treatments-xml/data/03/A2/87/03A287AEFF88FFC1FC9AFE7D8227FE80.xml

<document id="9E4A593724504A90D9731A110BEFCD40" ID-DOI="10.6620/ZS.2022.61-24" ID-ISSN="1810-522X" ID-PMC="PMC9537050" ID-PubMed="36330030" ID-Zenodo-Dep="12827680" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721933725735" checkinUser="felipe" docAuthor="Lin, Hsiang-Yun, Huang, Jun-Xuan, Liu, Hsi-Hsuan &amp; Chang, Chih-Han" docDate="2022" docId="03A287AEFF88FFC1FC9AFE7D8227FE80" docLanguage="en" docName="ZoolStud.61.24.1-13.pdf" docOrigin="Zoological Studies 61 (24)" docSource="http://dx.doi.org/10.5281/zenodo.12827270" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Garypus sanasai Lin, Huang &amp; Chang 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FF9BFFD6FF8DFFC9FFA8FFE18341FFD5" masterDocTitle="Tubuca paradussumieri" masterLastPageNumber="13" masterPageNumber="1" pageNumber="6" updateTime="1721938663393" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2493C42553E4502104208F838D7B811D">Tubuca paradussumieri</mods:title>
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<mods:namePart id="2813112C267E0D41990CA03B49380B5F">Lin, Hsiang-Yun</mods:namePart>
<mods:affiliation id="43CC61146062CAA6DBDE8B7474FDB333">Department of Life Science, National Taiwan Normal University, No. 88, Sec. 4, Tingzhou Rd., Taipei City 116059, Taiwan.</mods:affiliation>
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<mods:namePart id="2B349A0E1599BA1757597A9EAB975BCE">Huang, Jun-Xuan</mods:namePart>
<mods:affiliation id="96BD24ED37A5E171BA3C0B1077D2D027">Department of Entomology, National Taiwan University, No. 1, Sec. 4, Roosevelt Rd., Taipei City 106216, Taiwan.</mods:affiliation>
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<mods:namePart id="BC23C6AE955E61B4039889C6099E4E97">Liu, Hsi-Hsuan</mods:namePart>
<mods:affiliation id="8A4F229667EF25341DD8425A69DF95DB">Sacred Heart High School for Girls, No. 263, Sec. 1, Longmi Rd., New Taipei City 24931, Taiwan. E-mail: princekinliu @ gmail. com (Liu)</mods:affiliation>
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<mods:namePart id="1FF15954924204F3ACDE541B1A8DE606">Chang, Chih-Han</mods:namePart>
<mods:affiliation id="DD2B428DE270244761DB9357BBBF71B2">Department of Life Science, National Taiwan University, No. 1, Sec. 4, Roosevelt Rd., Taipei City 106216, Taiwan. &amp; Institute of Ecology and Evolutionary Biology, National Taiwan University, No. 1, Sec. 4, Roosevelt Rd., Taipei City 106216, Taiwan</mods:affiliation>
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<taxonomicName id="4C0B4D3BFF88FFCCFC9AFE7D8650FE63" authority="Lin, Huang &amp; Chang" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[818,1297,412,438]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sanasai" status="sp. nov.">
<emphasis id="B97FEAAAFF88FFCCFC9AFE7D8749FE63" bold="true" box="[818,1032,412,438]" italics="true" pageId="5" pageNumber="6">Garypus sanasai</emphasis>
Lin, Huang &amp; Chang
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, 
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<paragraph id="8BB436B8FF88FFCCFC4AFE5C87E8FDDF" blockId="5.[846,1380,445,522]" pageId="5" pageNumber="6">
(
<figureCitation id="13302A3DFF88FFCCFC42FE5C8706FE00" box="[1002,1095,445,469]" captionStart="Fig" captionStartId="3.[136,167,1114,1133]" captionTargetBox="[136,1413,682,1087]" captionTargetId="figure-299@3.[136,1413,682,1087]" captionTargetPageId="3" captionText="Fig. 1. Photos of the holotypes of the two new species of Garypus. The photos were taken when the individuals were still alive. (A) Garypus wilsoni sp. nov. Lin &amp; Chang (Photo by Bo-Xin Guo and Yi-Ting Chung with permission). (B) Garypus sanasai sp. nov. Lin, Huang &amp; Chang." figureDoi="http://doi.org/10.5281/zenodo.12827682" httpUri="https://zenodo.org/record/12827682/files/figure.png" pageId="5" pageNumber="6">Figs. 1B</figureCitation>
