,969,1745]" captionTargetPageId="24" captionText="FIGURE 6. A. turrialba, dorsal aspect, male: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 1.7; male SW = 3.2, female SW = 3.2; AL = 3.0." httpUri="https://zenodo.org/record/162810/files/figure.png" pageId="23" pageNumber="24">Figs. 6a, 6</figureCitation>
b, 6c, 6d)
</paragraph>
</subSubSection>
<subSubSection id="C31165577802FF98FEECFC199E45FC21" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BB436DC7802FF98FEECFC199E45FC21" blockId="23.[264,1325,923,1870]" pageId="23" pageNumber="24">
<emphasis id="B97FEACE7802FF98FEECFC199DB5FC4A" bold="true" box="[264,385,923,949]" pageId="23" pageNumber="24">
<typeStatus id="54B0887E7802FF98FEECFC199D48FC4A" box="[264,380,923,949]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<collectingCountry id="F31C764C7802FF98FE32FC1E9EBCFC49" box="[470,648,924,950]" name="Costa Rica" pageId="23" pageNumber="24">COSTA RICA</collectingCountry>
: Cartago,
<taxonomicName id="4C0B4D5F7802FF98FCE4FC1E9F5BFC49" box="[768,879,924,950]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
,
<quantity id="4CF39B397802FF98FC9FFC1E9FF5FC49" box="[891,961,924,950]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="24" unit="m" value="600.0">600m</quantity>
,
<geoCoordinate id="EE3F501B7802FF98FC2AFC1E9818FC49" box="[974,1068,924,950]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="9.883333">09º53'N</geoCoordinate>
<geoCoordinate id="EE3F501B7802FF98FBD1FC1E9896FC49" box="[1077,1186,924,950]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="-83.63333">83º 38'W</geoCoordinate>
, June (J. E. Wappes)(
<collectionCode id="ED1AAE197802FF98FE9DFC469DE4FC21" LSID="urn:lsid:biocol.org:col:34871" box="[377,464,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24">USNM</collectionCode>
: ADP58598).
</paragraph>
</subSubSection>
<subSubSection id="C31165577802FF98FEDCFC699D65FB81" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BB436DC7802FF98FEDCFC699D65FB81" blockId="23.[264,1325,923,1870]" pageId="23" pageNumber="24">
<emphasis id="B97FEACE7802FF98FEDCFC699D83FBFA" bold="true" box="[312,439,1003,1029]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
This is the only species of the cajennensis species­group in
<collectingCountry id="F31C764C7802FF98FB6FFC6E9938FBF9" box="[1163,1292,1004,1030]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
in which males have abdominal sternum IV elongate and medially lobed so that it covers partially sternum V; males also with hind femur deeply excavate and anterio­medially glabrous.
</paragraph>
</subSubSection>
<subSubSection id="C31165577802FF97FEDCFB09993AFD11" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="8BB436DC7802FF98FEDCFB099F88F9F1" blockId="23.[264,1325,923,1870]" pageId="23" pageNumber="24">
<emphasis id="B97FEACE7802FF98FEDCFB099DE4FB5A" bold="true" box="[312,464,1163,1189]" pageId="23" pageNumber="24">Description.</emphasis>
Color and luster: Piceous with bright rufous appendages, head rufinistic, femur slightly infuscated. Surface very shiny. Form: Medium­sized and narrow, females with slightly broader elytra. Head (
<figureCitation id="13302A597802FF98FD47FB5E9EC3FB09" box="[675,759,1244,1270]" captionStart="FIGURE 6. A" captionStartId="24.[264,368,1771,1795]" captionTargetBox="[458,1128,988,1737]" captionTargetId="figure@24.[450,1138,969,1745]" captionTargetPageId="24" captionText="FIGURE 6. A. turrialba, dorsal aspect, male: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 1.7; male SW = 3.2, female SW = 3.2; AL = 3.0." httpUri="https://zenodo.org/record/162810/files/figure.png" pageId="23" pageNumber="24">Fig. 6a</figureCitation>
