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<document id="1F1D3FB24DAF7DC0AE074ECFA86A0108" ID-CLB-Dataset="28899" ID-DOI="10.5852/ejt.2018.462" ID-GBIF-Dataset="328c4344-2529-4aae-9c38-a3f9cb576c35" ID-Zenodo-Dep="1467972" ID-ZooBank="urn:lsid:zoobank.org:pub:60992A95-1F59-4407-9774-B8FC1545A8B4" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1540146986426" checkinUser="plazi" docAuthor="Ruttapon Srisonchai, Henrik Enghoff, Natdanai Likhitrakarn &amp; Somsak Panha" docDate="2018" docId="03A287AE0D712274F774EA7BFAFC5C2D" docLanguage="en" docName="ejt-462_srisonchai_enghoff_likhitrkarn_panha.pdf.imf" docOrigin="European Journal of Taxonomy 462" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nagaxytes acantherpestes, gen. et comb. nov." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FF9BFFD60D7B2265F629EF21FF8A5847" masterDocTitle="A revision of dragon millipedes II: the new genus Nagaxytes gen. nov., with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="11" updateTime="1698580138259" updateUser="ExternalLinkService">
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<mods:title id="22B69B092D12778AE62B531E63AE313C">A revision of dragon millipedes II: the new genus Nagaxytes gen. nov., with the description of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="8CA87E483A0EDC314A8D8C1DFFA7C283">Ruttapon Srisonchai</mods:namePart>
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<mods:namePart id="DE81A4F0DF539C511C66D9A3AB360632">Henrik Enghoff</mods:namePart>
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<mods:namePart id="3ED8546C67CF360211DD027B2CE49860">Natdanai Likhitrakarn</mods:namePart>
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<mods:namePart id="01BAA9D4F193FACF41D97DF9EF1236F8">Somsak Panha</mods:namePart>
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<treatment id="03A287AE0D712274F774EA7BFAFC5C2D" ID-DOI="http://doi.org/10.5281/zenodo.5946402" ID-GBIF-Taxon="148885085" ID-Zenodo-Dep="5946402" LSID="urn:lsid:plazi:treatment:03A287AE0D712274F774EA7BFAFC5C2D" httpUri="http://treatment.plazi.org/id/03A287AE0D712274F774EA7BFAFC5C2D" lastPageId="17" lastPageNumber="18" pageId="10" pageNumber="11">
<subSubSection id="C31165330D71226FF774EA7BFB5D5D33" box="[349,1239,1369,1396]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BB436B80D71226FF774EA7BFB5D5D33" blockId="10.[349,1239,1369,1432]" box="[349,1239,1369,1396]" pageId="10" pageNumber="11">
<taxonomicName id="4C0B4D3B0D71226FF774EA7BFC725D34" ID-CoL="45KDP" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[349,1016,1369,1396]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D71226FF774EA7BFD1F5D33" bold="true" box="[349,661,1369,1396]" italics="true" pageId="10" pageNumber="11">Nagaxytes acantherpestes</emphasis>
(
<bibRefCitation id="EF9A4B490D71226FF48DEA78FC7A5D34" box="[676,1008,1369,1395]" pageId="10" pageNumber="11" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff, 1994</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D71226FF229EA7BFB5D5D33" box="[1024,1239,1369,1396]" pageId="10" pageNumber="11">gen. et comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31165330D71226FF4F8EA5FFCE85DDF" box="[721,866,1406,1432]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BB436B80D71226FF4F8EA5FFCE85DDF" blockId="10.[349,1239,1369,1432]" box="[721,866,1406,1432]" pageId="10" pageNumber="11">
<figureCitation id="13302A3D0D71226FF4F8EA5FFC965DDF" box="[721,796,1406,1432]" captionStart="Fig. 2" captionStartId="5.[189,232,1872,1898]" captionTargetBox="[192,1396,282,1802]" captionTargetId="figure@5.[189,1398,269,1817]" captionTargetPageId="5" captionText="Fig. 2. General body characters of Nagaxytes Srisonchai, Enghoff &amp; Panha gen. nov. – N. acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype, Prachuap Khiri Khan Province, Hua Hin District (ZMUC 000101457). A. Entire body. B. Anterior body part. C. Body rings 5–8, showing sternal lobe between coxae 4 and gonopods on ring 7. D. Head and antennae." httpUri="https://zenodo.org/record/1467976/files/figure.png" pageId="10" pageNumber="11">Figs 2</figureCitation>
, 
<figureCitation id="13302A3D0D71226FF501EA5FFCE85DDF" box="[808,866,1406,1432]" captionStart-0="Fig. 6" captionStart-1="Fig. 7" captionStart-2="Fig. 8" captionStart-3="Fig. 9" captionStart-4="Fig. 10" captionStartId-0="11.[189,232,1934,1960]" captionStartId-1="13.[189,232,1904,1930]" captionStartId-2="14.[189,232,1915,1941]" captionStartId-3="15.[189,232,1938,1964]" captionStartId-4="16.[189,232,1915,1941]" captionTargetBox-0="[189,1399,346,1894]" captionTargetBox-1="[192,1391,295,1832]" captionTargetBox-2="[193,1393,299,1846]" captionTargetBox-3="[189,1386,320,1856]" captionTargetBox-4="[189,1399,296,1846]" captionTargetId-0="figure@11.[189,1399,344,1894]" captionTargetId-1="figure@13.[189,1399,291,1841]" captionTargetId-2="figure@14.[189,1399,295,1846]" captionTargetId-3="figure@15.[189,1399,306,1856]" captionTargetId-4="figure@16.[189,1399,296,1846]" captionTargetPageId-0="11" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="Fig. 6. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov. A. Drawing of ♂, holotype (ZMUC 000101457). B. Parasitic mite. C. Habitat." captionText-1="Fig. 7. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Anterior body part. B. Body rings 8–10. C. Posteriormost body rings and telson. D. Head and antenna. E. ♂ leg 5 (right). F–G. Midbody ring." captionText-2="Fig. 8. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Body rings 8–10. B. Paraterga of ring 10. C–D. Last ring and telson. E. Hypoproct. F–G. Epiproct. H–J. Sternal lobe between coxae 4." captionText-3="Fig. 9. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., specimen from Ban Yan Sue (Thailand), right gonopod. A. Mesal view. B. Dorsal view. C. Lateral view. D. Ventral view." captionText-4="Fig. 10. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., specimen from Ban Yan Sue (Thailand), right gonopod. A. Lateral view. B. Mesal view. C. Ventral view. D, F. Subdorsal view. E. Dorsal view." httpUri-0="https://zenodo.org/record/1467984/files/figure.png" httpUri-1="https://zenodo.org/record/1467986/files/figure.png" httpUri-2="https://zenodo.org/record/1467988/files/figure.png" httpUri-3="https://zenodo.org/record/1467990/files/figure.png" httpUri-4="https://zenodo.org/record/1467992/files/figure.png" pageId="10" pageNumber="11">6–10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31165330D71226FF694EAE0FAFD5E5A" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BB436B80D71226FF694EAE0FBB05D9D" blockId="10.[189,1082,1472,1499]" box="[189,1082,1472,1499]" pageId="10" pageNumber="11">
<taxonomicName id="4C0B4D3B0D71226FF694EAE0FCF75D9C" ID-CoL="6CNL8" authority="Golovatch &amp; Enghoff, 1994: 51" authorityName="Golovatch &amp; Enghoff" authorityPageNumber="51" authorityYear="1994" box="[189,893,1472,1499]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D71226FF694EAE0FD8A5D9D" box="[189,512,1472,1498]" italics="true" pageId="10" pageNumber="11">Desmoxytes acantherpestes</emphasis>
<bibRefCitation id="EF9A4B490D71226FF421EAE1FCDF5D9D" box="[520,853,1472,1498]" pageId="10" pageNumber="11" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff, 1994</bibRefCitation>
: 51
</taxonomicName>
–53, figs 21–28.