, 
<figureCitation id="13302A3DFF88FFCCFBFAFE5C87D5FE00" box="[1106,1172,445,469]" captionStart="Fig" captionStartId="5.[137,168,1896,1915]" captionTargetBox="[136,732,976,1868]" captionTargetId="figure-656@5.[136,737,966,1868]" captionTargetPageId="5" captionText="Fig. 4. Dorsal view of carapace and the comparison of color patterns of the two species. (A, B) Garypus wilsoni Lin &amp; Chang sp. nov.; (C, D) Garypus sanasai Lin, Huang &amp; Chang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12827686" httpUri="https://zenodo.org/record/12827686/files/figure.png" pageId="5" pageNumber="6">4C, D</figureCitation>
, 
<figureCitation id="13302A3DFF88FFCCFB37FE5C87EFFE00" box="[1183,1198,445,469]" captionStart="Fig" captionStartId="6.[174,205,1954,1973]" captionTargetBox="[223,1402,1259,1929]" captionTargetId="figure-6@6.[203,1421,1212,1957]" captionTargetPageId="6" captionText="Fig. 5. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view; (B) ventral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827688" httpUri="https://zenodo.org/record/12827688/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
, 
<figureCitation id="13302A3DFF88FFCCFB12FE5C8789FE00" box="[1210,1224,445,469]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="5" pageNumber="6">6</figureCitation>
) 
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BB436B8FF88FFCCFCC1FDD087ACFD7C" blockId="5.[813,1414,560,1961]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAAFF88FFCCFCC1FDD08727FD9D" box="[873,1126,560,584]" italics="true" pageId="5" pageNumber="6">M a t e r i a l s e x a m i n e d</emphasis>
: H o l o t y p e: ò; Lüdao Township, 
<collectingRegion id="49CFF85AFF88FFCCFC1AFDB1872CFDBC" box="[946,1133,592,617]" country="Taiwan" name="Taitung" pageId="5" pageNumber="6">Taitung County</collectingRegion>
, 
<collectingCountry id="F31C7628FF88FFCCFBD4FDB18794FDBD" box="[1148,1237,592,616]" name="Taiwan" pageId="5" pageNumber="6">Taiwan</collectingCountry>
; 
<geoCoordinate id="EE3F507FFF88FFCCFB4DFDB086C5FDBC" box="[1253,1412,593,617]" degrees="22" direction="north" minutes="38" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="08.1" value="22.635584">22°38'08.1&quot;N</geoCoordinate>
<geoCoordinate id="EE3F507FFF88FFCCFC85FD918090FD5D" box="[813,977,624,648]" degrees="121" direction="east" minutes="29" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="55.8" value="121.49883">121°29'55.8&quot;E</geoCoordinate>
; coll. Jun-Xuan Huang; 
<date id="FFB51078FF88FFCCFB40FD9086C0FD5C" box="[1256,1409,625,649]" pageId="5" pageNumber="6" value="2020-09-01">01 Sep. 2020</date>
; GenBank: OL514113; NTUM-PSC-0002.
</paragraph>
<paragraph id="8BB436B8FF88FFCCFCC1FD51878CFC5D" blockId="5.[813,1414,560,1961]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAAFF88FFCCFCC1FD51809AFD1D" box="[873,987,688,712]" italics="true" pageId="5" pageNumber="6">Etymology</emphasis>
: The species is named after 
<taxonomicName id="4C0B4D3BFF88FFCCFAB2FD518636FD1D" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[1306,1399,688,712]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sanasai">“sanasai</taxonomicName>
”, a legendary name of the 
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locality. In the legends of many Taiwanese indigenous tribes, which are part of the Austronesian peoples, their ancestors arrived 
<collectingCountry id="F31C7628FF88FFCCFA9DFCF186C4FCFD" box="[1333,1413,784,808]" name="Taiwan" pageId="5" pageNumber="6">Taiwan</collectingCountry>
through a way station called 
<taxonomicName id="4C0B4D3BFF88FFCCFBD4FCD1879FFC9D" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[1148,1246,816,840]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sanasai">“sanasai</taxonomicName>
”, which, some tribes believe, is Green Island (Lüdao Township). The specific epithet is a noun in apposition.