) behind eyes small and narrow, rounded in both sexes, shorter than prothorax. Prothorax moderately short and medially swollen, gradually constricted with sides convergent to anterior collar. Elytron (
<figureCitation id="13302A597802FF98FC39FAAE9811FAB9" box="[989,1061,1324,1350]" captionStart="FIGURE 6. A" captionStartId="24.[264,368,1771,1795]" captionTargetBox="[458,1128,988,1737]" captionTargetId="figure@24.[450,1138,969,1745]" captionTargetPageId="24" captionText="FIGURE 6. A. turrialba, dorsal aspect, male: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 1.7; male SW = 3.2, female SW = 3.2; AL = 3.0." httpUri="https://zenodo.org/record/162810/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
b) narrow and moderately convex; apex markedly oblique, lateral tooth prominent, sutural tooth elongate, markedly acute; interneurs each a series of linear foveae. Abdominal sternum VI (
<figureCitation id="13302A597802FF98FB2BFAFE9921FA69" box="[1231,1301,1404,1430]" captionStart="FIGURE 6. A" captionStartId="24.[264,368,1771,1795]" captionTargetBox="[458,1128,988,1737]" captionTargetId="figure@24.[450,1138,969,1745]" captionTargetPageId="24" captionText="FIGURE 6. A. turrialba, dorsal aspect, male: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 1.7; male SW = 3.2, female SW = 3.2; AL = 3.0." httpUri="https://zenodo.org/record/162810/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
c) deeply round­notched in male, V­notched in female. Aedeagus (
<figureCitation id="13302A597802FF98FBF6FA269868FA41" box="[1042,1116,1444,1470]" captionStart="FIGURE 6. A" captionStartId="24.[264,368,1771,1795]" captionTargetBox="[458,1128,988,1737]" captionTargetId="figure@24.[450,1138,969,1745]" captionTargetPageId="24" captionText="FIGURE 6. A. turrialba, dorsal aspect, male: a) head; b) apex of elytron; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = 2.7; WE = 1.7; male SW = 3.2, female SW = 3.2; AL = 3.0." httpUri="https://zenodo.org/record/162810/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
d) with markedly short ostium, apex simple lobe; phallus ventrally with a longitudinal depression. Size: medium, 14.6 to 17.0mm in length,
<quantity id="4CF39B397802FF98FD4BFA769F7FF9F1" box="[687,843,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="23" pageNumber="24" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4 to 3.8mm</quantity>
in width.
</paragraph>
<paragraph id="8BB436DC7802FF97FEDCF999993AFD11" blockId="23.[264,1325,923,1870]" lastBlockId="24.[264,1324,284,910]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="B97FEACE7802FF98FEDCF9999EA2F9CA" bold="true" box="[312,662,1563,1589]" pageId="23" pageNumber="24">Other specimens examined.</emphasis>
<collectingCountry id="F31C764C7802FF98FD7BF99E9F12F9C9" box="[671,806,1564,1590]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
: 19
<typeStatus id="54B0887E7802FF98FCBFF99E9FFBF9C9" box="[859,975,1564,1590]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D0DFD557802FF98FC39F99E9805F9C9" box="[989,1073,1564,1590]" pageId="23" pageNumber="24" type="male">8 male</specimenCount>
,
<specimenCount id="9D0DFD557802FF98FBDBF99E9882F9C9" box="[1087,1206,1564,1590]" pageId="23" pageNumber="24" type="female">7 females</specimenCount>
, from the following Conservation Areas: Cordillera Volcánica Central, Guanacaste, Huetar Norte, Tortuguero.