</paragraph>
<paragraph id="8BB436B80D71226FF694E922FAFD5E5A" blockId="10.[189,1399,1538,1565]" box="[189,1399,1538,1565]" pageId="10" pageNumber="11">
<taxonomicName id="4C0B4D3B0D71226FF694E922FE765E5A" ID-CoL="6CNL8" authorityName="Golovatch &amp; Enghoff" authorityYear="1994" box="[189,508,1539,1565]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D71226FF694E922FE765E5A" box="[189,508,1539,1565]" italics="true" pageId="10" pageNumber="11">Desmoxytes acantherpestes</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EF9A4B490D71226FF43FE922FD335E5A" box="[534,697,1538,1565]" pageId="10" pageNumber="11" refId="ref11455" refString="Enghoff H. 2005. The millipedes of Thailand (Diplopoda). Steenstrupia 29 (1): 87 - 103. Available from http: // www. zmuc. dk / commonweb / JOURNALS / PDF / Vol 29 - 1 / Enghoff. pdf [accessed 22 Aug. 2018]." type="journal article">Enghoff 2005</bibRefCitation>
: 96. — 
<bibRefCitation id="EF9A4B490D71226FF538E922FC205E5A" box="[785,938,1539,1565]" pageId="10" pageNumber="11" refId="ref11383" refString="Decker P. 2010. Contributions to the myriapod fauna of Thailand - New records of millipedes and centipedes from Thailand (Myriapoda: Diplopoda, Chilopoda). Schubartiana 4: 23 - 34. Available from http: // schubartiana. de / issues / pdf / vol 4 / Decker- 2010 - Myriapod _ fauna _ of _ Thailand. pdf [accessed 22 Aug. 2018]." type="journal article">Decker 2010</bibRefCitation>
: 28. — 
<bibRefCitation id="EF9A4B490D71226FF228E922FAA15E5A" box="[1025,1323,1539,1565]" pageId="10" pageNumber="11" refId="ref11855" refString="Nguyen D. A. &amp; Sierwald P. 2013. A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9 (6): 1132 - 1353. https: // doi. org / 10.15560 / 9.6.1132" type="journal article">Nguyen &amp; Sierwald 2013</bibRefCitation>
: 1240.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D71226FF694E964FF755EBC" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BB436B80D71226FF694E964FEBC5E18" blockId="10.[189,310,1605,1631]" box="[189,310,1605,1631]" pageId="10" pageNumber="11">
<heading id="D0FC81D40D71226FF694E964FEBC5E18" bold="true" box="[189,310,1605,1631]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B97FEAAA0D71226FF694E964FEBC5E18" bold="true" box="[189,310,1605,1631]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D71226FF694E954FF755EBC" blockId="10.[189,1399,1653,1787]" pageId="10" pageNumber="11">
Metaterga 4–19 with one row of 2+2 posterior spines (anterior row absent). Similar in this respect to 
<taxonomicName id="4C0B4D3B0D71226FF694E9BBFEB35EF4" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[189,313,1689,1715]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gracilis" status="gen. et sp. nov.">
<emphasis id="B97FEAAA0D71226FF694E9BBFEB35EF4" box="[189,313,1689,1715]" italics="true" pageId="10" pageNumber="11">N. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D71226FF769E9BBFE7A5EF3" box="[320,496,1690,1716]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
but differs from this species by having paraterga shorter; degree of elevation of paraterga lower; sternal lobe between male coxae 4 subtrapeziform; lamina lateralis swollen, stout, short.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D712269F694E805FE5859C8" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BB436B80D71226FF694E805FE205F79" blockId="10.[189,426,1828,1854]" box="[189,426,1828,1854]" pageId="10" pageNumber="11">
<heading id="D0FC81D40D71226FF694E805FE205F79" bold="true" box="[189,426,1828,1854]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B97FEAAA0D71226FF694E805FE205F79" bold="true" box="[189,426,1828,1854]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D71226FF6C3E875FBE05FF0" blockId="10.[189,1399,1876,1975]" pageId="10" pageNumber="11">
<materialsCitation id="3B633CE50D71226FF6C3E875FBE05FF0" ID-GBIF-Occurrence="1931420962" collectingDate="1979-08-08" collectionCode="ZMUC" collectorName="B. Petersen" country="Thailand" county="Hua Hin District" location="western coast of Siam Gulf (Gulf of Thailand), Hua Hin [Prachuap Khiri Khan Province" pageId="10" pageNumber="11" specimenCode="ZMUC 000101457" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B97FEAAA0D71226FF6C3E875FED65F29" bold="true" box="[234,348,1876,1902]" pageId="10" pageNumber="11">
<typeStatus id="54B0881A0D71226FF6C3E875FED65F29" box="[234,348,1876,1902]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F31C76280D71226FF694E859FED35FD5" box="[189,345,1912,1938]" name="Thailand" pageId="10" pageNumber="11">THAILAND</collectingCountry>
: 
<specimenCount id="9D0DFD310D71226FF743E859FE0B5FD5" box="[362,385,1912,1938]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
, 
<location id="8ED460630D71226FF7BDE859FEA25FF1" LSID="urn:lsid:plazi:treatment:03A287AE0D712274F774EA7BFAFC5C2D:8ED460630D71226FF7BDE859FEA25FF1" country="Thailand" county="Hua Hin District" name="western coast of Siam Gulf (Gulf of Thailand), Hua Hin [Prachuap Khiri Khan Province" pageId="10" pageNumber="11">western coast of Siam Gulf (Gulf of Thailand), Hua Hin [Prachuap Khiri Khan Province</location>
, 
<collectingCounty id="62D54E340D71226FF71DE8BCFE775FF1" box="[308,509,1947,1975]" pageId="10" pageNumber="11">Hua Hin District</collectingCounty>
], 
<collectingDate id="EFF1E9900D71226FF439E8BDFD285FF1" box="[528,674,1948,1974]" pageId="10" pageNumber="11" value="1979-08-08">
<date id="FFB510780D71226FF439E8BDFD285FF1" box="[528,674,1948,1974]" pageId="10" pageNumber="11" value="1979-08-08">8 Aug. 1979</date>
</collectingDate>
, 
<collectorName id="26FE536E0D71226FF486E8BCFCBE5FF0" box="[687,820,1949,1975]" pageId="10" pageNumber="11">B. Petersen</collectorName>
leg. (
<specimenCode id="DBAD9EC30D71226FF55FE8BDFBD45FF0" box="[886,1118,1948,1975]" pageId="10" pageNumber="11">ZMUC 000101457</specimenCode>
).
