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<document id="ACB9E1FA387DD24AACEE4D445F2A0288" ID-DOI="10.5252/geodiversitas2021v43a22" ID-ISSN="1638-9395" ID-Zenodo-Dep="5764379" ID-ZooBank="urn:lsid:zoobank.org:pub:6D1643AD-66A5-4678-9397-CE08E610D641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1638890191441" checkinUser="felipe" docAuthor="Vasseur, Raphaël &amp; Lathuilière, Bernard" docDate="2021" docId="03A287A248723107993D65CAFED4FC3F" docLanguage="en" docName="Geodiversitas.43.22.1187-1291.pdf" docOrigin="Geodiversitas 43 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Neorylstonia pseudocolumellata" docType="treatment" docVersion="7" lastPageNumber="1238" masterDocId="FF9BFFDA484331329A586119FFC1FFD0" masterDocTitle="Pliensbachian corals from the Western Tethys" masterLastPageNumber="1291" masterPageNumber="1187" pageNumber="1234" updateTime="1699441156090" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="0440229079B48DCC8C4D985113546A0D">Pliensbachian corals from the Western Tethys</mods:title>
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<mods:name id="FF7A4E7CF1033DC71585BD59C2180C4B" type="personal">
<mods:role id="79549A507CE43CF9CD1C334F424F4FE9">
<mods:roleTerm id="025E65017DE4E70BF80BAED56B05EA87">Author</mods:roleTerm>
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<mods:namePart id="B04661BA06A052C801617ED3B5149EBF">Vasseur, Raphaël</mods:namePart>
<mods:affiliation id="F955A3365E76C4FA723D3A94940408CE">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
<mods:nameIdentifier id="A77D1D5918B546053091050548F82BEC" type="email">raphaelvasseur@wanadoo.fr</mods:nameIdentifier>
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<mods:namePart id="78793EBC121B9511B6EF160FE964C956">Lathuilière, Bernard</mods:namePart>
<mods:affiliation id="1AD9BD58A89D1F865CCF8D0838080C0A">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
<mods:nameIdentifier id="E99007025D57B64A40A2761B7F673DCF" type="email">bernard.lathuiliere@univ-lorraine.fr</mods:nameIdentifier>
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<mods:title id="3AE70610E085258B0F3AC241AF884DA9">Geodiversitas</mods:title>
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<mods:date id="8FE2C5397BE145CFD53043539659751A">2021</mods:date>
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<mods:number id="AF5D2DDD89F6E51406A99494CC5B2465">2021-11-23</mods:number>
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<mods:number id="58BB300AD32B5917D9F1E3E37ECA36DF">43</mods:number>
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<treatment id="03A287A248723107993D65CAFED4FC3F" ID-DOI="http://doi.org/10.5281/zenodo.5765008" ID-Zenodo-Dep="5765008" LSID="urn:lsid:plazi:treatment:03A287A248723107993D65CAFED4FC3F" httpUri="http://treatment.plazi.org/id/03A287A248723107993D65CAFED4FC3F" lastPageId="53" lastPageNumber="1238" pageId="49" pageNumber="1234">
<subSubSection id="C311653F48723103993D65CAFAB6FB3F" box="[869,1399,1235,1263]" pageId="49" pageNumber="1234" type="nomenclature">
<paragraph id="8BB436B448723103993D65CAFAB6FB3F" blockId="49.[869,1399,1235,1294]" box="[869,1399,1235,1263]" pageId="49" pageNumber="1234">
<taxonomicName id="4C0B4D3748723103993D65CAFAB6FB3F" ID-CoL="d11bfea7-f4f3-4074-8b85-9cc3e2332e43" authority="(Beauvais, 1986)" baseAuthorityName="Beauvais" baseAuthorityYear="1986" box="[869,1399,1235,1263]" class="Anthozoa" family="Epismiliidae" genus="Neorylstonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Scleractinia" pageId="49" pageNumber="1234" phylum="Cnidaria" rank="species" species="pseudocolumellata">
<emphasis id="B97FEAA648723103993D65CAFB7AFB3E" bold="true" box="[869,1211,1235,1262]" italics="true" pageId="49" pageNumber="1234">Neorylstonia pseudocolumellata</emphasis>
(