</paragraph>
<paragraph id="8BB436B8FF88FFCCFCC1FC7080E0F9FD" blockId="5.[813,1414,560,1961]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAAFF88FFCCFCC1FC70809CFC7D" box="[873,989,913,936]" italics="true" pageId="5" pageNumber="6">Diagnosis</emphasis>
: The color pattern on carapace and tergites of the new species is most similar to that of 
<taxonomicName id="4C0B4D3BFF88FFCCFC85FC308720FC3C" authority="Harvey." authorityName="Harvey." box="[813,1121,976,1001]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="B97FEAAAFF88FFCCFC85FC3080BAFC3D" box="[813,1019,976,1000]" italics="true" pageId="5" pageNumber="6">G. schwendingeri</emphasis>
Harvey.
</taxonomicName>
The new species can be differentiated from 
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<emphasis id="B97FEAAAFF88FFCCFBABFC108782FBDD" box="[1027,1219,1008,1032]" italics="true" pageId="5" pageNumber="6">G. schwendingeri</emphasis>
</taxonomicName>
by the pattern of trichobothria on chela (
<emphasis id="B97FEAAAFF88FFCCFB8CFBF0877CFBFD" box="[1060,1085,1041,1064]" italics="true" pageId="5" pageNumber="6">ist</emphasis>
and 
<emphasis id="B97FEAAAFF88FFCCFBD8FBF187CEFBFD" box="[1136,1167,1040,1064]" italics="true" pageId="5" pageNumber="6">isb</emphasis>
on the same horizontal plane in the new species, but not in 
<taxonomicName id="4C0B4D3BFF88FFCCFB10FBD08637FB9D" box="[1208,1398,1072,1096]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="B97FEAAAFF88FFCCFB10FBD08637FB9D" box="[1208,1398,1072,1096]" italics="true" pageId="5" pageNumber="6">G. schwendingeri</emphasis>
</taxonomicName>
). The species shares some common characters with 
<taxonomicName id="4C0B4D3BFF88FFCCFAC3FBB080B6FB5C" authority="Harvey" authorityName="Harvey" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="necopinus">
<emphasis id="B97FEAAAFF88FFCCFAC3FBB080DCFB5D" italics="true" pageId="5" pageNumber="6">G. necopinus</emphasis>
Harvey
</taxonomicName>
and 
<taxonomicName id="4C0B4D3BFF88FFCCFB9AFB908606FB5D" authority="Mahnert" authorityName="Mahnert" authorityYear="1982" box="[1074,1351,1136,1160]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="marmoratus">
<emphasis id="B97FEAAAFF88FFCCFB9AFB90879CFB5D" box="[1074,1245,1137,1160]" italics="true" pageId="5" pageNumber="6">G. marmoratus</emphasis>
Mahnert
</taxonomicName>
, with trichobothrium 
<emphasis id="B97FEAAAFF88FFCCFC7FFB7080A8FB7D" box="[983,1001,1169,1192]" italics="true" pageId="5" pageNumber="6">st</emphasis>
midway between 
<emphasis id="B97FEAAAFF88FFCCFB1CFB71878DFB7D" box="[1204,1228,1168,1192]" italics="true" pageId="5" pageNumber="6">sb</emphasis>
and 
<emphasis id="B97FEAAAFF88FFCCFAADFB70864DFB7D" box="[1285,1292,1169,1192]" italics="true" pageId="5" pageNumber="6">t</emphasis>
or slightly closer to 
<emphasis id="B97FEAAAFF88FFCCFC3DFB5180ECFB1D" box="[917,941,1200,1224]" italics="true" pageId="5" pageNumber="6">sb</emphasis>
, tooth rows not touching each other when fingers of palps are closed, tergites mostly pale yellow in color with some brown spots. The three species can be differentiated by the color pattern of carapace and the shape of galea and rallum. Compared to 
<taxonomicName id="4C0B4D3BFF88FFCCFAB8FAD0863EFA9D" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" box="[1296,1407,1328,1352]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B97FEAAAFF88FFCCFAB8FAD0863EFA9D" box="[1296,1407,1328,1352]" italics="true" pageId="5" pageNumber="6">G. wilsoni</emphasis>
</taxonomicName>
, the color is darker than 
<taxonomicName id="4C0B4D3BFF88FFCCFB87FAB087DFFABD" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" box="[1071,1182,1360,1384]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B97FEAAAFF88FFCCFB87FAB087DFFABD" box="[1071,1182,1360,1384]" italics="true" pageId="5" pageNumber="6">G. wilsoni</emphasis>
</taxonomicName>
and the color pattern of carapace is quite different on both lateral sides, also the color pattern of tergites, 