</materialsCitation>
</paragraph>
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<paragraph id="8BB436B80D70226EF694E8AFFC0B5F8C" blockId="11.[189,1399,1934,1996]" pageId="11" pageNumber="12">
<emphasis id="B97FEAAA0D70226EF694E8AFFE875FEF" bold="true" box="[189,269,1934,1960]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
<taxonomicName id="4C0B4D3B0D70226EF73FE8AEFC3E5FEF" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[278,948,1934,1960]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D70226EF73FE8AEFDC35FEF" box="[278,585,1934,1960]" italics="true" pageId="11" pageNumber="12">Nagaxytes acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D70226EF594E8AEFB105FEE" box="[957,1178,1934,1961]" pageId="11" pageNumber="12">gen. et comb. nov.</taxonomicNameLabel>
<emphasis id="B97FEAAA0D70226EF28BE8AFFB3D5FEF" bold="true" box="[1186,1207,1934,1960]" pageId="11" pageNumber="12">A</emphasis>
. Drawing of ♂, holotype 
<emphasis id="B97FEAAA0D70226EF702E890FEBF5F8C" bold="true" box="[299,309,1969,1995]" pageId="11" pageNumber="12">(</emphasis>
ZMUC 000101457). 
<emphasis id="B97FEAAA0D70226EF407E893FDC85F8B" bold="true" box="[558,578,1970,1996]" pageId="11" pageNumber="12">B</emphasis>
. Parasitic mite. 
<emphasis id="B97FEAAA0D70226EF4D6E890FC9E5F8C" bold="true" box="[767,788,1969,1995]" pageId="11" pageNumber="12">C</emphasis>
. Habitat.
</paragraph>
</caption>
<paragraph id="8BB436B80D772269F6C3EE2AFE2C5962" blockId="12.[189,1398,267,399]" box="[234,422,267,293]" pageId="12" pageNumber="13">
<emphasis id="B97FEAAA0D772269F6C3EE2AFE2C5962" bold="true" box="[234,422,267,293]" pageId="12" pageNumber="13">Other material</emphasis>
</paragraph>
<paragraph id="8BB436B80D772269F694EE0FFE5859C8" blockId="12.[189,1398,267,399]" pageId="12" pageNumber="13">
<materialsCitation id="3B633CE50D772269F694EE0FFE5859C8" ID-GBIF-Occurrence="1931420900" collectingDate="2007-08-31" collectionCode="CUMZ" collectorName="ASRU members leg" country="Thailand" elevation="147" latitude="12.053333" location="Hat Kham Subdistrict" longLatPrecision="21" longitude="99.63111" municipality="Kui Buri District" pageId="12" pageNumber="13" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Prachuap Khiri Khan Province">
<collectingCountry id="F31C76280D772269F694EE0FFED3590F" box="[189,345,302,328]" name="Thailand" pageId="12" pageNumber="13">THAILAND</collectingCountry>
: 
<specimenCount id="9D0DFD310D772269F74CEE0FFE21590E" box="[357,427,302,329]" pageId="12" pageNumber="13" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="9D0DFD310D772269F790EE0FFE6C590E" box="[441,486,302,329]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
(gonopods lost), 
<specimenCount id="9D0DFD310D772269F491EE0FFD6F590E" box="[696,741,302,329]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="49CFF85A0D772269F4DCEE0FFBEF590F" box="[757,1125,302,328]" country="Thailand" name="Prachuap Khiri Khan" pageId="12" pageNumber="13">Prachuap Khiri Khan Province</collectingRegion>
, 
<collectingMunicipality id="6BD0ACC20D772269F25BEE0FFAB5590F" box="[1138,1343,302,328]" pageId="12" pageNumber="13">Kui Buri District</collectingMunicipality>
, 
<location id="8ED460630D772269F365EE0EFE18592C" LSID="urn:lsid:plazi:treatment:03A287AE0D712274F774EA7BFAFC5C2D:8ED460630D772269F365EE0EFE18592C" country="Thailand" latitude="12.053333" longLatPrecision="21" longitude="99.63111" municipality="Kui Buri District" name="Hat Kham Subdistrict" pageId="12" pageNumber="13" stateProvince="Prachuap Khiri Khan Province">Hat Kham Subdistrict</location>
, 
<location id="8ED460630D772269F788EE73FDC9592C" LSID="urn:lsid:plazi:treatment:03A287AE0D712274F774EA7BFAFC5C2D:8ED460630D772269F788EE73FDC9592C" box="[417,579,337,364]" country="Thailand" latitude="12.053333" longLatPrecision="21" longitude="99.63111" municipality="Kui Buri District" name="Ban Yan Sue" pageId="12" pageNumber="13" stateProvince="Prachuap Khiri Khan Province">Ban Yan Sue</location>
, 
<geoCoordinate id="EE3F507F0D772269F47AEE6EFD66592B" box="[595,748,335,364]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="12.053333">12°03′12″ N</geoCoordinate>
, 
<geoCoordinate id="EE3F507F0D772269F4D5EE6EFC1B592B" box="[764,913,335,364]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="99.63111">99°37′52″ E</geoCoordinate>
, ca. 