<bibRefCitation id="EF9A4B45487231039E9165CDFAAFFB3F" author="BEAUVAIS L." box="[1225,1390,1236,1263]" pageId="49" pageNumber="1234" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C311653F487231039E6765EDFB5CFADE" box="[1087,1181,1267,1294]" pageId="49" pageNumber="1234" type="description">
<paragraph id="8BB436B4487231039E6765EDFB5CFADE" blockId="49.[869,1399,1235,1294]" box="[1087,1181,1267,1294]" pageId="49" pageNumber="1234">
(
<figureCitation id="13302A31487231039E1065EAFB55FADE" box="[1096,1172,1267,1294]" captionStart="FIG" captionStartId="52.[132,143,1894,1911]" captionTargetBox="[253,1333,215,1877]" captionTargetId="figure-17@52.[253,1334,215,1877]" captionTargetPageId="52" captionText="FIG. 32. — Neorylstonia pseudocolumellata (Beauvais, 1986): A: transverse section of the specimen CPUN AM16164, showing a type 2 pseudocolumella with visible pseudocolumellar septa; B: transverse section of the specimen CPUN AM175 showing a type 1 pseudocolumella; C: transverse section in the specimen CPUN DA2905E1.1 showing the true epitheca surrounding the septal apparatus and the stereome deposits on septa; D: detail of the ghosts of septal structure in transverse section (specimen CPUN 2110A2) and the stereome deposits in “ogives” or “glove fingers”;E: longitudinal section of the specimen MNHN.F.A57331-2 showing the endothecal organization in tabular dissepiments and the axial porosity of the pseudocolumella; F, detail of the parathecal wall in continuity with the tabular dissepiments (longitudinal section of the specimen CPUN 2110A2). Scale bars: A, B, E, F, 5 mm; C, 4 mm; D, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5764467" httpUri="https://zenodo.org/record/5764467/files/figure.png" pageId="49" pageNumber="1234">Fig. 32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311653F4872310399756434FB02FA08" pageId="49" pageNumber="1234" type="materials_examined">
<paragraph id="8BB436B44872310399756434FB02FA08" blockId="49.[813,1455,1324,1497]" pageId="49" pageNumber="1234">
<materialsCitation id="3B633CE94872310399756434FB02FA08" collectionCode="TYPE, MNHN" country="Morocco" location="Beauvais" pageId="49" pageNumber="1234" specimenCount="4" typeStatus="holotype">
<collectionCode id="ED1AAE714872310399756434FC9CFA92" box="[813,861,1325,1348]" pageId="49" pageNumber="1234">TYPE</collectionCode>
MATERIAL. —
<typeStatus id="54B088164872310399A96435FB90FA94" box="[1009,1105,1324,1348]" pageId="49" pageNumber="1234" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED1AAE71487231039E046434FB71FA94" box="[1116,1200,1325,1348]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="49" pageNumber="1234">MNHN</collectionCode>
.F.R11604 (original designation),
<bibRefCitation id="EF9A4B454872310399DA645EFBFBFA8F" author="BEAUVAIS L." box="[898,1082,1351,1375]" pageId="49" pageNumber="1234" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">
<location id="8ED4606F4872310399DA645EFC17FA8F" LSID="urn:lsid:plazi:treatment:03A287A248723107993D65CAFED4FC3F:8ED4606F4872310399DA645EFC17FA8F" box="[898,982,1351,1375]" country="Morocco" name="Beauvais" pageId="49" pageNumber="1234">Beauvais</location>
1986: 20
</bibRefCitation>
, pl. 4, fig. 1a-1b;
<typeStatus id="54B08816487231039EB4645EFA89FA8F" box="[1260,1352,1351,1375]" pageId="49" pageNumber="1234" type="paratype">paratypes</typeStatus>
figured in
<bibRefCitation id="EF9A4B454872310399766478FC7BFAA9" author="BEAUVAIS L." box="[814,954,1377,1401]" pageId="49" pageNumber="1234" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">
<location id="8ED4606F4872310399766478FC43FAA9" LSID="urn:lsid:plazi:treatment:03A287A248723107993D65CAFED4FC3F:8ED4606F4872310399766478FC43FAA9" box="[814,898,1377,1401]" country="Morocco" name="Beauvais" pageId="49" pageNumber="1234">Beauvais</location>
1986
</bibRefCitation>
: pl. 4, fig. 1c, d and e.