<taxonomicName id="4C0B4D3BFF88FFCCFBCAFA708790FA7D" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" box="[1122,1233,1424,1448]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B97FEAAAFF88FFCCFBCAFA708790FA7D" box="[1122,1233,1424,1448]" italics="true" pageId="5" pageNumber="6">G. wilsoni</emphasis>
</taxonomicName>
with less brown color, the setae of poseterior genital operculum, 
<taxonomicName id="4C0B4D3BFF88FFCCFAC3FA50803BFA3D" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B97FEAAAFF88FFCCFAC3FA50803BFA3D" italics="true" pageId="5" pageNumber="6">G. wilsoni</emphasis>
</taxonomicName>
is not in a row, 
<emphasis id="B97FEAAAFF88FFCCFB8BFA318779FA3D" box="[1059,1080,1488,1512]" italics="true" pageId="5" pageNumber="6">ib</emphasis>
is situated between 
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and 
<emphasis id="B97FEAAAFF88FFCCFACAFA3186C4FA3D" box="[1378,1413,1488,1512]" italics="true" pageId="5" pageNumber="6">esb</emphasis>
in 
<taxonomicName id="4C0B4D3BFF88FFCCFCE0FA1080FBF9DD" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[840,954,1521,1544]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sanasai">
<emphasis id="B97FEAAAFF88FFCCFCE0FA1080FBF9DD" box="[840,954,1521,1544]" italics="true" pageId="5" pageNumber="6">G. sanasai</emphasis>
</taxonomicName>
, 
<emphasis id="B97FEAAAFF88FFCCFC61FA11809FF9DD" box="[969,990,1520,1544]" italics="true" pageId="5" pageNumber="6">ib</emphasis>
and 
<emphasis id="B97FEAAAFF88FFCCFBBAFA118777F9DD" box="[1042,1078,1520,1544]" italics="true" pageId="5" pageNumber="6">esb</emphasis>
on the same horizontal pane in 
<taxonomicName id="4C0B4D3BFF88FFCCFC85F9F080DBF9FD" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" box="[813,922,1552,1576]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="B97FEAAAFF88FFCCFC85F9F080DBF9FD" box="[813,922,1552,1576]" italics="true" pageId="5" pageNumber="6">G. wilsoni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB436B8FF88FFCCFCC1F9D08664F93D" blockId="5.[813,1414,560,1961]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAAFF88FFCCFCC1F9D080A6F99D" box="[873,999,1585,1608]" italics="true" pageId="5" pageNumber="6">Description</emphasis>
: Color in ethanol (
<figureCitation id="13302A3DFF88FFCCFB19F9D187B3F99D" box="[1201,1266,1584,1608]" captionStart="Fig" captionStartId="6.[174,205,1954,1973]" captionTargetBox="[223,1402,1259,1929]" captionTargetId="figure-6@6.[203,1421,1212,1957]" captionTargetPageId="6" captionText="Fig. 5. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view; (B) ventral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827688" httpUri="https://zenodo.org/record/12827688/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
). Mostly pale yellow; pedipalp dark yellow with dark brown chela, anterior, posterolateral, and posteromedial carapace dark brown, legs pale white, tergites mostly light yellow with 4 brown spots at lateral and middle side, and with some brown color at the middle of tergites I‒V.
</paragraph>
<paragraph id="8BB436B8FF88FFCFFCC1F9118106FEE0" blockId="5.[813,1414,560,1961]" lastBlockId="6.[174,775,253,1205]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Chelicera: Hand with 5 setae, movable finger with 1 distal setae; 1 dorsal and 1 ventral lyrifissure on the basal of fixed finger; fixed finger with 3 teeth and 4 smaller teeth, movable finger with 1 apical tooth; galea present (
<figureCitation id="13302A3DFF88FFCCFC22F891809FF85D" box="[906,990,1904,1928]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="5" pageNumber="6">Fig. 6E</figureCitation>
), with 4 rami, 3 rami at distal and 1 at lateral; rallum (
<figureCitation id="13302A3DFF88FFCCFC7DF871876BF87D" box="[981,1066,1936,1960]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="5" pageNumber="6">Fig. 6D</figureCitation>
) with 3 blades, each blade with several small lateral spinules; serrula exterior with 33 blades, serrula interior with 21 blades.