<elevation id="0026D18B0D772269F5E7EE70FB97592B" box="[974,1053,337,364]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="12" pageNumber="13" unit="m" value="147.0">
<quantity id="4CF39B5D0D772269F5E7EE70FB97592B" box="[974,1053,337,364]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="12" pageNumber="13" unit="m" value="147.0">147 m</quantity>
</elevation>
a.s.l., 
<collectingDate id="EFF1E9900D772269F258EE70FA91592C" box="[1137,1307,337,363]" pageId="12" pageNumber="13" value="2007-08-31">
<date id="FFB510780D772269F258EE70FA91592C" box="[1137,1307,337,363]" pageId="12" pageNumber="13" value="2007-08-31">31 Aug. 2007</date>
</collectingDate>
, 
<collectorName id="26FE536E0D772269F301EE70FEDD59C9" pageId="12" pageNumber="13">ASRU members leg</collectorName>
. (
<collectionCode id="ED1AAE7D0D772269F742EE54FE4D59C8" box="[363,455,373,399]" country="Thailand" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="12" pageNumber="13">CUMZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31165330D772269F694EE97FD925F41" pageId="12" pageNumber="13" type="description">
<paragraph id="8BB436B80D772269F694EE97FEE15997" blockId="12.[189,1399,438,581]" box="[189,363,438,464]" pageId="12" pageNumber="13">
<heading id="D0FC81D40D772269F694EE97FEE15997" bold="true" box="[189,363,438,464]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B97FEAAA0D772269F694EE97FEE15997" bold="true" box="[189,363,438,464]" pageId="12" pageNumber="13">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D772269F694EEC4FE5D5A02" blockId="12.[189,1399,438,581]" pageId="12" pageNumber="13">
SIZE. Length 
<quantity id="4CF39B5D0D772269F776EEC4FE6959B8" box="[351,483,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="12" pageNumber="13" unit="mm" value="33.0" valueMax="34.0" valueMin="32.0">32–34 mm</quantity>
(male), 
<quantity id="4CF39B5D0D772269F467EEC5FD2F59B8" box="[590,677,484,511]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="13" unit="mm" value="36.0">36 mm</quantity>
(female); width of midbody metazona ca. 
<quantity id="4CF39B5D0D772269F290EEC4FA9D59B8" box="[1209,1303,485,511]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="mm" value="2.2">2.2 mm</quantity>
(male), 3.0 mm (female). Width of head &lt;body ring 2 = 3 &lt;collum &lt;4 &lt;5–16, thereafter body gradually tapering towards telson.
</paragraph>
<paragraph id="8BB436B80D772269F694ED4DFE8C5A8A" blockId="12.[189,1398,620,717]" pageId="12" pageNumber="13">COLOUR. Specimens in life brown. Colour in alcohol: after 29 years changed to pale brown; head, antennae, collum, metaterga, surface below paraterga, sterna, epiproct pale brown; paraterga brownish white.</paragraph>
<paragraph id="8BB436B80D772269F694EDD5FAFD5B76" blockId="12.[189,1399,756,818]" pageId="12" pageNumber="13">
COLLUM (
<figureCitation id="13302A3D0D772269F705EDD5FE015B49" box="[300,395,756,783]" captionStart="Fig. 7" captionStartId="13.[189,232,1904,1930]" captionTargetBox="[192,1391,295,1832]" captionTargetId="figure@13.[189,1399,291,1841]" captionTargetPageId="13" captionText="Fig. 7. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Anterior body part. B. Body rings 8–10. C. Posteriormost body rings and telson. D. Head and antenna. E. ♂ leg 5 (right). F–G. Midbody ring." httpUri="https://zenodo.org/record/1467986/files/figure.png" pageId="12" pageNumber="13">Fig. 7A</figureCitation>
). With setae in anterior row and intermediate row, spines in posterior row; paraterga of collum elevated at ca. 35–40°, directed caudolaterad, with two conspicuous notches at lateral margin.
</paragraph>
<paragraph id="8BB436B80D772269F694EC78FE505BD1" blockId="12.[189,1398,857,918]" pageId="12" pageNumber="13">
ANTENNAE (
<figureCitation id="13302A3D0D772269F76CEC78FE2E5B34" box="[325,420,857,883]" captionStart="Fig. 7" captionStartId="13.[189,232,1904,1930]" captionTargetBox="[192,1391,295,1832]" captionTargetId="figure@13.[189,1399,291,1841]" captionTargetPageId="13" captionText="Fig. 7. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Anterior body part. B. Body rings 8–10. C. Posteriormost body rings and telson. D. Head and antenna. E. ♂ leg 5 (right). F–G. Midbody ring." httpUri="https://zenodo.org/record/1467986/files/figure.png" pageId="12" pageNumber="13">Fig. 7D</figureCitation>
). Moderately long and slender, reaching to body ring 5 or 6 (male) and 5 (female) when stretched dorsally.
</paragraph>
<paragraph id="8BB436B80D772269F694EC9FFBDE5B90" blockId="12.[189,1108,957,984]" box="[189,1108,957,984]" pageId="12" pageNumber="13">TEGUMENT. Quite dull. Stricture between prozona and metazona shallow, wide.</paragraph>
<paragraph id="8BB436B80D772269F694ECDEFE635C27" blockId="12.[189,1399,1023,1120]" pageId="12" pageNumber="13">
METATERGA (
<figureCitation id="13302A3D0D772269F779ECDEFE5A5C5E" box="[336,464,1023,1049]" captionStart="Fig. 7" captionStartId="13.[189,232,1904,1930]" captionTargetBox="[192,1391,295,1832]" captionTargetId="figure@13.[189,1399,291,1841]" captionTargetPageId="13" captionText="Fig. 7. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Anterior body part. B. Body rings 8–10. C. Posteriormost body rings and telson. D. Head and antenna. E. ♂ leg 5 (right). F–G. Midbody ring." httpUri="https://zenodo.org/record/1467986/files/figure.png" pageId="12" pageNumber="13">Fig. 7A–C</figureCitation>
). With one or two transverse rows of setae and spines; metatergum 2 with?1+?1 anterior setae (inconspicuous) and 2+2 posterior spines; metaterga 3–19 with 2+2 posterior spines (lateral spines very long).
</paragraph>
<paragraph id="8BB436B80D772269F694EBA6FAFD5CE6" blockId="12.[189,1399,1159,1185]" box="[189,1399,1159,1185]" pageId="12" pageNumber="13">PLEUROSTERNAL CARINAE. On body ring 2 long, crest-like; on body ring 3 a small ridges; thereafter missing.</paragraph>
<paragraph id="8BB436B80D772269F694EBE8FCF25D42" blockId="12.[189,1398,1224,1286]" pageId="12" pageNumber="13">
PARATERGA (
<figureCitation id="13302A3D0D772269F765EBE9FE425CA5" box="[332,456,1224,1251]" captionStart="Fig. 7" captionStartId="13.[189,232,1904,1930]" captionTargetBox="[192,1391,295,1832]" captionTargetId="figure@13.[189,1399,291,1841]" captionTargetPageId="13" captionText="Fig. 7. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Anterior body part. B. Body rings 8–10. C. Posteriormost body rings and telson. D. Head and antenna. E. ♂ leg 5 (right). F–G. Midbody ring." httpUri="https://zenodo.org/record/1467986/files/figure.png" pageId="12" pageNumber="13">Figs 7F–G</figureCitation>
, 
<figureCitation id="13302A3D0D772269F7FEEBE9FE765CA5" box="[471,508,1224,1250]" captionStart="Fig. 8" captionStartId="14.[189,232,1915,1941]" captionTargetBox="[193,1393,299,1846]" captionTargetId="figure@14.[189,1399,295,1846]" captionTargetPageId="14" captionText="Fig. 8. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Body rings 8–10. B. Paraterga of ring 10. C–D. Last ring and telson. E. Hypoproct. F–G. Epiproct. H–J. Sternal lobe between coxae 4." httpUri="https://zenodo.org/record/1467988/files/figure.png" pageId="12" pageNumber="13">8B</figureCitation>
). Directed caudolaterad on body rings 2–18, elevated at ca. 60° (male) 50°– 60° (female), directed increasingly caudad on body ring 19.