<typeStatus id="54B08816487231039975648BFC9CFA77" box="[813,861,1426,1449]" pageId="49" pageNumber="1234">TYPE</typeStatus>
HORIZON. —
<location id="8ED4606F4872310399B46488FBB7FA79" LSID="urn:lsid:plazi:treatment:03A287A248723107993D65CAFED4FC3F:8ED4606F4872310399B46488FBB7FA79" box="[1004,1142,1425,1449]" country="Morocco" name="Pliensbachian" pageId="49" pageNumber="1234">Pliensbachian</location>
.
<typeStatus id="54B0881648723103997564D8FC9CFA06" box="[813,861,1473,1496]" pageId="49" pageNumber="1234">TYPE</typeStatus>
LOCALITY. —
<location id="8ED4606F4872310399B364D8FB9AFA09" LSID="urn:lsid:plazi:treatment:03A287A248723107993D65CAFED4FC3F:8ED4606F4872310399B364D8FB9AFA09" box="[1003,1115,1473,1497]" country="Morocco" name="Beni Tadjit" pageId="49" pageNumber="1234">Beni Tadjit</location>
,
<collectingCountry id="F31C7624487231039E3C64D8FB7EFA08" box="[1124,1215,1473,1496]" name="Morocco" pageId="49" pageNumber="1234">Morocco</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311653F48723103997564E9FB7FF982" pageId="49" pageNumber="1234" type="reference_group">
<paragraph id="8BB436B448723103997564E9FA71F9D8" blockId="49.[813,1456,1520,1544]" box="[813,1456,1520,1544]" pageId="49" pageNumber="1234">
<taxonomicName id="4C0B4D3748723103997564E9FAD4F9D8" authority="Beauvais, 1986: 20" authorityName="Beauvais" authorityPageNumber="20" authorityYear="1986" box="[813,1301,1520,1544]" class="Florideophyceae" family="Hapalidiaceae" genus="Mesophyllum" kingdom="Plantae" order="Corallinales" pageId="49" pageNumber="1234" phylum="Rhodophyta" rank="species" species="pseudocolumellatum">
<emphasis id="B97FEAA648723103997564E9FB9EF9D8" box="[813,1119,1520,1544]" italics="true" pageId="49" pageNumber="1234">Mesophyllum pseudocolumellatum</emphasis>
<bibRefCitation id="EF9A4B45487231039E3964E9FAD4F9D8" author="BEAUVAIS L." box="[1121,1301,1520,1544]" pageId="49" pageNumber="1234" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986: 20</bibRefCitation>
</taxonomicName>
, pl. 4, fig. 1a-1b.
</paragraph>
<paragraph id="8BB436B44872310399756739FB7FF982" blockId="49.[812,1455,1568,1618]" pageId="49" pageNumber="1234">
<taxonomicName id="4C0B4D374872310399756739FB91F9E8" ID-CoL="d11bfea7-f4f3-4074-8b85-9cc3e2332e43" baseAuthorityName="Beauvais" baseAuthorityYear="1986" box="[813,1104,1568,1592]" class="Anthozoa" family="Epismiliidae" genus="Neorylstonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Scleractinia" pageId="49" pageNumber="1234" phylum="Cnidaria" rank="species" species="pseudocolumellata">
<emphasis id="B97FEAA64872310399756739FB91F9E8" box="[813,1104,1568,1592]" italics="true" pageId="49" pageNumber="1234">Neorylstonia pseudocolumellata</emphasis>
</taxonomicName>
<bibRefCitation id="EF9A4B45487231039E306739FA8CF9E8" author="VASSEUR R." box="[1128,1357,1568,1592]" pageId="49" pageNumber="1234" refId="ref55769" refString="VASSEUR R. 2018. - Extinctions et recouvrements de coraux au cours de la crise Pliensbachien-Toarcien. These de doctorat, Universite de Lorraine, France, 542 p." type="book" year="2018">Vasseur 2018: 259-260</bibRefCitation>
, fig. 3.63;
<bibRefCitation id="EF9A4B454872310399746722FBE7F982" author="VASSEUR R. &amp; BOIVIN S. &amp; LATHUILIERE B. &amp; LAZAR I. &amp; DURLET C. &amp; MARTINDALE R. - C. &amp; BODIN S. &amp; ELHMIDI K." box="[812,1062,1594,1618]" pageId="49" pageNumber="1234" pagination="2" refId="ref55800" refString="VASSEUR R., BOIVIN S., LATHUILIERE B., LAZAR I., DURLET C., MARTINDALE R. - C., BODIN S. &amp; ELHMIDI K. 2019. - Lower Jurassic corals from Morocco with skeletal structures convergent with those of Paleozoic rugosan corals. Palaeontologia Electronica 22 (2). https: // doi. org / 10.26879 / 874" type="journal article" year="2019">
Vasseur
<emphasis id="B97FEAA64872310399246722FC6AF982" box="[892,939,1594,1618]" italics="true" pageId="49" pageNumber="1234">et al.</emphasis>
2019: 9 -17
</bibRefCitation>
, fig. 2- 6, 13.6.
