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<document id="D9F364D744C9840510A861E6CE4F30F6" ID-DOI="10.11646/phytotaxa.66.1.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="5065874" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625288901423" checkinUser="felipe" docAuthor="Melai, Martina, Marchetti, Dino, Bernardello, Remo &amp; Peruzzi, Lorenzo" docDate="2012" docId="03A287C1653FFFB0FCE047B0EE16FA0A" docLanguage="en" docName="Phytotaxa.66.27-37.pdf" docOrigin="Phytotaxa 66 (1)" docSource="http://www.biotaxa.org/Phytotaxa/article/view/phytotaxa.66.1.5" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Leucanthemum ligusticum Marchetti, Bernardello, Melai &amp; Peruzzi 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FF9BFFB9653EFFB4FC774142EF06FFD5" masterDocTitle="A new diploid species of Leucanthemum (Asteraceae, Anthemideae) from Liguria (northwestern Italy)" masterLastPageNumber="37" masterPageNumber="27" pageNumber="28" updateTime="1698987123099" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4F4DA6221898E08AA8B5AD1A4A996C06">A new diploid species of Leucanthemum (Asteraceae, Anthemideae) from Liguria (northwestern Italy)</mods:title>
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<mods:namePart id="4AD6D4D9CCA3EEE2F85907B2F9A70D03">Melai, Martina</mods:namePart>
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<mods:namePart id="7F32AECD26C1801A8B7457E56C0A55D7">Marchetti, Dino</mods:namePart>
<mods:affiliation id="4642C193236CA42AD437DC292D558232">Via Isonzo 6, 54100, Massa, Italy</mods:affiliation>
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<mods:namePart id="87DB5001043C52D4DF00E2A67E71DA16">Bernardello, Remo</mods:namePart>
<mods:affiliation id="206202C17CDFBB1653F75379F8E7BC34">Via Isonzo 6, 54100, Massa, Italy</mods:affiliation>
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<mods:namePart id="07A01231D972C4C57F85D429FE782FDB">Peruzzi, Lorenzo</mods:namePart>
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<mods:title id="D4E55BB18D6FCC8E4501A9AEC6ACD0F4">Phytotaxa</mods:title>
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<mods:date id="7AF7FE954702F388574AF3EBF8D7935F">2012</mods:date>
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<treatment id="03A287C1653FFFB0FCE047B0EE16FA0A" ID-DOI="http://doi.org/10.5281/zenodo.5067389" ID-Zenodo-Dep="5067389" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A" httpUri="http://treatment.plazi.org/id/03A287C1653FFFB0FCE047B0EE16FA0A" lastPageId="4" lastPageNumber="31" pageId="1" pageNumber="28">
<subSubSection id="C311655C653FFFB5FCE047B0EA32F8DA" box="[151,1332,1778,1807]" pageId="1" pageNumber="28" type="nomenclature">
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<heading id="D0FC81BB653FFFB5FCE047B0EA32F8DA" box="[151,1332,1778,1807]" fontSize="12" level="2" pageId="1" pageNumber="28" reason="4">
<emphasis id="B97FEAC5653FFFB5FCE047B0EA32F8DA" bold="true" box="[151,1332,1778,1807]" pageId="1" pageNumber="28">
<taxonomicName id="4C0B4D54653FFFB5FCE047B0EB29F8DA" ID-CoL="6PS7K" authority="Marchetti, Bernardello, Melai &amp; Peruzzi" authorityName="Marchetti, Bernardello, Melai &amp; Peruzzi" authorityYear="2012" box="[151,1071,1778,1807]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="28" phylum="Tracheophyta" rank="species" species="ligusticum" status="sp. nov.">
<emphasis id="B97FEAC5653FFFB5FCE047B0EEFCF8DB" bold="true" box="[151,506,1778,1806]" italics="true" pageId="1" pageNumber="28">Leucanthemum ligusticum</emphasis>
Marchetti, Bernardello, Melai &amp; Peruzzi
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,
<emphasis id="B97FEAC5653FFFB5F84A47B0EBA6F8DB" bold="true" box="[1085,1184,1778,1806]" italics="true" pageId="1" pageNumber="28">