</paragraph>
<paragraph id="8BB436B80D772269F694EA0CFA925DCA" blockId="12.[189,1399,1325,1422]" pageId="12" pageNumber="13">
TELSON (
<figureCitation id="13302A3D0D772269F702EA0CFE3F5D00" box="[299,437,1325,1351]" captionStart="Fig. 8" captionStartId="14.[189,232,1915,1941]" captionTargetBox="[193,1393,299,1846]" captionTargetId="figure@14.[189,1399,295,1846]" captionTargetPageId="14" captionText="Fig. 8. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Body rings 8–10. B. Paraterga of ring 10. C–D. Last ring and telson. E. Hypoproct. F–G. Epiproct. H–J. Sternal lobe between coxae 4." httpUri="https://zenodo.org/record/1467988/files/figure.png" pageId="12" pageNumber="13">Fig. 8C–G</figureCitation>
). Epiproct quite short; tip usually subtruncate (in some specimens slightly emarginate); lateral setiferous tubercles conspicuous, long, digitiform; apical tubercles conspicuous. Hypoproct subtriangular; caudal margin subtriangular, with inconspicuous setiferous tubercles.
</paragraph>
<paragraph id="8BB436B80D772269F694EA94FB4F5DB5" blockId="12.[189,1398,1460,1522]" pageId="12" pageNumber="13">
STERNA (
<figureCitation id="13302A3D0D772269F702EA95FE205D88" box="[299,426,1460,1487]" captionStart="Fig. 8" captionStartId="14.[189,232,1915,1941]" captionTargetBox="[193,1393,299,1846]" captionTargetId="figure@14.[189,1399,295,1846]" captionTargetPageId="14" captionText="Fig. 8. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., ♂, holotype (ZMUC 000101457). A. Body rings 8–10. B. Paraterga of ring 10. C–D. Last ring and telson. E. Hypoproct. F–G. Epiproct. H–J. Sternal lobe between coxae 4." httpUri="https://zenodo.org/record/1467988/files/figure.png" pageId="12" pageNumber="13">Fig. 8H–J</figureCitation>
). Cross-impressions quite deep. Sternal lobe between male coxae 4 swollen, subtrapeziform when seen in caudal view; base broad; tip slightly or deeply emarginate.
</paragraph>
<paragraph id="8BB436B80D772269F694E938FD925F41" blockId="12.[189,1399,1561,1798]" pageId="12" pageNumber="13">
GONOPODS (
<figureCitation id="13302A3D0D772269F764E938FE175E74" box="[333,413,1561,1587]" captionStart="Fig. 9" captionStartId="15.[189,232,1938,1964]" captionTargetBox="[189,1386,320,1856]" captionTargetId="figure@15.[189,1399,306,1856]" captionTargetPageId="15" captionText="Fig. 9. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., specimen from Ban Yan Sue (Thailand), right gonopod. A. Mesal view. B. Dorsal view. C. Lateral view. D. Ventral view." httpUri="https://zenodo.org/record/1467990/files/figure.png" pageId="12" pageNumber="13">Figs 9</figureCitation>
, 
<figureCitation id="13302A3D0D772269F787E938FE5A5E74" box="[430,464,1561,1587]" captionStart="Fig. 10" captionStartId="16.[189,232,1915,1941]" captionTargetBox="[189,1399,296,1846]" captionTargetId="figure@16.[189,1399,296,1846]" captionTargetPageId="16" captionText="Fig. 10. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov., specimen from Ban Yan Sue (Thailand), right gonopod. A. Lateral view. B. Mesal view. C. Ventral view. D, F. Subdorsal view. E. Dorsal view." httpUri="https://zenodo.org/record/1467992/files/figure.png" pageId="12" pageNumber="13">10</figureCitation>
). Coxa subequal in length to femur. Cannula long and slender. Telopodite quite long and slender. Prefemoral part ca. ¾ as long as femur. Femur quite long and slender. Mesal sulcus conspicuous, deep, wide; lateral sulcus conspicuous, very deep. Postfemoral part conspicuous, quite short. Solenophore well-developed: lamina lateralis swollen, stout, short, compact: lamina medialis very long; with a swollen base, gradually becoming thinner towards tip; apically fringed with several small spines; tip directed mesoventrad. Solenomere quite long, curved, tip directed ventrad.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D772274F694E80FFD485A85" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BB436B80D772269F694E80FFE645F0F" blockId="12.[189,494,1838,1864]" box="[189,494,1838,1864]" pageId="12" pageNumber="13">
<heading id="D0FC81D40D772269F694E80FFE645F0F" bold="true" box="[189,494,1838,1864]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B97FEAAA0D772269F694E80FFE645F0F" bold="true" box="[189,494,1838,1864]" pageId="12" pageNumber="13">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D772269F694E87DFD5C5FA7" blockId="12.[189,1399,1884,2017]" pageId="12" pageNumber="13">
This species is known from two locations (Hua Hin and Ban Yan Sue) in Prachuap Khirikhan Province. We believe that the 
<typeStatus id="54B0881A0D772269F783E85EFD9B5FDE" box="[426,529,1919,1945]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
was probably collected from limestone habitats because the 
<typeStatus id="54B0881A0D772269F2CDE8A1FA9D5FDD" box="[1252,1303,1920,1946]" pageId="12" pageNumber="13">type</typeStatus>
locality (near Hua Hin) lies in a limestone area. Specimens collected by us were from limestone habitats. It has been found crawling on rocks or on leaf litter.
</paragraph>
<caption id="DF7466300D762268F694E851FCA95F96" httpUri="https://zenodo.org/record/1467986/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1904,1930]" targetBox="[192,1391,295,1832]" targetPageId="13">
<paragraph id="8BB436B80D762268F694E851FCA95F96" blockId="13.[189,1399,1904,2002]" pageId="13" pageNumber="14">
<emphasis id="B97FEAAA0D762268F694E851FE835FCC" bold="true" box="[189,265,1904,1931]" pageId="13" pageNumber="14">Fig. 7.</emphasis>
<taxonomicName id="4C0B4D3B0D762268F739E853FC155FCC" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[272,927,1905,1931]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D762268F739E853FDCA5FCC" box="[272,576,1905,1931]" italics="true" pageId="13" pageNumber="14">Nagaxytes acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D762268F58FE850FBF75FCC" box="[934,1149,1905,1931]" pageId="13" pageNumber="14">gen. et comb. nov.</taxonomicNameLabel>
, ♂, holotype (ZMUC 000101457). 