</paragraph>
</subSubSection>
<subSubSection id="C311653F4872310399756773FA71F8CB" pageId="49" pageNumber="1234" type="materials_examined">
<paragraph id="8BB436B44872310399756773FBBDF94D" blockId="49.[813,1455,1642,1693]" pageId="49" pageNumber="1234">
STUDIED SAMPLES. —
<specimenCount id="9D0DFD3D4872310399A36773FBB8F952" box="[1019,1145,1642,1666]" pageId="49" pageNumber="1234" type="generic">91 specimens</specimenCount>
described in
<bibRefCitation id="EF9A4B45487231039EA96773FA6EF952" author="VASSEUR R. &amp; BOIVIN S. &amp; LATHUILIERE B. &amp; LAZAR I. &amp; DURLET C. &amp; MARTINDALE R. - C. &amp; BODIN S. &amp; ELHMIDI K." box="[1265,1455,1642,1666]" pageId="49" pageNumber="1234" pagination="2" refId="ref55800" refString="VASSEUR R., BOIVIN S., LATHUILIERE B., LAZAR I., DURLET C., MARTINDALE R. - C., BODIN S. &amp; ELHMIDI K. 2019. - Lower Jurassic corals from Morocco with skeletal structures convergent with those of Paleozoic rugosan corals. Palaeontologia Electronica 22 (2). https: // doi. org / 10.26879 / 874" type="journal article" year="2019">
Vasseur
<emphasis id="B97FEAA6487231039F666772FAABF952" box="[1342,1386,1642,1666]" italics="true" pageId="49" pageNumber="1234">et al.</emphasis>
(2019)
</bibRefCitation>
(collections MNHN and CPUN).
</paragraph>
<paragraph id="8BB436B448723103997567ADFA71F8CB" blockId="49.[813,1456,1716,1819]" pageId="49" pageNumber="1234">
GEOGRAPHIC AND STRATIGRAPHIC RANGE. — Sinemurian of Amellagou (Jebel Serdrar, High Atlas) and Pliensbachian of Amellagou, Dades Valley, Koudiat Ech Chehem and Taililout (High and Middle Atlas) in
<collectingCountry id="F31C76244872310399FE661DFBC0F8CB" box="[934,1025,1796,1819]" name="Morocco" pageId="49" pageNumber="1234">Morocco</collectingCountry>
; Pliensbachian of
<collectingCountry id="F31C7624487231039EEB661AFB1FF8CB" box="[1203,1246,1795,1819]" name="Italy" pageId="49" pageNumber="1234">Italy</collectingCountry>
(Caloveto,
<collectingRegion id="49CFF856487231039F16661AFA64F8CB" box="[1358,1445,1795,1819]" country="Italy" name="Calabria" pageId="49" pageNumber="1234">Calabria</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C311653F487231079975664BFED4FC3F" lastPageId="53" lastPageNumber="1238" pageId="49" pageNumber="1234" type="description">
<paragraph id="8BB436B4487231039975664BFC7CF8BA" blockId="49.[811,1456,1874,2028]" box="[813,957,1874,1900]" pageId="49" pageNumber="1234">DESCRIPTION</paragraph>
<paragraph id="8BB436B44872310799756668FD7EFDE1" blockId="49.[811,1456,1874,2028]" lastBlockId="53.[130,777,215,848]" lastPageId="53" lastPageNumber="1238" pageId="49" pageNumber="1234">
Solitary coral. Corallum ceratoid, straight or curved. The calice is subcircular in outline. Costosepta straight or slightly curved (possibly post-mortem distortion), free, compact, occasionally rhopaloid. Some first-order septa (S1) rarely reach the axial structure but in most cases the inner edge abruptly dips in the fossa. Lateral faces and distal edge are smooth. No pali. Radial elements are commonly arranged in three distinct size orders with different length and width in transverse sections but S1 and S2 are occasionally indistinct because of similar sizes. In contrast, some other samples show clearly four distinguishable size orders of septa. In the latter case, the first and second size orders geometrically correspond to a subdivision of the S1. Lastly, the S3 are sometimes missing. On the basis of the described arrangement of septal length and thicknesses a very classical model of septal insertion can be proposed, namely 96 septa distributed in five ontogenetical cycles according to 6+6+12+24+48 septa. This model of septal insertion is statistically grounded by univariate statistical analysis in which the number 96 shows a well-marked mode (