<collectingCountry id="F31C764C7802FF98FE7CF9EE9E2FF979" box="[408,539,1644,1670]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
, Cartago, 1f, A.C. Amistad, Monumento Nacional Guayabo,
<taxonomicName id="4C0B4D5F7802FF98FB0CF9EE9D96F951" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">
Turrialba,
<quantity id="4CF39B397802FF98FEABF9169D96F951" box="[335,418,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="24" unit="m" value="1100.0">1100m</quantity>
</taxonomicName>
, LN 217200,570300 #3202, September, (G. Fonseca) (
<taxonomicName id="4C0B4D5F7802FF98FBBFF9169887F951" box="[1115,1203,1684,1710]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">INBIO</taxonomicName>
: CRI002­ 039094), 1f, June, (G. Ekis)(USNM: ADP47280), 1f, February, (H.F. Howden &amp; A. Howden)(UASM: ADP54795), 1f,
<taxonomicName id="4C0B4D5F7802FF98FD20F9669F07F901" box="[708,819,1764,1790]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
, 3.0 km SE,
<taxonomicName id="4C0B4D5F7802FF98FC0BF9669871F901" box="[1007,1093,1764,1790]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="51" phylum="Arthropoda" rank="species" species="catie">CATIE</taxonomicName>
,
<geoCoordinate id="EE3F501B7802FF98FBBFF96698F6F901" box="[1115,1218,1764,1790]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="-83.63333">83°38'W</geoCoordinate>
<geoCoordinate id="EE3F501B7802FF98FB37F966991CF901" box="[1235,1320,1764,1790]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="9.883333">9°53'N</geoCoordinate>
, December, (F.T. Hovore)(FTHC: ADP6381),
<quantity id="4CF39B397802FF98FC85F88E9FBDF8D9" box="[865,905,1804,1830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1.0">1m</quantity>
, August, (D.W. Alsop)(CUNY: ADP58621),
<quantity id="4CF39B397802FF98FE4AF8B69DE1F8B1" box="[430,469,1844,1870]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1.0">1m</quantity>
, June, (J.E. Wappes)(USNM: ADP58598),
<quantity id="4CF39B397802FF98FBE6F8B6981DF8B1" box="[1026,1065,1844,1870]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1.0">1m</quantity>
, June, (E. Giesbert) (FSCA: ADP
<date id="FFB5101C780DFF97FE56FE9E9E3FFEC9" box="[434,523,284,310]" pageId="24" pageNumber="25" value="15-10-31">103115</date>
),
<quantity id="4CF39B39780DFF97FDCDFE9E9E65FEC9" box="[553,593,284,310]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, Tuis,
<quantity id="4CF39B39780DFF97FD50FE9E9ECDFEC9" box="[692,761,284,310]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="24" pageNumber="25" unit="m" value="732.0">732m</quantity>
,
<geoCoordinate id="EE3F501B780DFF97FCEBFE9E9F43FEC9" box="[783,887,284,310]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-83.8">83°48'W</geoCoordinate>
<geoCoordinate id="EE3F501B780DFF97FC6CFE9E9FE9FEC9" box="[904,989,284,310]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="9.833333">9°50'N</geoCoordinate>
, (C.H. Lankester)(USNM: ADP10014), 1f, Tucurrique,
<geoCoordinate id="EE3F501B780DFF97FD75FEC69ECCFEA1" box="[657,760,324,350]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-83.71667">83°43'W</geoCoordinate>
<geoCoordinate id="EE3F501B780DFF97FCF5FEC69F52FEA1" box="[785,870,324,350]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="9.85">9°51'N</geoCoordinate>
, (P. Schild &amp; Burgdorf)(USNM: ADP10013); Guanacaste,
<quantity id="4CF39B39780DFF97FDDAFEEE9E52FE79" box="[574,614,364,390]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, P.N. Guanacaste, Estacíon
<taxonomicName id="4C0B4D5F780DFF97FC57FEEE9FC8FE79" box="[947,1020,364,390]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="54" phylum="Arthropoda" rank="species" species="pitilla">Pitilla</taxonomicName>
, Santa Cecilia, 9.0 km S,
<quantity id="4CF39B39780DFF97FEECFE169D7AFE51" box="[264,334,404,430]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="25" unit="m" value="700.0">700m</quantity>
, 85°25'40'W
<geoCoordinate id="EE3F501B780DFF97FDE6FE169EBAFE51" box="[514,654,404,430]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="15" value="10.990556">10°59'26&quot;N</geoCoordinate>
, LN 330200,380200, November, (C. Moraga &amp; P. Rios)(
<taxonomicName id="4C0B4D5F780DFF97FEB5FE3E9D93FE29" box="[337,423,444,470]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">INBIO</taxonomicName>
: CRI000­
<date id="FFB5101C780DFF97FDC1FE3E9E4BFE29" box="[549,639,444,470]" pageId="24" pageNumber="25" value="44-11-27">112744</date>
),