<emphasis id="B97FEAAA0D762268F771E8B5FEE75FE9" bold="true" box="[344,365,1940,1966]" pageId="13" pageNumber="14">A</emphasis>
. Anterior body part. 
<emphasis id="B97FEAAA0D762268F475E8B5FDFA5FE9" bold="true" box="[604,624,1940,1966]" pageId="13" pageNumber="14">B</emphasis>
. Body rings 8–10. 
<emphasis id="B97FEAAA0D762268F560E8B5FCD45FE9" bold="true" box="[841,862,1940,1966]" pageId="13" pageNumber="14">C</emphasis>
. Posteriormost body rings and telson. 
<emphasis id="B97FEAAA0D762268F33FE8B5FAA15FE9" bold="true" box="[1302,1323,1940,1966]" pageId="13" pageNumber="14">D</emphasis>
. Head and antenna. 
<emphasis id="B97FEAAA0D762268F771E896FEE65F96" bold="true" box="[344,364,1975,2001]" pageId="13" pageNumber="14">E</emphasis>
. ♂ leg 5 (right). 
<emphasis id="B97FEAAA0D762268F41AE896FDE05F96" bold="true" box="[563,618,1975,2002]" pageId="13" pageNumber="14">F–G</emphasis>
. Midbody ring.
</paragraph>
</caption>
<caption id="DF7466300D75226BF694E85AFC925F9B" httpUri="https://zenodo.org/record/1467988/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1915,1941]" targetBox="[193,1393,299,1846]" targetPageId="14">
<paragraph id="8BB436B80D75226BF694E85AFC925F9B" blockId="14.[189,1399,1915,2012]" pageId="14" pageNumber="15">
<emphasis id="B97FEAAA0D75226BF694E85AFE835FD2" bold="true" box="[189,265,1915,1941]" pageId="14" pageNumber="15">Fig. 8.</emphasis>
<taxonomicName id="4C0B4D3B0D75226BF739E85DFC155FD2" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[272,927,1915,1941]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D75226BF739E85DFDCA5FD2" box="[272,576,1915,1941]" italics="true" pageId="14" pageNumber="15">Nagaxytes acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D75226BF58FE85DFBF75FD1" box="[934,1149,1915,1942]" pageId="14" pageNumber="15">gen. et comb. nov.</taxonomicNameLabel>
, ♂, holotype (ZMUC 000101457). 
<emphasis id="B97FEAAA0D75226BF775E8BFFEFB5FFF" bold="true" box="[348,369,1950,1976]" pageId="14" pageNumber="15">A</emphasis>
. Body rings 8–10. 
<emphasis id="B97FEAAA0D75226BF475E8BEFDFA5FFE" bold="true" box="[604,624,1951,1977]" pageId="14" pageNumber="15">B</emphasis>
. Paraterga of ring 10. 
<emphasis id="B97FEAAA0D75226BF5ABE8BFFC315FFE" bold="true" box="[898,955,1950,1977]" pageId="14" pageNumber="15">C–D</emphasis>
. Last ring and telson. 
<emphasis id="B97FEAAA0D75226BF2E4E8BEFB6B5FFE" bold="true" box="[1229,1249,1951,1977]" pageId="14" pageNumber="15">E</emphasis>
. Hypoproct. 
<emphasis id="B97FEAAA0D75226BF694E8E3FF7F5F9C" bold="true" box="[189,245,1985,2012]" pageId="14" pageNumber="15">F–G</emphasis>
. Epiproct. 
<emphasis id="B97FEAAA0D75226BF75EE8E3FE215F9B" bold="true" box="[375,427,1986,2012]" pageId="14" pageNumber="15">H–J</emphasis>
. Sternal lobe between coxae 4.
</paragraph>
</caption>
<caption id="DF7466300D74226AF694E8B3FAD75F88" httpUri="https://zenodo.org/record/1467990/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1938,1964]" targetBox="[189,1386,320,1856]" targetPageId="15">
<paragraph id="8BB436B80D74226AF694E8B3FAD75F88" blockId="15.[189,1398,1938,2000]" pageId="15" pageNumber="16">
<emphasis id="B97FEAAA0D74226AF694E8B3FE805FEB" bold="true" box="[189,266,1938,1964]" pageId="15" pageNumber="16">Fig. 9.</emphasis>
<taxonomicName id="4C0B4D3B0D74226AF738E8B2FC2F5FEB" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[273,933,1938,1964]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D74226AF738E8B2FDC95FEB" box="[273,579,1938,1964]" italics="true" pageId="15" pageNumber="16">Nagaxytes acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D74226AF585E8B3FB0F5FEB" box="[940,1157,1938,1964]" pageId="15" pageNumber="16">gen. et comb. nov.</taxonomicNameLabel>
, specimen from Ban Yan Sue (Thailand), right gonopod. 
<emphasis id="B97FEAAA0D74226AF44DE894FDF35F88" bold="true" box="[612,633,1973,1999]" pageId="15" pageNumber="16">A</emphasis>
. Mesal view. 
<emphasis id="B97FEAAA0D74226AF534E894FCBB5F88" bold="true" box="[797,817,1973,1999]" pageId="15" pageNumber="16">B</emphasis>
. Dorsal view. 
<emphasis id="B97FEAAA0D74226AF5F3E894FC655F88" bold="true" box="[986,1007,1973,1999]" pageId="15" pageNumber="16">C</emphasis>
. Lateral view. 
<emphasis id="B97FEAAA0D74226AF2B7E894FB395F88" bold="true" box="[1182,1203,1973,1999]" pageId="15" pageNumber="16">D</emphasis>
. Ventral view.
</paragraph>
</caption>
<caption id="DF7466300D6B2275F694E85AFE335F9B" httpUri="https://zenodo.org/record/1467992/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1915,1941]" targetBox="[189,1399,296,1846]" targetPageId="16">
<paragraph id="8BB436B80D6B2275F694E85AFE335F9B" blockId="16.[189,1398,1915,2012]" pageId="16" pageNumber="17">
<emphasis id="B97FEAAA0D6B2275F694E85AFE925FD2" bold="true" box="[189,280,1915,1941]" pageId="16" pageNumber="17">Fig. 10.</emphasis>
<taxonomicName id="4C0B4D3B0D6B2275F737E85DFC265FD2" authority="(Golovatch &amp; Enghoff, 1994)" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[286,940,1915,1941]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D6B2275F737E85DFDC45FD2" box="[286,590,1915,1941]" italics="true" pageId="16" pageNumber="17">Nagaxytes acantherpestes</emphasis>
(Golovatch &amp; Enghoff, 1994)
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D6B2275F59AE85DFB035FD1" box="[947,1161,1915,1942]" pageId="16" pageNumber="17">gen. et comb. nov.</taxonomicNameLabel>
, specimen from Ban Yan Sue (Thailand), right gonopod. 