<bibRefCitation id="EF9A4B45487031019E8167AAFA5EF91C" author="VASSEUR R. &amp; BOIVIN S. &amp; LATHUILIERE B. &amp; LAZAR I. &amp; DURLET C. &amp; MARTINDALE R. - C. &amp; BODIN S. &amp; ELHMIDI K." box="[1241,1439,1714,1741]" pageId="51" pageNumber="1236" pagination="2" refId="ref55800" refString="VASSEUR R., BOIVIN S., LATHUILIERE B., LAZAR I., DURLET C., MARTINDALE R. - C., BODIN S. &amp; ELHMIDI K. 2019. - Lower Jurassic corals from Morocco with skeletal structures convergent with those of Paleozoic rugosan corals. Palaeontologia Electronica 22 (2). https: // doi. org / 10.26879 / 874" type="journal article" year="2019">
Vasseur
<emphasis id="B97FEAA6487031019F7767AAFAA3F91C" box="[1327,1378,1714,1740]" italics="true" pageId="51" pageNumber="1236">et al.</emphasis>
2019
</bibRefCitation>
). Ghosts of septal structures can sometimes be observed in thin sections. They consist in transverse section of a darker axial zone within the septum, a brighter zigzag line or some dark spots in a line considered as remains of “centers of calcification” or centers of rapid accretion (
<bibRefCitation id="EF9A4B45487031019EF06648FA85F8BC" author="STOLARSKI J." box="[1192,1348,1873,1900]" pageId="51" pageNumber="1236" pagination="497 - 530" refId="ref54971" refString="STOLARSKI J. 2003. - Three-dimensional micro- and nanostructural characteristics of the scleractinian coral skeleton: a biocalcification proxy. Acta Palaeontologica Polonica 48: 497 - 530." type="journal article" year="2003">Stolarski 2003</bibRefCitation>
). Growth lines in “ogive” or “gloves fingers” can be seen in longitudinal sections, resulting from the deposition of different successive layers of stereome thickening the septa (thickening deposits of
<bibRefCitation id="EF9A4B4548703101991466C8FC2DF83B" author="STOLARSKI J." box="[844,1004,2001,2027]" pageId="51" pageNumber="1236" pagination="497 - 530" refId="ref54971" refString="STOLARSKI J. 2003. - Three-dimensional micro- and nanostructural characteristics of the scleractinian coral skeleton: a biocalcification proxy. Acta Palaeontologica Polonica 48: 497 - 530." type="journal article" year="2003">Stolarski 2003</bibRefCitation>
). Axial structure present, in the form of a Rugose-like calicular boss made of septa and endotheca. According to
<bibRefCitation id="EF9A4B45487631079B4E61EEFD63FEC1" author="MILNE EDWARDS H. &amp; HAIME J." box="[278,674,247,274]" pageId="53" pageNumber="1238" pagination="465 - 469" refId="ref53302" refString="MILNE EDWARDS H. &amp; HAIME J. 1848. - Observations sur les polypiers de la famille des astreides. Comptes rendus hebdomadaires des Seances de l'Academie des Sciences 27: 465 - 469. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 29840 / f 469. item" type="journal article" year="1848">Milne Edwards &amp; Haime (1848: 78)</bibRefCitation>
and later
<bibRefCitation id="EF9A4B45487631079ADC600EFE6EFEE2" author="BERKOWSKI B. &amp; WEYER D." box="[132,431,279,306]" pageId="53" pageNumber="1238" pagination="245 - 253" refId="ref49134" refString="BERKOWSKI B. &amp; WEYER D. 2012. - Hamaraxonia, a new pseudocolumellate genus of Middle Devonian deep-water Rugosa (Anthozoa) from Morocco, in DENAYER J., ARETZ M. &amp; POTY E. (eds), Proceedings of the XIth International Symposium on Fossil Cnidaria and Porifera, Geologica Belgica 15 (4): 245 - 253. https: // popups. uliege. be / 1374 - 8505 / index. php? id = 3753" type="journal article" year="2012">Berkowski &amp; Weyer (2012)</bibRefCitation>