<taxonomicNameLabel id="A24C57BE653FFFB5F84A47B0EBA6F8DB" box="[1085,1184,1778,1806]" pageId="1" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13302A52653FFFB5F8C647B0EA0AF8DA" box="[1201,1292,1778,1807]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,244,1981,2002]" captionStartId-1="3.[151,246,1766,1787]" captionTargetBox-0="[151,1436,375,1960]" captionTargetBox-1="[151,1436,932,1746]" captionTargetId-0="figure-151@2.[151,1436,375,1960]" captionTargetId-1="figure-455@3.[151,1436,932,1746]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Leucanthemum ligusticum sp. nov. General view of the plant (A); closeup of a capitulum in top view (B) and bottom view (C)." captionText-1="FIGURE 2. Leucanthemum ligusticum sp. nov. Variability of basal leaves in individuals from Ponte di Lagoscuro (A), Rocche di Valletti (B), Rocchetta di Vara (C). Variability of cauline leaves, upper ones (sx) to lower ones (dx), from Ponte di Lagoscuro (D), Rocche di Valletti (E), Rocchetta di Vara (F). Scale bars: 10 cm." figureDoi-0="http://doi.org/10.5281/zenodo.5065878" figureDoi-1="http://doi.org/10.5281/zenodo.5065880" httpUri-0="https://zenodo.org/record/5065878/files/figure.png" httpUri-1="https://zenodo.org/record/5065880/files/figure.png" pageId="1" pageNumber="28">Figs. 1</figureCitation>
–2)
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</paragraph>
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<paragraph id="8BB436D7653FFFB5FCE04675EDECF8BA" blockId="1.[151,1436,1847,1903]" pageId="1" pageNumber="28">
<emphasis id="B97FEAC5653FFFB5FCE04675EFC9F89A" bold="true" box="[151,207,1847,1871]" pageId="1" pageNumber="28">Type</emphasis>
:—
<materialsCitation id="3B633C8A653FFFB5FC9B467AEDECF8BA" collectingDate="2011-05-07" collectorName="di Vara &amp; La Spezia &amp; M. Nero &amp; D. Marchetti" country="Italy" county="Val di Vara" elevation="225" location="Rocchetta di Vara" municipality="La Spezia" pageId="1" pageNumber="28" specimenCount="1" stateProvince="Liguria" typeStatus="holotype">
<collectingCountry id="F31C7647653FFFB5FC9B467AEE3DF89A" box="[236,315,1848,1871]" name="Italy" pageId="1" pageNumber="28">ITALY</collectingCountry>
,
<emphasis id="B97FEAC5653FFFB5FD314675EEA5F89A" bold="true" box="[326,419,1847,1871]" pageId="1" pageNumber="28">
<collectingRegion id="49CFF835653FFFB5FD314675EE98F89A" box="[326,414,1847,1871]" country="Italy" name="Liguria" pageId="1" pageNumber="28">Liguria</collectingRegion>
.
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<collectingCounty id="62D54E5B653FFFB5FDDC467AED25F89A" box="[427,547,1848,1871]" pageId="1" pageNumber="28">
Val
<collectorName id="26FE5301653FFFB5FDA1467AED25F89A" box="[470,547,1848,1871]" pageId="1" pageNumber="28">di Vara</collectorName>
</collectingCounty>
(
<collectorName id="26FE5301653FFFB5FE45467AEDA4F89A" box="[562,674,1848,1871]" pageId="1" pageNumber="28">
<collectingMunicipality id="6BD0ACAD653FFFB5FE45467AEDA4F89A" box="[562,674,1848,1871]" pageId="1" pageNumber="28">La Spezia</collectingMunicipality>
</collectorName>
),
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Rocchetta
<collectorName id="26FE5301653FFFB5FF5F467AEC7EF89A" box="[808,888,1848,1871]" pageId="1" pageNumber="28">di Vara</collectorName>
</location>
, alla base del
<collectorName id="26FE5301653FFFB5F866467AEB69F89A" box="[1041,1135,1848,1871]" pageId="1" pageNumber="28">M. Nero</collectorName>
, ca.
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<elevation id="0026D1E4653FFFB5F8D7467AEBE0F89A" box="[1184,1254,1848,1871]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="1" pageNumber="28" unit="m" value="225.0">225 m</elevation>
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,
<date id="FFB51017653FFFB5F885467AEA71F89A" box="[1266,1399,1848,1871]" pageId="1" pageNumber="28" value="2011-05-07">
<collectingDate id="EFF1E9FF653FFFB5F885467AEA71F89A" box="[1266,1399,1848,1871]" pageId="1" pageNumber="28" value="2011-05-07">7 May 2011</collectingDate>
</date>
,
<emphasis id="B97FEAC5653FFFB5F9F5467AEE5CF8BA" italics="true" pageId="1" pageNumber="28">
<collectorName id="26FE5301653FFFB5F9F5467AEE29F8BA" pageId="1" pageNumber="28">D. Marchetti</collectorName>
s.n.
</emphasis>
(
<typeStatus id="54B08875653FFFB5FD1E461AEECAF8BA" box="[361,460,1880,1903]" pageId="1" pageNumber="28" type="holotype">holotype</typeStatus>
: PI!; isotype: B!, FI!, GE!)
</materialsCitation>
</paragraph>
<paragraph id="8BB436D7653FFFB6FCE046DAED9CFE9A" blockId="1.[151,1436,1944,2031]" lastBlockId="2.[199,1437,152,335]" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
<emphasis id="B97FEAC5653FFFB5FCE046DAEBD8F83A" italics="true" pageId="1" pageNumber="28">
A
<taxonomicName id="4C0B4D54653FFFB5FCC146DAEE7AF87A" authority="Pau" authorityName="Pau" box="[182,380,1944,1967]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="28" phylum="Tracheophyta" rank="species" species="plurifloro">L. plurifloro Pau</taxonomicName>
flore ligulato 2.5–3.8(4.8) non (3.0)3.5–5.5(6.0) mm lato, flore tubuloso max.