</paragraph>
<caption id="DF746630FF88FFCCFF21F8898177F866" ID-DOI="http://doi.org/10.5281/zenodo.12827686" ID-Zenodo-Dep="12827686" httpUri="https://zenodo.org/record/12827686/files/figure.png" pageId="5" pageNumber="6" startId="5.[137,168,1896,1915]" targetBox="[136,732,976,1868]" targetPageId="5" targetType="figure">
<paragraph id="8BB436B8FF88FFCCFF21F8898177F866" blockId="5.[136,739,1896,1971]" pageId="5" pageNumber="6">
<emphasis id="B97FEAAAFF88FFCCFF21F8898383F8AE" bold="true" box="[137,194,1896,1915]" pageId="5" pageNumber="6">Fig. 4.</emphasis>
Dorsal view of carapace and the comparison of color patterns of the two species. (A, B) 
<taxonomicName id="4C0B4D3BFF88FFCCFEC5F8658133F842" authority="Lin &amp; Chang" authorityName="Lin &amp; Huang &amp; Liu &amp; Chang" authorityYear="2022" box="[365,626,1924,1943]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
<emphasis id="B97FEAAAFF88FFCCFEC5F86582BBF842" box="[365,506,1924,1943]" italics="true" pageId="5" pageNumber="6">Garypus wilsoni</emphasis>
Lin &amp; Chang
</taxonomicName>
<taxonomicNameLabel id="A24C57D1FF88FFCCFDD0F86581FFF842" box="[632,702,1924,1943]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
; (C, D) 
<taxonomicName id="4C0B4D3BFF88FFCCFF0CF84182AFF866" authority="Lin, Huang &amp; Chang" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[164,494,1952,1971]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sanasai" status="sp. nov.">
<emphasis id="B97FEAAAFF88FFCCFF0CF8418275F867" box="[164,308,1952,1971]" italics="true" pageId="5" pageNumber="6">Garypus sanasai</emphasis>
Lin, Huang &amp; Chang
</taxonomicName>
<taxonomicNameLabel id="A24C57D1FF88FFCCFE5BF8418177F866" box="[499,566,1952,1971]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB436B8FF8BFFCFFF42FEDC82A4FC80" blockId="6.[174,775,253,1205]" pageId="6" pageNumber="7">
Pedipalp (
<figureCitation id="13302A3DFF8BFFCFFEC8FEDC82FEFE80" box="[352,447,317,341]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="6" pageNumber="7">Fig. 6A</figureCitation>
): Granulated in whole chela including the pedicel; Femur slender, 4.15 times longer than broad, patella 3.19 longer than broad; 2 lyrifissure situated at basal part of patella; both fingers slightly curved in lateral view (
<figureCitation id="13302A3DFF8BFFCFFE6DFE5C815EFE00" box="[453,543,445,469]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="6" pageNumber="7">Fig. 6F</figureCitation>
), fixed finger more curved than movable; fixed finger with 8 trichobothria, movable finger with 4; 
<emphasis id="B97FEAAAFF8BFFCFFE07FE1C828FFDC0" box="[431,462,509,533]" italics="true" pageId="6" pageNumber="7">isb</emphasis>
, 
<emphasis id="B97FEAAAFF8BFFCFFE75FE1C82B3FDC0" box="[477,498,509,533]" italics="true" pageId="6" pageNumber="7">ib</emphasis>
, 
<emphasis id="B97FEAAAFF8BFFCFFDA9FE1C8164FDC0" box="[513,549,509,533]" italics="true" pageId="6" pageNumber="7">esb</emphasis>
, 
<emphasis id="B97FEAAAFF8BFFCFFD9CFE1C810CFDC0" box="[564,589,509,533]" italics="true" pageId="6" pageNumber="7">eb</emphasis>
in a row, 
<emphasis id="B97FEAAAFF8BFFCFFD16FE1F8196FDC0" box="[702,727,510,533]" italics="true" pageId="6" pageNumber="7">ist</emphasis>
and 
<emphasis id="B97FEAAAFF8BFFCFFF06FDFC8382FDE0" box="[174,195,541,565]" italics="true" pageId="6" pageNumber="7">ib</emphasis>
in a different row, 
<emphasis id="B97FEAAAFF8BFFCFFE30FDFF82EBFDE0" box="[408,426,542,565]" italics="true" pageId="6" pageNumber="7">st</emphasis>
much closer to 
<emphasis id="B97FEAAAFF8BFFCFFDF5FDFC8134FDE0" box="[605,629,541,565]" italics="true" pageId="6" pageNumber="7">sb</emphasis>
than to 