it corresponds to the definition of a pseudocolumella. This pseudocolumella presents different morphologies during the polyp life, giving the fossils a very characteristic aspect in transverse section. It is made of 16 to 26 septa not organized in a strict radial symmetry, more often 21 to 22, organized even in two crescents separated by an axial cavity (
<typeStatus id="54B08816487631079B7060CFFE99FE20" box="[296,344,470,496]" pageId="53" pageNumber="1238">type</typeStatus>
2) or in the form of a star with opposite septa linked in the calicular axial zone (
<typeStatus id="54B0881648763107984160EFFD89FDC0" box="[537,584,502,528]" pageId="53" pageNumber="1238">type</typeStatus>
1). Between these two extreme morphologies are some transition states.
</paragraph>
<caption id="DF74663C487131009ADC66B3FBE1F839" ID-DOI="http://doi.org/10.5281/zenodo.5764461" ID-Zenodo-Dep="5764461" httpUri="https://zenodo.org/record/5764461/files/figure.png" pageId="50" pageNumber="1235" startId="50.[132,143,1962,1979]" targetBox="[135,1451,215,1828]" targetPageId="50">
<paragraph id="8BB436B4487131009ADC66B3FBE1F839" blockId="50.[132,1457,1962,2025]" pageId="50" pageNumber="1235">
FIG. 30. —
<taxonomicName id="4C0B4D37487131009A8766B3FB8AF86B" authority="Boivin, Vasseur, Lathuiliere, Lazar, Durlet, Martindale, El Hmidi &amp; Martini, 2019" authorityName="Boivin, Vasseur, Lathuiliere, Lazar, Durlet, Martindale, El Hmidi &amp; Martini" authorityYear="2019" box="[223,1099,1962,1980]" class="Anthozoa" family="Amphiastraeidae" genus="Hispaniastraea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="50" pageNumber="1235" phylum="Cnidaria" rank="species" species="ousriorum">
<emphasis id="B97FEAA6487131009A8766B3FE72F86B" box="[223,435,1962,1979]" italics="true" pageId="50" pageNumber="1235">Hispaniastraea ousriorum</emphasis>
<bibRefCitation id="EF9A4B45487131009BE066B3FB8AF86B" author="BOIVIN S. &amp; VASSEUR R. &amp; LATHUILIERE B. &amp; LAZAR I. &amp; DURLET C. &amp; MARTINDALE R. C. &amp; EL HMIDI K. &amp; MARTINI R." box="[440,1099,1962,1980]" pageId="50" pageNumber="1235" pagination="1 - 24" refId="ref49395" refString="BOIVIN S., VASSEUR R., LATHUILIERE B., LAZAR I., DURLET C., MARTINDALE R. C., EL HMIDI K. &amp; MARTINI R. 2019 - A little walk between Early Jurassic sponges and corals: A confusing morphological convergence. Geobios 57: 1 - 24. https: // doi. org / 10.1016 / j. geobios. 2019.10.001" type="journal article" year="2019">Boivin, Vasseur, Lathuilière, Lazăr, Durlet, Martindale, El Hmidi &amp; Martini, 2019</bibRefCitation>
</taxonomicName>
:
<emphasis id="B97FEAA6487131009E0A66B3FB9EF86B" bold="true" box="[1106,1119,1962,1979]" pageId="50" pageNumber="1235">A</emphasis>
, specimen CPUN CDAm1;
<emphasis id="B97FEAA6487131009F1E66B3FA92F86B" bold="true" box="[1350,1363,1962,1979]" pageId="50" pageNumber="1235">B</emphasis>
, specimen CPUN AM16188-5;
<emphasis id="B97FEAA6487131009B7166D8FEF6F802" bold="true" box="[297,311,1985,2002]" pageId="50" pageNumber="1235">C</emphasis>
, transverse polished section of the specimen CPUN AM16160-3;
<emphasis id="B97FEAA648713100990D66D8FCA2F802" bold="true" box="[853,867,1985,2002]" pageId="50" pageNumber="1235">D</emphasis>
, distal view of the calicular surface of the specimen CPUN AM16188-5;
<emphasis id="B97FEAA6487131009ADC66C1FF51F839" bold="true" box="[132,144,2008,2025]" pageId="50" pageNumber="1235">E</emphasis>
, transverse thin section of the specimen CPUN CDAm1. Scale bars: A, 5 mm; B, 1 cm; C, 2 mm; D, E, 4 mm.