<quantity id="4CF39B32653FFFB5F95646DAEA62F87A" box="[1313,1380,1944,1967]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="28" unit="mm" value="3.0">3 mm</quantity>
non (3.1)3.8–4.6(5.0) mm longo, squama involucrali (0.9)1.0–1.5(1.9) non (0.8)1.8–2.2(2.5) lata, petiolo foliorum caulis inferiorum [(15)20–41(49) pro (13)16–21(22) mm] longiore, dentibus foliorum caulis inferiorum
</emphasis>
(
<emphasis id="B97FEAC5653FFFB6F898469AEEFAFF1A" italics="true" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
1.0)1.5–2.7(3.0) mm non (2.7)
<quantity id="4CF39B32653CFFB6FD2341DAEED4FF7A" box="[340,466,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.3" metricValueMin="2.8" pageId="2" pageNumber="29" unit="mm" value="3.55" valueMax="4.3" valueMin="2.8">2.8–4.3 mm</quantity>
longis, foliis caulis superioribus longioribus [(18)19–28(33) pro (7)8–19(21) mm], differt. A
<taxonomicName id="4C0B4D54653CFFB6FC9641FAEE8DFF1A" box="[225,395,184,207]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="monspeliensi">L. monspeliensi</taxonomicName>
(L.) H. J.
</emphasis>
Coste
<emphasis id="B97FEAC5653CFFB6FE3C41FAED9CFE9A" italics="true" pageId="2" pageNumber="29">
flore ligulato 2.5–3.8(4.8) non (3.3)3.4–3.9(4.0) mm lato, squama involucrali (0.9)1.0–1.5(1.9) non (1.2)1.3–1.9(2) mm lata, foliis basalibus longioribus [(31)32–68(80) vs. (16)17–29(32) mm] et latioribus [(19)22–40.8(58) vs. (9)
<quantity id="4CF39B32653CFFB6FE2241BAEDD8FEDA" box="[597,734,248,271]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.75" metricValueMin="1.0" pageId="2" pageNumber="29" unit="mm" value="13.75" valueMax="17.5" valueMin="10.0">10–17.5 mm</quantity>
], pluribus dentibus praeditis [(24)25–72(78) pro (12)13–16(17)] et petiolo longiore [(32)38–96(103) pro (27)30–44(45) mm], foliis caulis inferioribus petiolum longiorem ferentibus [(15)20–41(49) pro (17)18–28(29)], differt.
</emphasis>
</paragraph>
<caption id="DF74665F653CFFB6FCE046FFEFBBF83A" ID-DOI="http://doi.org/10.5281/zenodo.5065878" ID-Zenodo-Dep="5065878" httpUri="https://zenodo.org/record/5065878/files/figure.png" pageId="2" pageNumber="29" startId="2.[151,244,1981,2002]" targetBox="[151,1436,375,1960]" targetPageId="2">
<paragraph id="8BB436D7653CFFB6FCE046FFEFBBF83A" blockId="2.[151,1435,1981,2031]" pageId="2" pageNumber="29">
<emphasis id="B97FEAC5653CFFB6FCE046FFEE0AF807" bold="true" box="[151,268,1981,2002]" pageId="2" pageNumber="29">FIGURE 1.</emphasis>
<taxonomicName id="4C0B4D54653CFFB6FD6646FFED0BF807" authority="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi, 2012" authorityName="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi" authorityYear="2012" box="[273,525,1981,2002]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="ligusticum" status="sp. nov.">
<emphasis id="B97FEAC5653CFFB6FD6646FFED0BF807" box="[273,525,1981,2002]" italics="true" pageId="2" pageNumber="29">Leucanthemum ligusticum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57BE653CFFB6FE6446FFED5BF807" box="[531,605,1981,2002]" pageId="2" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
General view of the plant (A); closeup of a capitulum in top view (B) and bottom view (C).