<quantity id="4CF39B39780DFF97FD79FE3E9EF1FE29" box="[669,709,444,470]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, vcn. Dos Rios, January, (E. Giesbert)(LACM: ADP58594); Heredia,
<quantity id="4CF39B39780DFF97FDF6FE669E0EFE01" box="[530,570,484,510]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, Estacíon La Selva, Puerto Viejo, 3.0 km S,
<quantity id="4CF39B39780DFF97FBAEFE669883FE01" box="[1098,1207,484,510]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="24" pageNumber="25" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50­150m</quantity>
,
<geoCoordinate id="EE3F501B780DFF97FB20FE66991FFE01" box="[1220,1323,484,510]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-83.98333">83°59'W</geoCoordinate>
<geoCoordinate id="EE3F501B780DFF97FEECFD8E9D5FFDD9" box="[264,363,524,550]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="10.45">10°27'N</geoCoordinate>
, April, (INBIO­OET: CRI002­268518), M/10/388, Bosque primario,
<quantity id="4CF39B39780DFF97FB73FD8E988BFDD9" box="[1175,1215,524,550]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, November, (INBIO­OET: CRI002­232086), FVK/32/09,
<taxonomicName id="4C0B4D5F780DFF97FC86FDB6982FFDB2" box="[866,1051,564,589]" class="Magnoliopsida" family="Myristicaceae" genus="Virola" kingdom="Plantae" order="Magnoliales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="koschnyi">
<emphasis id="B97FEACE780DFF97FC86FDB6982FFDB2" box="[866,1051,564,589]" italics="true" pageId="24" pageNumber="25">Virola koschnyi</emphasis>
</taxonomicName>
, 1f, January, (INBIO­ OET: CRI002­724110), at light, L/00/666, 1f, Puerto Viejo, 3.0 km S, Finca La Selva,
<geoCoordinate id="EE3F501B780DFF97FEECFD069D5BFD61" box="[264,367,644,670]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-84.01667">84°01'W</geoCoordinate>
<geoCoordinate id="EE3F501B780DFF97FE66FD069DD1FD61" box="[386,485,644,670]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="10.433333">10°26'N</geoCoordinate>
, June, (H.
<taxonomicName id="4C0B4D5F780DFF97FD97FD069F75FD61" box="[627,833,644,670]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="32" phylum="Arthropoda" rank="species" species="hespenheide">A. Hespenheide</taxonomicName>
)(HAH: ADP76792),
<quantity id="4CF39B39780DFF97FBB4FD06984CFD61" box="[1104,1144,644,670]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="1.0">1m</quantity>
, April, (H.
<taxonomicName id="4C0B4D5F780DFF97FAEAFD069D91FD39" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="32" phylum="Arthropoda" rank="species" species="hespenheide">A. Hespenheide</taxonomicName>
)(HAH: ADP5095); Limón, 1f, R.B. Hitoy Cerere, Estacíon Hitoy Cerere, Send. Espavel, 200M, LN 400750,570200, June, (W. Arana)(
<taxonomicName id="4C0B4D5F780DFF97FC33FD569818FD11" box="[983,1068,724,750]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="67" phylum="Arthropoda" rank="species" species="inbio">INBIO</taxonomicName>
: INB0003167631).
</paragraph>
</subSubSection>
<subSubSection id="C3116557780DFF97FEDCFD79984EFCC1" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BB436DC780DFF97FEDCFD79984EFCC1" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
<emphasis id="B97FEACE780DFF97FEDCFD799DCAFCEA" bold="true" box="[312,510,763,789]" pageId="24" pageNumber="25">Specific epithet.</emphasis>
The specific epithet,
<taxonomicName id="4C0B4D5F780DFF97FD1AFD7E9F5CFCEA" box="[766,872,764,789]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">
<emphasis id="B97FEACE780DFF97FD1AFD7E9F5CFCEA" box="[766,872,764,789]" italics="true" pageId="24" pageNumber="25">turrialba</emphasis>
</taxonomicName>
, is used as a noun in apposition based on the town
<taxonomicName id="4C0B4D5F780DFF97FE7EFCA69E32FCC1" box="[410,518,804,830]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="turrialba">Turrialba</taxonomicName>
in Cartago Province, in reference to the
<typeStatus id="54B0887E780DFF97FC06FCA69820FCC1" box="[994,1044,804,830]" pageId="24" pageNumber="25">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<paragraph id="8BB436DC780DFF97FEDCFCC99E0AFC71" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
<emphasis id="B97FEACE780DFF97FEDCFCC99DB1FC9A" bold="true" box="[312,389,843,869]" pageId="24" pageNumber="25">Notes.</emphasis>
This species is widespread in
<collectingCountry id="F31C764C780DFF97FD03FCCE9F53FC99" box="[743,871,844,870]" name="Costa Rica" pageId="24" pageNumber="25">Costa Rica</collectingCountry>
from sea level to 1100 meters altitude, in both dry and wet forest.
</paragraph>
</treatment>
</document>