<emphasis id="B97FEAAA0D6B2275F445E8BFFD0B5FFF" bold="true" box="[620,641,1950,1976]" pageId="16" pageNumber="17">A</emphasis>
. Lateral view. 
<emphasis id="B97FEAAA0D6B2275F51DE8BEFCC25FFE" bold="true" box="[820,840,1951,1977]" pageId="16" pageNumber="17">B</emphasis>
. Mesal view. 
<emphasis id="B97FEAAA0D6B2275F5D9E8BFFB8F5FFF" bold="true" box="[1008,1029,1950,1976]" pageId="16" pageNumber="17">C</emphasis>
. Ventral view. 
<emphasis id="B97FEAAA0D6B2275F293E8BEFB455FFE" bold="true" box="[1210,1231,1951,1977]" pageId="16" pageNumber="17">D</emphasis>
, 
<emphasis id="B97FEAAA0D6B2275F2F6E8BEFB7B5FFE" bold="true" box="[1247,1265,1951,1977]" pageId="16" pageNumber="17">F</emphasis>
. Subdorsal view. 
<emphasis id="B97FEAAA0D6B2275F72AE8E3FE9D5F9B" bold="true" box="[259,279,1986,2012]" pageId="16" pageNumber="17">E</emphasis>
. Dorsal view.
</paragraph>
</caption>
<paragraph id="8BB436B80D6A2274F694EE2AFEAB59C8" blockId="17.[189,1398,267,399]" pageId="17" pageNumber="18">
This species is likely to be syntopic with 
<taxonomicName id="4C0B4D3B0D6A2274F4B4EE2DFBC95961" authority="(Pocock, 1895)" authorityName="Pocock" authorityYear="1895" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[669,1091,267,294]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="planata">
<emphasis id="B97FEAAA0D6A2274F4B4EE2DFC005962" box="[669,906,267,293]" italics="true" pageId="17" pageNumber="18">Desmoxytes planata</emphasis>
(Pocock, 1895)
</taxonomicName>
at Ban Yan Sue; however, we noticed that the habitats of these two species are clearly different. 
<taxonomicName id="4C0B4D3B0D6A2274F5C6EE0EFAAA590E" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[1007,1312,303,329]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D6A2274F5C6EE0EFAAA590E" box="[1007,1312,303,329]" italics="true" pageId="17" pageNumber="18">Nagaxytes acantherpestes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D6A2274F301EE0EFECA592B" pageId="17" pageNumber="18">gen. et comb. nov.</taxonomicNameLabel>
lives on the ground or on limestone rocks while 
<taxonomicName id="4C0B4D3B0D6A2274F5A2EE72FB81592B" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[907,1035,338,364]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="planata">
<emphasis id="B97FEAAA0D6A2274F5A2EE72FB81592B" box="[907,1035,338,364]" italics="true" pageId="17" pageNumber="18">D. planata</emphasis>
</taxonomicName>
was found on tree trunks near garbage.
</paragraph>
<paragraph id="8BB436B80D6A2274F694EE97FAFD5A1A" blockId="17.[189,1399,438,606]" pageId="17" pageNumber="18">
<bibRefCitation id="EF9A4B490D6A2274F694EE97FEED5996" box="[189,359,438,465]" pageId="17" pageNumber="18" refId="ref11383" refString="Decker P. 2010. Contributions to the myriapod fauna of Thailand - New records of millipedes and centipedes from Thailand (Myriapoda: Diplopoda, Chilopoda). Schubartiana 4: 23 - 34. Available from http: // schubartiana. de / issues / pdf / vol 4 / Decker- 2010 - Myriapod _ fauna _ of _ Thailand. pdf [accessed 22 Aug. 2018]." type="journal article">Decker (2010)</bibRefCitation>
identified one male in NHMG, collected from Thanboke Khorani National Park, 
<collectingRegion id="49CFF85A0D6A2274F31DEE97FEAE59B3" country="Thailand" name="Krabi" pageId="17" pageNumber="18">Krabi Province</collectingRegion>
(
<geoCoordinate id="EE3F507F0D6A2274F71FEEF9FE3559B3" box="[310,447,472,500]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="8.386666">8°23′12″ N</geoCoordinate>
, 
<geoCoordinate id="EE3F507F0D6A2274F7E4EEF9FDD559B3" box="[461,607,472,500]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="98.73778">98°44′16″ E</geoCoordinate>
) as 
<taxonomicName id="4C0B4D3B0D6A2274F4BBEEFBFCEC59B3" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[658,870,474,500]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D6A2274F4BBEEFBFCEC59B3" box="[658,870,474,500]" italics="true" pageId="17" pageNumber="18">D. acantherpestes</emphasis>
</taxonomicName>
. This specimen might perhaps be ‘ 
<emphasis id="B97FEAAA0D6A2274F322EEFBFAFC59B4" box="[1291,1398,474,500]" italics="true" pageId="17" pageNumber="18">D.’ gigas</emphasis>
<bibRefCitation id="EF9A4B490D6A2274F694EEDCFD815A50" box="[189,523,509,535]" pageId="17" pageNumber="18" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff, 1994</bibRefCitation>
(in the ‘ 
<emphasis id="B97FEAAA0D6A2274F45EEEDFFD3C5A50" box="[631,694,510,535]" italics="true" pageId="17" pageNumber="18">gigas</emphasis>
’ group) because the location falls in the distribution range for this group in South 
<collectingCountry id="F31C76280D6A2274F7E7ED01FDBF5A7D" box="[462,565,544,570]" name="Thailand" pageId="17" pageNumber="18">Thailand</collectingCountry>
whereas 
<taxonomicName id="4C0B4D3B0D6A2274F48CED00FC945A7D" authorityName="Srisonchai, Enghoff &amp; Panha" authorityYear="2018" box="[677,798,545,570]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97FEAAA0D6A2274F48CED00FC945A7D" box="[677,798,545,570]" italics="true" pageId="17" pageNumber="18">Nagaxytes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D6A2274F50FED00FC1A5A7C" box="[806,912,545,571]" pageId="17" pageNumber="18" rank="genus">gen. nov.</taxonomicNameLabel>
(the ‘ 
<taxonomicName id="4C0B4D3B0D6A2274F5FEED01FB0F5A7D" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[983,1157,544,570]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D6A2274F5FEED01FB0F5A7D" box="[983,1157,544,570]" italics="true" pageId="17" pageNumber="18">acantherpestes</emphasis>
</taxonomicName>
’ group) is limited to western 
<collectingCountry id="F31C76280D6A2274F734ED62FE025A1A" box="[285,392,579,605]" name="Thailand" pageId="17" pageNumber="18">Thailand</collectingCountry>
. Moreover, Thanboke Khorani National Park and Hua Hin are far apart, about 
<quantity id="4CF39B5D0D6A2274F33EED62FAF95A1A" box="[1303,1395,579,606]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="500.0">500 km</quantity>
.