</paragraph>
</caption>
<paragraph id="8BB436D7653DFFB7FCE041DAECC1FCBF" blockId="3.[151,1437,152,874]" pageId="3" pageNumber="30">
Short-lived perennial herb (34)43–72(100) cm tall, sparsely hairy (especially on the lower part of stem and under the capitula), with rosettes arising from an aerial, oblique, violet woody rootstock, up to
<quantity id="4CF39B32653DFFB7F97A41FCEA51FF0D" box="[1293,1367,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="30" unit="cm" value="20.0">20 cm</quantity>
long. Stems several, erect-decumbent, (0.9)1.0–3.3(6.0) mm thick at the base, divided above into several branches, each bearing 1 capitulum. Basal leaves (31)32–68(80) mm long, (19)22–41(58) mm wide, thin, hairless, ovate-spathulate, pinnatipartite or, rarely, bipinnatipartite, with dentate or deeply dentate margins and (24)25–72(78) teeth, which are (1.8)
<quantity id="4CF39B32653DFFB7FE1D401BEDEDFEA6" box="[618,747,345,371]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.5" metricValueMin="2.0" pageId="3" pageNumber="30" unit="mm" value="3.25" valueMax="4.5" valueMin="2.0">2–4.5 mm</quantity>
long; petioles (32)38–96(103) mm long, withering at flowering. Lower cauline leaves (15)19–32(40) mm long, (9.5)12–21(25) mm wide, spathulate, pinnatipartite or, rarely, bipinnatipartite, with dentate or deeply dentate margins and (12)15–24(34) teeth, which are (1.0)1.5–2.7(3.0) mm long; petioles slightly winged, (15)20–41(49) mm long. Middle cauline leaves sessile, (21.5)23.7–47.9(59.0) mm long, (9)10–20(22) mm wide, oblong-linear, pinnatipartite or pinnatifid, with laciniate margins and (6)7–24(31) laciniae, which are (1.7)2.3–4.0(5.0) mm long. Upper cauline leaves sessile, (18)19–28(33) mm long, (2.0)2.7–8.0(9.0) mm wide, linear with laciniate margins and (3)4–9(11) laciniae, which are (1.2)1.6–3.0 mm long. Involucre (10.0)12.7–19.3(21.0) mm wide, with pluriseriate, triangular (external) to linear (internal), green phyllaries, (4.0)4.4–5.4(7.0) mm long, (0.9)1.0–1.5(1.9) mm wide, with brownish scarious margins. Receptacle glabrous, slightly convex. Capitula up to
<quantity id="4CF39B32653DFFB7F95343F7EA7CFD1A" box="[1316,1402,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="30" unit="mm" value="53.0">53 mm</quantity>
in diameter. Ligulate florets up to 31, white, with tridentate apex, (8.5)10.0–17.6(20.0) mm long, 2.5–3.8(4.8) mm wide. Disk florets numerous, yellow, up to
<quantity id="4CF39B32653DFFB7FEB14240EC0FFCC9" box="[710,777,770,796]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="30" unit="mm" value="3.0">3 mm</quantity>
long. Ray cypselae up to
<quantity id="4CF39B32653DFFB7F84E4240EB79FCC9" box="[1081,1151,770,796]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="30" unit="mm" value="3.0">3 mm</quantity>
, brown, with 10 whitish ribs and a tubulose crown-shaped pappus. Disk cypselae similar to ray cypselae, lacking pappus, (1.7)1.9–2.3(2.7) mm long,
<quantity id="4CF39B32653DFFB7FDAB4212ED80FCBF" box="[476,646,848,874]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.4" metricValueMin="7.0" pageId="3" pageNumber="30" unit="mm" value="0.72" valueMax="0.74" valueMin="0.7">0.70–0.74 mm</quantity>
wide,
<quantity id="4CF39B32653DFFB7FEA34212EC7BFCBF" box="[724,893,848,874]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.1" metricValueMin="5.5" pageId="3" pageNumber="30" unit="mm" value="0.58" valueMax="0.61" valueMin="0.55">0.55–0.61 mm</quantity>
thick.
</paragraph>
<caption id="DF74665F653DFFB7FCE047A4EDFBF8E2" ID-DOI="http://doi.org/10.5281/zenodo.5065880" ID-Zenodo-Dep="5065880" httpUri="https://zenodo.org/record/5065880/files/figure.png" pageId="3" pageNumber="30" startId="3.[151,246,1766,1787]" targetBox="[151,1436,932,1746]" targetPageId="3">
<paragraph id="8BB436D7653DFFB7FCE047A4EDFBF8E2" blockId="3.[151,1436,1766,1847]" pageId="3" pageNumber="30">
<emphasis id="B97FEAC5653DFFB7FCE047A4EE09F92E" bold="true" box="[151,271,1766,1787]" pageId="3" pageNumber="30">FIGURE 2.</emphasis>
<taxonomicName id="4C0B4D54653DFFB7FD6147A5ED1CF929" authority="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi, 2012" authorityName="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi" authorityYear="2012" box="[278,538,1767,1788]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="ligusticum" status="sp. nov.">
<emphasis id="B97FEAC5653DFFB7FD6147A5ED1CF929" box="[278,538,1767,1788]" italics="true" pageId="3" pageNumber="30">Leucanthemum ligusticum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57BE653DFFB7FE5747A5ED6BF929" box="[544,621,1767,1788]" pageId="3" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
Variability of basal leaves in individuals from Ponte di Lagoscuro (A), Rocche di Valletti (B), Rocchetta di Vara (C). Variability of cauline leaves, upper ones (sx) to lower ones (dx), from Ponte di Lagoscuro (D), Rocche di Valletti (E), Rocchetta di Vara (F). Scale bars: 10 cm.