<emphasis id="B97FEAAAFF8BFFCFFD7AFDFF8198FDE0" box="[722,729,542,565]" italics="true" pageId="6" pageNumber="7">t</emphasis>
, 
<emphasis id="B97FEAAAFF8BFFCFFD41FDFF8046FDE0" box="[745,775,542,565]" italics="true" pageId="6" pageNumber="7">est</emphasis>
closer to 
<emphasis id="B97FEAAAFF8BFFCFFEBCFDDF8266FD80" box="[276,295,574,597]" italics="true" pageId="6" pageNumber="7">et</emphasis>
than 
<emphasis id="B97FEAAAFF8BFFCFFECFFDDC82C7FD80" box="[359,390,573,597]" italics="true" pageId="6" pageNumber="7">isb</emphasis>
, 
<emphasis id="B97FEAAAFF8BFFCFFE3DFDDF82EFFD80" box="[405,430,574,597]" italics="true" pageId="6" pageNumber="7">ist</emphasis>
and 
<emphasis id="B97FEAAAFF8BFFCFFE4EFDDC8144FD80" box="[486,517,573,597]" italics="true" pageId="6" pageNumber="7">isb</emphasis>
on the same horizontal plane, 
<emphasis id="B97FEAAAFF8BFFCFFF57FDBC8254FDA0" box="[255,277,605,629]" italics="true" pageId="6" pageNumber="7">ib</emphasis>
between 
<emphasis id="B97FEAAAFF8BFFCFFE22FDBC82EAFDA0" box="[394,427,605,629]" italics="true" pageId="6" pageNumber="7">isb</emphasis>
and 
<emphasis id="B97FEAAAFF8BFFCFFE42FDBC8151FDA0" box="[490,528,605,629]" italics="true" pageId="6" pageNumber="7">esb</emphasis>
; 
<emphasis id="B97FEAAAFF8BFFCFFD8CFDBF8179FDA0" box="[548,568,606,629]" italics="true" pageId="6" pageNumber="7">et</emphasis>
without adjacent patch of microsetae, with 18 microsetae near 
<emphasis id="B97FEAAAFF8BFFCFFD19FD9F81F9FD40" box="[689,696,638,661]" italics="true" pageId="6" pageNumber="7">t</emphasis>
, and 7 microsetae between 
<emphasis id="B97FEAAAFF8BFFCFFE24FD7C82E5FD60" box="[396,420,669,693]" italics="true" pageId="6" pageNumber="7">sb</emphasis>
and 
<emphasis id="B97FEAAAFF8BFFCFFE74FD7C82A8FD60" box="[476,489,669,693]" italics="true" pageId="6" pageNumber="7">b</emphasis>
; venom apparatus present on both fingers; gaps present while fingers closed; Fixed finger with 116 straight round teeth, movable finger with 89 teeth, 37 round and small teeth at the basal part, 52 retrose triangular teeth at the distal part, a gap present between 37th and 38th tooth.
</paragraph>
<paragraph id="8BB436B8FF8BFFCFFF42FCBC81E8FC20" blockId="6.[174,775,253,1205]" pageId="6" pageNumber="7">
Cephalothorax: Triangular carapace (
<figureCitation id="13302A3DFF8BFFCFFD0DFCBC8387FC43" captionStart="Fig" captionStartId="5.[137,168,1896,1915]" captionTargetBox="[136,732,976,1868]" captionTargetId="figure-656@5.[136,737,966,1868]" captionTargetPageId="5" captionText="Fig. 4. Dorsal view of carapace and the comparison of color patterns of the two species. (A, B) Garypus wilsoni Lin &amp; Chang sp. nov.; (C, D) Garypus sanasai Lin, Huang &amp; Chang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12827686" httpUri="https://zenodo.org/record/12827686/files/figure.png" pageId="6" pageNumber="7">Fig. 4C, D</figureCitation>
); 2 pairs of eyes; anterior and posterior furrow present, posterior furrow clearer than the anterior; the manducatory process with 5 apical setae, 10 setae on palp coxa; coxa I‒IV setae arranged: 5:8:10:17.
</paragraph>
<paragraph id="8BB436B8FF8BFFCFFF42FC1F816DFBA0" blockId="6.[174,775,253,1205]" pageId="6" pageNumber="7">
Legs (
<figureCitation id="13302A3DFF8BFFCFFE99FC1C82ECFBC3" box="[305,429,1021,1046]" captionStart="Fig" captionStartId="7.[136,167,1925,1944]" captionText="Fig. 6. Garypus sanasai Lin, Huang &amp; Chang sp. nov. (A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm." pageId="6" pageNumber="7">Fig. 6B, C</figureCitation>
): Metatarsi and tarsi not fused together in all legs, metatarsi slightly longer than tarsus in Leg I, but same length in Leg IV, transverse structure present between metatarsi and tarsi.