</paragraph>
<paragraph id="8BB436B80D6A2274F694EDA4FD485A85" blockId="17.[189,1398,645,706]" pageId="17" pageNumber="18">
This species is distributed in a narrow area; we regard 
<taxonomicName id="4C0B4D3B0D6A2274F515EDA4FB845AD8" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[828,1038,645,671]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acantherpestes" status="gen. et comb. nov.">
<emphasis id="B97FEAAA0D6A2274F515EDA4FB845AD8" box="[828,1038,645,671]" italics="true" pageId="17" pageNumber="18">N. acantherpestes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57D10D6A2274F23CEDA4FB665AD8" box="[1045,1260,645,671]" pageId="17" pageNumber="18">gen. et comb. nov.</taxonomicNameLabel>
as endemic to 
<collectingRegion id="49CFF85A0D6A2274F6F2ED89FDCC5A85" box="[219,582,680,706]" country="Thailand" name="Prachuap Khiri Khan" pageId="17" pageNumber="18">Prachuap Khiri Khan Province</collectingRegion>
, 
<collectingCountry id="F31C76280D6A2274F47BED89FD345A85" box="[594,702,680,706]" name="Thailand" pageId="17" pageNumber="18">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D6A2274F694EDC8FAFD5BA5" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BB436B80D6A2274F694EDC8FEA55B44" blockId="17.[189,303,745,771]" box="[189,303,745,771]" pageId="17" pageNumber="18">
<heading id="D0FC81D40D6A2274F694EDC8FEA55B44" bold="true" box="[189,303,745,771]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B97FEAAA0D6A2274F694EDC8FEA55B44" bold="true" box="[189,303,745,771]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB436B80D6A2274F694EC39FAFD5BA5" blockId="17.[189,1399,792,994]" pageId="17" pageNumber="18">
In the original description, 
<bibRefCitation id="EF9A4B490D6A2274F439EC39FCF95B75" box="[528,883,792,818]" pageId="17" pageNumber="18" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff (1994)</bibRefCitation>
did not comment on the live colouration of this species. We did not photograph living specimens, but our collector noticed a brown colouration. 
<bibRefCitation id="EF9A4B490D6A2274F77CEC7FFD3E5B3F" box="[341,692,862,888]" pageId="17" pageNumber="18" refId="ref11554" refString="Golovatch S. I. &amp; Enghoff H. 1994. Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 20 (2): 45 - 71." type="journal article">Golovatch &amp; Enghoff (1994)</bibRefCitation>
also described 
<taxonomicName id="4C0B4D3B0D6A2274F55CEC7EFBC75B3F" baseAuthorityName="Golovatch &amp; Enghoff" baseAuthorityYear="1994" box="[885,1101,862,888]" class="Diplopoda" family="Paradoxosomatidae" genus="Nagaxytes" higherTaxonomySource="CoL" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acantherpestes">
<emphasis id="B97FEAAA0D6A2274F55CEC7EFBC75B3F" box="[885,1101,862,888]" italics="true" pageId="17" pageNumber="18">D. acantherpestes</emphasis>
</taxonomicName>
as lacking pleurosternal 
<taxonomicName id="4C0B4D3B0D6A2274F694ECA0FE9F5BDC" authorityName="" authorityYear="1992" box="[189,277,897,923]" class="Insecta" family="Eccoptarthridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Carinae">carinae</taxonomicName>
, but after we re-examined the 
<typeStatus id="54B0881A0D6A2274F455ECA0FD695BDC" box="[636,739,897,923]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
and examined all newly collected specimens, we found the pleurosternal 
<taxonomicName id="4C0B4D3B0D6A2274F7A3EC84FE555BF8" authorityName="" authorityYear="1992" box="[394,479,933,959]" class="Insecta" family="Eccoptarthridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subFamily" subFamily="Carinae">carinae</taxonomicName>
as crest-like on ring 2, as small ridges on ring 3, thereafter absent. There is a variation in the tip of the epiproct which in some specimens is subtruncate, in others slightly emarginate.
</paragraph>
</subSubSection>
<subSubSection id="C31165330D6A2274F694EB28FAFC5C2D" pageId="17" pageNumber="18" type="biology_ecology">
<paragraph id="8BB436B80D6A2274F694EB28FAFC5C2D" blockId="17.[189,1398,1033,1130]" pageId="17" pageNumber="18">
On some specimens we found parasitic mites, probably of the genus 
<emphasis id="B97FEAAA0D6A2274F5F9EB2BFB965C64" box="[976,1052,1034,1059]" italics="true" pageId="17" pageNumber="18">Leptus</emphasis>
Latreille, 1795 (
<figureCitation id="13302A3D0D6A2274F2F2EB28FABF5C63" box="[1243,1333,1033,1060]" captionStart="Fig. 6" captionStartId="11.[189,232,1934,1960]" captionTargetBox="[189,1399,346,1894]" captionTargetId="figure@11.[189,1399,344,1894]" captionTargetPageId="11" captionText="Fig. 6. Nagaxytes acantherpestes (Golovatch &amp; Enghoff, 1994) gen. et comb. nov. A. Drawing of ♂, holotype (ZMUC 000101457). B. Parasitic mite. C. Habitat." httpUri="https://zenodo.org/record/1467984/files/figure.png" pageId="17" pageNumber="18">Fig. 6B</figureCitation>
). The mites infested the millipede on the metaterga. This association reminds us of what 
<bibRefCitation id="EF9A4B490D6A2274F242EB0DFAFC5C00" box="[1131,1398,1068,1095]" pageId="17" pageNumber="18" refId="ref12078" refString="Srisonchai R., Enghoff H., Likhitrakarn N. &amp; Panha S. 2018. A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 761: 1 - 177. https: // doi. org / 10.3897 / zookeys. 761.24214" type="journal article">
Srisonchai 
<emphasis id="B97FEAAA0D6A2274F2C3EB0CFAA95C01" box="[1258,1315,1068,1094]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(2018)
</bibRefCitation>
found in 
<taxonomicName id="4C0B4D3B0D6A2274F708EB70FD305C2D" authority="(Pocock, 1895)" authorityName="Pocock" authorityYear="1895" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[289,698,1103,1130]" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cervina">
<emphasis id="B97FEAAA0D6A2274F708EB70FD8E5C2D" box="[289,516,1105,1130]" italics="true" pageId="17" pageNumber="18">Desmoxytes cervina</emphasis>
(Pocock, 1895)
</taxonomicName>
: several engorged 
<emphasis id="B97FEAAA0D6A2274F5A0EB70FC5F5C2D" box="[905,981,1105,1130]" italics="true" pageId="17" pageNumber="18">Leptus</emphasis>
mites attached to the millipede body.
</paragraph>
</subSubSection>
</treatment>
</document>