</paragraph>
</caption>
<paragraph id="8BB436D7653DFFB0FCB14627ECDFFEF3" blockId="3.[151,1436,1893,2036]" lastBlockId="4.[151,1437,152,564]" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
<materialsCitation id="3B633C8A653DFFB0FCB14627ECD3FEF3" collectingDate="2008-05-16" collectingDateMax="2010-07-21" collectingDateMin="2008-05-16" collectorName="di Vara &amp; La Spezia &amp; D. Marchetti &amp; M. Melai &amp; R. Bernardello &amp; G. Trombetti &amp; di Valletti &amp; di Lagoscuro" country="Italy" county="Rocchetta di Vara" elevation="220" lastPageId="4" lastPageNumber="31" location="Rocchetta di Vara" municipality="La Spezia" pageId="3" pageNumber="30" specimenCount="2" stateProvince="Liguria" typeStatus="paratype">
<emphasis id="B97FEAC5653DFFB7FCB14627ED59F8AA" bold="true" box="[198,607,1893,1919]" pageId="3" pageNumber="30">Additional specimens examined</emphasis>
(
<typeStatus id="54B08875653DFFB7FE024624EDEFF855" box="[629,745,1894,1920]" pageId="3" pageNumber="30" type="paratype">paratypes</typeStatus>
):
<emphasis id="B97FEAC5653DFFB7FE8C4624EC13F8AA" box="[763,789,1894,1919]" italics="true" pageId="3" pageNumber="30">—</emphasis>
<collectingCountry id="F31C7647653DFFB7FF624624EC61F855" box="[789,871,1894,1920]" name="Italy" pageId="3" pageNumber="30">ITALY</collectingCountry>
,
<emphasis id="B97FEAC5653DFFB7FF0E4627ECD9F8AA" bold="true" box="[889,991,1893,1919]" pageId="3" pageNumber="30">
<collectingRegion id="49CFF835653DFFB7FF0E4627ECDCF8AA" box="[889,986,1893,1919]" country="Italy" name="Liguria" pageId="3" pageNumber="30">Liguria</collectingRegion>
.
</emphasis>
<collectingCounty id="62D54E5B653DFFB7FF9B4624EBCCF855" box="[1004,1226,1894,1920]" pageId="3" pageNumber="30">
Rocchetta
<collectorName id="26FE5301653DFFB7F8074624EBCCF855" box="[1136,1226,1894,1920]" pageId="3" pageNumber="30">di Vara</collectorName>
</collectingCounty>
(province of
<collectorName id="26FE5301653DFFB7F90A4624EFECF873" pageId="3" pageNumber="30">
<collectingMunicipality id="6BD0ACAD653DFFB7F90A4624EFECF873" pageId="3" pageNumber="30">La Spezia</collectingMunicipality>
</collectorName>
), on siliceous substrate (&quot;diaspro&quot;),
<elevation id="0026D1E4653DFFB7FEE546CEEC50F873" box="[658,854,1932,1958]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">
<quantity id="4CF39B32653DFFB7FEE546CEEC10F873" box="[658,790,1932,1958]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">200–240 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB51017653DFFB7FF1346CEEB05F873" box="[868,1027,1932,1958]" pageId="3" pageNumber="30" value="2008-05-16">
<collectingDate id="EFF1E9FF653DFFB7FF1346CEEB05F873" box="[868,1027,1932,1958]" pageId="3" pageNumber="30" value="2008-05-16">16 May 2008</collectingDate>
</date>
,
<emphasis id="B97FEAC5653DFFB7F86746CFEBD1F873" box="[1040,1239,1933,1958]" italics="true" pageId="3" pageNumber="30">
<collectorName id="26FE5301653DFFB7F86746CFEBA0F873" box="[1040,1190,1933,1958]" pageId="3" pageNumber="30">D. Marchetti</collectorName>
s.n.
</emphasis>
(PI!);
<location id="8ED4600C653DFFB7F95146CEEFF6F818" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A:8ED4600C653DFFB7F95146CEEFF6F818" country="Italy" county="Rocchetta di Vara" municipality="La Spezia" name="Rocchetta di Vara" pageId="3" pageNumber="30" stateProvince="Liguria">
Rocchetta
<collectorName id="26FE5301653DFFB7FCE046F1EFF6F818" box="[151,240,1971,1997]" pageId="3" pageNumber="30">di Vara</collectorName>
</location>
(province of
<collectorName id="26FE5301653DFFB7FDEA46F1ED1BF818" box="[413,541,1971,1997]" pageId="3" pageNumber="30">La Spezia</collectorName>
), on siliceous substrate (&quot;diaspro&quot;),
<elevation id="0026D1E4653DFFB7FF9046F1EBB1F818" box="[999,1207,1971,1997]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">
<quantity id="4CF39B32653DFFB7FF9046F1EB77F818" box="[999,1137,1971,1997]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">200–240 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB51017653DFFB7F8B046F1EA77F818" box="[1223,1393,1971,1997]" pageId="3" pageNumber="30" value="2008-05-17">
<collectingDate id="EFF1E9FF653DFFB7F8B046F1EA77F818" box="[1223,1393,1971,1997]" pageId="3" pageNumber="30" value="2008-05-17">17 May 2008</collectingDate>
</date>
,
<emphasis id="B97FEAC5653DFFB7F90946F1EE3FF826" italics="true" pageId="3" pageNumber="30">
<collectorName id="26FE5301653DFFB7F90946F1EE0FF826" pageId="3" pageNumber="30">D. Marchetti</collectorName>
s.n.