</paragraph>
<paragraph id="8BB436B8FF8BFFCFFF42FB9C80A6FE00" blockId="6.[174,775,253,1205]" lastBlockId="6.[850,1452,253,1206]" pageId="6" pageNumber="7">Abdomen. Tergites I–X and sternites I–X with median structure; setae at the bottom of each tergites; color pattern of tergites related to the presence of the setae; tergites I–XII setae arranged 6: 12: 11: 13: 9: 11: 11: 6: 7: 11: 8: 2, sternite IV–XII setae arranged 9: 10: 11: 12: 14: 12: 12: 4: 2; setae on genitalia (sternite II) smaller than the setae on other sternites, the setae at bottom - middle of sternite largest; Pleural membrane without setae.</paragraph>
<paragraph id="8BB436B8FF8BFFCFFC27FE3F80C3FD80" blockId="6.[850,1452,253,1206]" pageId="6" pageNumber="7">Genitalia: Not visible after incubation in Proteinase K for two days, 36 setae and 2 lyrifissure on anterior genital operculum, posterior with 9 setae in a row.</paragraph>
<paragraph id="8BB436B8FF8BFFCFFC27FDBC8745FC60" blockId="6.[850,1452,253,1206]" pageId="6" pageNumber="7">Dimensions (in mm): Body length (excluding chelicerae) 4.71. Chelicera 0.47/0.26; movable finger length 0.38. Pedipalp: trochanter 0.70/0.505; femur 1.66/0.40; patella 1.47/0.46; chela with pedicel 3.21/0.83; chela without pedicel 3.07 long; hand without pedicel 1.22 long; movable finger length 2.02. Carapace 1.36/1.40; anterior eye 0.14; posterior eye 0.12. Leg I: femur 0.80/0.23; patella 0.46/0.23; tibia 0.63/0.17; metatarsus 0.38/0.12; tarsus 0.32/0.10. Leg IV: femur + patella 1.40/0.32; tibia 1.10/0.16; metatarsus 0.45/0.15; tarsus 0.45/0.15.</paragraph>
<paragraph id="8BB436B8FF8BFFC1FC27FC5C83B5FEC0" blockId="6.[850,1452,253,1206]" lastBlockId="8.[174,775,253,342]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<emphasis id="B97FEAAAFF8BFFCFFC27FC5C80ADFC00" box="[911,1004,957,981]" italics="true" pageId="6" pageNumber="7">Remarks</emphasis>
: Green Island is a volcanic island located about 
<quantity id="4CF39B5DFF8BFFCFFC3DFC3F8098FC20" box="[917,985,989,1014]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="7" unit="km" value="33.0">33 km</quantity>
east of 
<collectingCountry id="F31C7628FF8BFFCFFB86FC3C873DFC20" box="[1070,1148,989,1013]" name="Taiwan" pageId="6" pageNumber="7">Taiwan</collectingCountry>
and covers an area of about 
<quantity id="4CF39B5DFF8BFFCFFCFAFC1C80DAFBC0" box="[850,923,1021,1045]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="7" unit="km" value="16.0">16 km</quantity>
<superScript id="7C7E9BF0FF8BFFCFFC34FC1980E5FBD3" attach="right" box="[924,932,1016,1030]" fontSize="6" pageId="6" pageNumber="7">2</superScript>
. The coasts on the island are mostly formed by coral reef and are scattered with stretches of sandy beaches. The specimen of 
<taxonomicName id="4C0B4D3BFF8BFFCFFBC2FBDF865DFB80" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[1130,1308,1086,1109]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sanasai">
<emphasis id="B97FEAAAFF8BFFCFFBC2FBDF865DFB80" box="[1130,1308,1086,1109]" italics="true" pageId="6" pageNumber="7">Garypus sanasai</emphasis>
</taxonomicName>
was found in Ziping. Located in the southernmost part of the island, Ziping is a tidal pool and the most complete lagoon on Green Island. The beach of the lagoon is formed by finely divided coral reefs, on which are small patches of the low-growing plant “Reef 
<taxonomicName id="4C0B4D3BFF8AFFCEFDA8FEFC8122FEE0" box="[512,611,285,309]" class="Magnoliopsida" family="Lythraceae" genus="Pemphis" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">Pemphis</taxonomicName>
” (
<taxonomicName id="4C0B4D3BFF8AFFCEFD29FEFC829EFE83" authority="JR Forst and G Forst" authorityName="JR Forst and G Forst" class="Magnoliopsida" family="Lythraceae" genus="Pemphis" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="acidula">
<emphasis id="B97FEAAAFF8AFFCEFD29FEFC839DFE80" italics="true" pageId="7" pageNumber="8">Pemphis acidula</emphasis>
JR Forst and G Forst
</taxonomicName>
). The coral reef beach was scattered with stones about 
<quantity id="4CF39B5DFF8AFFCEFE57FEBF8176FEA0" box="[511,567,350,373]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="8" unit="cm" value="7.0">7 cm</quantity>
in radius. The new species was found hiding under one of them. The individual kept its palps and legs close to its body and did not move when the stone was removed, probably to avoid being detected. No other animals were found under the same stone. After searching for other similarsized stones nearby, no other pseudoscorpions were found.