</emphasis>
(PI!);
<location id="8ED4600C653DFFB7FDFF4698ED5EF821" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A:8ED4600C653DFFB7FDFF4698ED5EF821" box="[392,600,2010,2036]" country="Italy" county="Rocchetta di Vara" municipality="La Spezia" name="Rocchetta di Vara" pageId="3" pageNumber="30" stateProvince="Liguria">
Rocchetta
<collectorName id="26FE5301653DFFB7FE724698ED5EF821" box="[517,600,2010,2036]" pageId="3" pageNumber="30">di Vara</collectorName>
</location>
(province of
<collectorName id="26FE5301653DFFB7FE8F4698EC77F821" box="[760,881,2010,2036]" pageId="3" pageNumber="30">La Spezia</collectorName>
), on siliceous substrate (&quot;diaspro&quot;),
<elevation id="0026D1E4653DFFB0F96F4698EFD7FF67" lastPageId="4" lastPageNumber="31" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">
<quantity id="4CF39B32653DFFB7F96F4698EA9AF821" box="[1304,1436,2010,2036]" metricMagnitude="2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="30" unit="m" value="220.0" valueMax="240.0" valueMin="200.0">200–240 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB51017653AFFB0FC9641DAEE81FF67" box="[225,391,152,178]" pageId="4" pageNumber="31" value="2010-06-23">
<collectingDate id="EFF1E9FF653AFFB0FC9641DAEE81FF67" box="[225,391,152,178]" pageId="4" pageNumber="31" value="2010-06-23">23 June 2010</collectingDate>
</date>
,
<emphasis id="B97FEAC5653AFFB0FDE241DAEB6AFF64" box="[405,1132,152,177]" italics="true" pageId="4" pageNumber="31">
<collectorName id="26FE5301653AFFB0FDE241DAED0FFF64" box="[405,521,152,177]" pageId="4" pageNumber="31">M. Melai</collectorName>
,
<collectorName id="26FE5301653AFFB0FE6141DAEDB0FF64" box="[534,694,152,177]" pageId="4" pageNumber="31">D. Marchetti</collectorName>
,
<collectorName id="26FE5301653AFFB0FEB341DAEC7FFF64" box="[708,889,152,177]" pageId="4" pageNumber="31">R. Bernardello</collectorName>
et
<collectorName id="26FE5301653AFFB0FFD741DAEB3EFF64" box="[928,1080,152,177]" pageId="4" pageNumber="31">G. Trombetti</collectorName>
s.n.
</emphasis>
(PI!);
<location id="8ED4600C653AFFB0F8B741DAEA9AFF67" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A:8ED4600C653AFFB0F8B741DAEA9AFF67" box="[1216,1436,152,178]" country="Italy" county="Rocchetta di Vara" municipality="La Spezia" name="Rocche di Valletti" pageId="4" pageNumber="31" stateProvince="Liguria">
Rocche
<collectorName id="26FE5301653AFFB0F95441DAEA9AFF67" box="[1315,1436,152,178]" pageId="4" pageNumber="31">di Valletti</collectorName>
</location>
(province of
<collectorName id="26FE5301653AFFB0FD5941FCEEA1FF0D" box="[302,423,190,216]" pageId="4" pageNumber="31">La Spezia</collectorName>
), on siliceous substrate (&quot;diaspro&quot;),
<elevation id="0026D1E4653AFFB0FF3941FCECD1FF0D" box="[846,983,190,216]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="31" unit="m" value="800.0">
<quantity id="4CF39B32653AFFB0FF3941FCEC9EFF0D" box="[846,920,190,216]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="31" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB51017653AFFB0FF9241FCEB79FF0D" box="[997,1151,190,216]" pageId="4" pageNumber="31" value="2010-07-21">
<collectingDate id="EFF1E9FF653AFFB0FF9241FCEB79FF0D" box="[997,1151,190,216]" pageId="4" pageNumber="31" value="2010-07-21">21 July 2010</collectingDate>
</date>
,
<emphasis id="B97FEAC5653AFFB0F8FC41FDEE9CFF2B" italics="true" pageId="4" pageNumber="31">
<collectorName id="26FE5301653AFFB0F8FC41FDEBFFFF0D" box="[1163,1273,191,216]" pageId="4" pageNumber="31">M. Melai</collectorName>
,
<collectorName id="26FE5301653AFFB0F97241FDEA9AFF0D" box="[1285,1436,191,216]" pageId="4" pageNumber="31">D. Marchetti</collectorName>
et
<collectorName id="26FE5301653AFFB0FCC241A7EE60FF2B" box="[181,358,229,254]" pageId="4" pageNumber="31">R. Bernardello</collectorName>
s.n.