</paragraph>
<caption id="DF746630FF8BFFCFFF06F843870BF863" ID-DOI="http://doi.org/10.5281/zenodo.12827688" ID-Zenodo-Dep="12827688" box="[174,1098,1954,1974]" httpUri="https://zenodo.org/record/12827688/files/figure.png" pageId="6" pageNumber="7" startId="6.[174,205,1954,1973]" targetBox="[223,1402,1259,1929]" targetPageId="6" targetType="figure">
<paragraph id="8BB436B8FF8BFFCFFF06F843870BF863" blockId="6.[174,1098,1954,1974]" box="[174,1098,1954,1974]" pageId="6" pageNumber="7">
<emphasis id="B97FEAAAFF8BFFCFFF06F84383A7F860" bold="true" box="[174,230,1954,1973]" pageId="6" pageNumber="7">Fig. 5.</emphasis>
<taxonomicName id="4C0B4D3BFF8BFFCFFF59F842817AF860" authority="Lin, Huang &amp; Chang" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[241,571,1954,1974]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sanasai" status="sp. nov.">
<emphasis id="B97FEAAAFF8BFFCFFF59F84282C0F860" box="[241,385,1955,1974]" italics="true" pageId="6" pageNumber="7">Garypus sanasai</emphasis>
Lin, Huang &amp; Chang
</taxonomicName>
<taxonomicNameLabel id="A24C57D1FF8BFFCFFDE8F84281C2F863" box="[576,643,1955,1974]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(A) dorsal view; (B) ventral view. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF746630FF8AFFCEFF20F86480F2F861" pageId="7" pageNumber="8" startId="7.[136,167,1925,1944]" targetType="figure">
<paragraph id="8BB436B8FF8AFFCEFF20F86480F2F861" blockId="7.[136,1413,1925,1972]" pageId="7" pageNumber="8">
<emphasis id="B97FEAAAFF8AFFCEFF20F8648380F84D" bold="true" box="[136,193,1925,1944]" pageId="7" pageNumber="8">Fig. 6.</emphasis>
<taxonomicName id="4C0B4D3BFF8AFFCEFF67F864815EF84D" authority="Lin, Huang &amp; Chang" authorityName="Lin, Huang &amp; Chang" authorityYear="2022" box="[207,543,1925,1944]" class="Arachnida" family="Garypidae" genus="Garypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sanasai" status="sp. nov.">
<emphasis id="B97FEAAAFF8AFFCEFF67F8648221F842" box="[207,352,1925,1944]" italics="true" pageId="7" pageNumber="8">Garypus sanasai</emphasis>
Lin, Huang &amp; Chang
</taxonomicName>
<taxonomicNameLabel id="A24C57D1FF8AFFCEFD8DF8648128F84D" box="[549,617,1925,1944]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
(A) dorsal view of left palp; (B) left leg IV; (C) left leg I; (D) rallum of left chelicera; (E) galea of left chelicera; (F) lateral view of right palp. Scale bars: A–D = 1 mm; E, F = 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BB436B8FF85FFC1FF42FEFC8227FE80" blockId="8.[174,775,253,342]" pageId="8" pageNumber="9">
<emphasis id="B97FEAAAFF85FFC1FF42FEFC822DFEE0" box="[234,364,285,309]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
: 
<typeStatus id="54B0881AFF85FFC1FED4FEFF82F0FEE3" box="[380,433,286,310]" pageId="8" pageNumber="9">Type</typeStatus>
locality (Green Island, 
<collectingRegion id="49CFF85AFF85FFC1FD1EFEFC83BFFE83" country="Taiwan" name="Taitung" pageId="8" pageNumber="9">Taitung County</collectingRegion>
, 
<collectingRegion id="49CFF85AFF85FFC1FEA0FEDC8223FE80" box="[264,354,317,341]" country="China" name="Taiwan)" pageId="8" pageNumber="9">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>