</emphasis>
(PI!);
<location id="8ED4600C653AFFB0FD9C41A7ED92FF2A" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A:8ED4600C653AFFB0FD9C41A7ED92FF2A" box="[491,660,229,255]" country="Italy" county="Rocchetta di Vara" municipality="La Spezia" name="Val Graveglia" pageId="4" pageNumber="31" stateProvince="Liguria">Val Graveglia</location>
, sotto Nascio (province of Genova),
<location id="8ED4600C653AFFB0F82241A7EA44FF2A" LSID="urn:lsid:plazi:treatment:03A287C1653FFFB0FCE047B0EE16FA0A:8ED4600C653AFFB0F82241A7EA44FF2A" box="[1109,1346,229,255]" country="Italy" county="Rocchetta di Vara" municipality="La Spezia" name="Ponte di Lagoscuro" pageId="4" pageNumber="31" stateProvince="Liguria">
Ponte
<collectorName id="26FE5301653AFFB0F8D641A7EA44FF2A" box="[1185,1346,229,255]" pageId="4" pageNumber="31">di Lagoscuro</collectorName>
</location>
,
<elevation id="0026D1E4653AFFB0F93841A7EFC8FEF3" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="31" unit="m" value="250.0">
<quantity id="4CF39B32653AFFB0F93841A7EA9AFF2A" box="[1359,1436,229,255]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="31" unit="m" value="250.0">250 m</quantity>
a.s.l.
</elevation>
,
<date id="FFB51017653AFFB0FCAB404EEE70FEF3" box="[220,374,268,294]" pageId="4" pageNumber="31" value="2010-07-21">
<collectingDate id="EFF1E9FF653AFFB0FCAB404EEE70FEF3" box="[220,374,268,294]" pageId="4" pageNumber="31" value="2010-07-21">21 July 2010</collectingDate>
</date>
,
<emphasis id="B97FEAC5653AFFB0FDF5404EEC95FEF0" box="[386,915,268,293]" italics="true" pageId="4" pageNumber="31">
<collectorName id="26FE5301653AFFB0FDF5404EEEF6FEF0" box="[386,496,268,293]" pageId="4" pageNumber="31">M. Melai</collectorName>
,
<collectorName id="26FE5301653AFFB0FD8B404EED94FEF0" box="[508,658,268,293]" pageId="4" pageNumber="31">D. Marchetti</collectorName>
et
<collectorName id="26FE5301653AFFB0FEC1404EEC65FEF0" box="[694,867,268,293]" pageId="4" pageNumber="31">R. Bernardello</collectorName>
s.n.
</emphasis>
(PI!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311655C653AFFB0FCB14070EC79FE4F" pageId="4" pageNumber="31" type="biology_ecology">
<paragraph id="8BB436D7653AFFB0FCB14070EC79FE4F" blockId="4.[151,1437,152,564]" pageId="4" pageNumber="31">
<emphasis id="B97FEAC5653AFFB0FCB14070EEFBFE99" bold="true" box="[198,509,306,332]" pageId="4" pageNumber="31">Ecology and Phenology:</emphasis>
—
<taxonomicName id="4C0B4D54653AFFB0FE6D4070EC55FE9E" authorityName="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi" authorityYear="2012" box="[538,851,306,331]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="31" phylum="Tracheophyta" rank="species" species="ligusticum">
<emphasis id="B97FEAC5653AFFB0FE6D4070EC55FE9E" box="[538,851,306,331]" italics="true" pageId="4" pageNumber="31">Leucanthemum ligusticum</emphasis>
</taxonomicName>
grows in open rocky places (
<quantity id="4CF39B32653AFFB0F8B04070EA62FE99" box="[1223,1380,306,332]" metricMagnitude="2" metricUnit="m" metricValue="6.3" metricValueMax="10.6" metricValueMin="2.0" pageId="4" pageNumber="31" unit="m" value="630.0" valueMax="1060.0" valueMin="200.0">200–1060 m</quantity>
), on siliceous substrates and deep-oceanic siliceous sedimentary rocks (&quot;diaspri&quot;). It is a hemicryptophyte, flowering from the beginning of May until the first half of July.
</paragraph>
</subSubSection>
<subSubSection id="C311655C653AFFB0FCB140E4EE16FA0A" pageId="4" pageNumber="31" type="distribution">
<paragraph id="8BB436D7653AFFB0FCB140E4EED2FDE1" blockId="4.[151,1437,152,564]" pageId="4" pageNumber="31">
<emphasis id="B97FEAC5653AFFB0FCB140E4EE6FFE15" bold="true" box="[198,361,422,448]" pageId="4" pageNumber="31">Distribution:</emphasis>
—
<taxonomicName id="4C0B4D54653AFFB0FDF140E5EDC4FE15" authorityName="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi" authorityYear="2012" box="[390,706,423,448]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="31" phylum="Tracheophyta" rank="species" species="ligusticum">
<emphasis id="B97FEAC5653AFFB0FDF140E5EDC4FE15" box="[390,706,423,448]" italics="true" pageId="4" pageNumber="31">Leucanthemum ligusticum</emphasis>
</taxonomicName>
is currently known from few localities in eastern
<collectingRegion id="49CFF835653AFFB0F93240E4EA9DFE15" box="[1349,1435,422,448]" country="Italy" name="Liguria" pageId="4" pageNumber="31">Liguria</collectingRegion>
(provinces of Genova and La Spezia;
<figureCitation id="13302A52653AFFB0FE1F408FEDB4FE32" box="[616,690,461,487]" captionStart="FIGURE 3" captionStartId="4.[151,244,1242,1263]" captionTargetBox="[166,1423,609,1225]" captionTargetId="figure-467@4.[151,1436,609,1252]" captionTargetPageId="4" captionText="FIGURE 3. Leucanthemum ligusticum sp. nov. Known geographical distribution in eastern Liguria (northwestern Italy Italy)." figureDoi="http://doi.org/10.5281/zenodo.5065882" httpUri="https://zenodo.org/record/5065882/files/figure.png" pageId="4" pageNumber="31">Fig. 3</figureCitation>
). Besides the three studied populations, the new species was observed near Bargone (Genova province), Mounts Roccagrande-Tregìn,
<quantity id="4CF39B32653AFFB0F85740B6EBACFDDB" box="[1056,1194,500,526]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="8.5" metricValueMin="4.0" pageId="4" pageNumber="31" unit="m" value="625.0" valueMax="850.0" valueMin="400.0">400–850 m</quantity>
and near the top of Mount Verruga, at
<quantity id="4CF39B32653AFFB0FD034358EED6FDE1" box="[372,464,538,564]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="31" unit="m" value="1060.0">1060 m</quantity>
.
</paragraph>
<caption id="DF74665F653AFFB0FCE04598EA4BFB25" ID-DOI="http://doi.org/10.5281/zenodo.5065882" ID-Zenodo-Dep="5065882" box="[151,1357,1242,1264]" httpUri="https://zenodo.org/record/5065882/files/figure.png" pageId="4" pageNumber="31" startId="4.[151,244,1242,1263]" targetBox="[166,1423,609,1225]" targetPageId="4">
<paragraph id="8BB436D7653AFFB0FCE04598EA4BFB25" blockId="4.[151,1357,1242,1264]" box="[151,1357,1242,1264]" pageId="4" pageNumber="31">
<emphasis id="B97FEAC5653AFFB0FCE04598EE0AFB3A" bold="true" box="[151,268,1242,1263]" pageId="4" pageNumber="31">FIGURE 3.</emphasis>
<taxonomicName id="4C0B4D54653AFFB0FD654599ED08FB25" authority="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi, 2012" authorityName="Melai &amp; Marchetti &amp; Bernardello &amp; Peruzzi" authorityYear="2012" box="[274,526,1243,1264]" class="Magnoliopsida" family="Asteraceae" genus="Leucanthemum" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="31" phylum="Tracheophyta" rank="species" species="ligusticum" status="sp. nov.">
<emphasis id="B97FEAC5653AFFB0FD654599ED08FB25" box="[274,526,1243,1264]" italics="true" pageId="4" pageNumber="31">Leucanthemum ligusticum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57BE653AFFB0FE634599ED59FB25" box="[532,607,1243,1264]" pageId="4" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
Known geographical distribution in eastern Liguria (northwestern Italy Italy).
</paragraph>
</caption>
<paragraph id="8BB436D7653AFFB0FCB1445DEE16FA0A" blockId="4.[151,1436,1311,1503]" pageId="4" pageNumber="31">
<emphasis id="B97FEAC5653AFFB0FCB1445DEE52FAEC" bold="true" box="[198,340,1311,1337]" pageId="4" pageNumber="31">Karyology:</emphasis>
—The new species is diploid, with 2n = 18 chromosomes, at all three localities studied (
<collectingRegion id="49CFF835653AFFB0FCE8440BEFFAFAB6" box="[159,252,1353,1379]" country="Italy" name="Liguria" pageId="4" pageNumber="31">Liguria</collectingRegion>
;
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). The karyotype formula is 2n = 2x = 18 =
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at Ponte di Lagoscuro, 2n = 2x = 18 =
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at Rocche di Valletti and 2n = 2x = 18 =
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at Rocchetta di Vara (
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). Chromosome size ranges from
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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