</paragraph>
</caption>
<caption id="DF74663C487031019ADC6717FBBDF9E6" ID-DOI="http://doi.org/10.5281/zenodo.5764465" ID-Zenodo-Dep="5764465" httpUri="https://zenodo.org/record/5764465/files/figure.png" pageId="51" pageNumber="1236" startId="51.[132,143,1550,1567]" targetBox="[134,1454,218,1508]" targetPageId="51">
<paragraph id="8BB436B4487031019ADC6717FBBDF9E6" blockId="51.[132,1455,1550,1590]" pageId="51" pageNumber="1236">
FIG. 31. — Gen. et sp. indet.:
<emphasis id="B97FEAA6487031019B2B6717FE41F9CF" bold="true" box="[371,384,1550,1567]" pageId="51" pageNumber="1236">A</emphasis>
, distal view of the specimen CPUN DA3105E1;
<emphasis id="B97FEAA64870310199506717FCD4F9CF" bold="true" box="[776,789,1550,1567]" pageId="51" pageNumber="1236">B</emphasis>
, detail in longitudinal thin section of pennular structures of septa in the specimen CPUN DA3105E1;
<emphasis id="B97FEAA6487031019B79673CFEEEF9E6" bold="true" box="[289,303,1573,1590]" pageId="51" pageNumber="1236">C</emphasis>
, transverse thin section in the specimen CPUN DA3105E1. Scale bars: A, 3 cm; B, 0.5 mm; C, 4 mm.
</paragraph>
</caption>
<caption id="DF74663C487731069ADC667FFB4AF839" ID-DOI="http://doi.org/10.5281/zenodo.5764467" ID-Zenodo-Dep="5764467" httpUri="https://zenodo.org/record/5764467/files/figure.png" pageId="52" pageNumber="1237" startId="52.[132,143,1894,1911]" targetBox="[253,1333,215,1877]" targetPageId="52">
<paragraph id="8BB436B4487731069ADC667FFB4AF839" blockId="52.[132,1456,1894,2025]" pageId="52" pageNumber="1237">
FIG. 32. —
<taxonomicName id="4C0B4D37487731069AB8667FFDBEF8A7" authority="(Beauvais, 1986)" baseAuthorityName="Beauvais" baseAuthorityYear="1986" box="[224,639,1894,1911]" class="Anthozoa" family="Epismiliidae" genus="Neorylstonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Scleractinia" pageId="52" pageNumber="1237" phylum="Cnidaria" rank="species" species="pseudocolumellata">
<emphasis id="B97FEAA6487731069AB8667FFE2AF8A7" box="[224,491,1894,1911]" italics="true" pageId="52" pageNumber="1237">Neorylstonia pseudocolumellata</emphasis>
(
<bibRefCitation id="EF9A4B45487731069BAD667FFDBBF8A7" author="BEAUVAIS L." box="[501,634,1894,1911]" pageId="52" pageNumber="1237" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986</bibRefCitation>
)
</taxonomicName>
:
<emphasis id="B97FEAA64877310698DF667FFD55F8A7" bold="true" box="[647,660,1894,1911]" pageId="52" pageNumber="1237">A</emphasis>
: transverse section of the specimen CPUN AM16164, showing a type 2 pseudocolumella with visible pseudocolumellar septa;
<emphasis id="B97FEAA6487731069BC86664FE5CF85E" bold="true" box="[400,413,1917,1934]" pageId="52" pageNumber="1237">B</emphasis>
: transverse section of the specimen CPUN AM175 showing a type 1 pseudocolumella;
<emphasis id="B97FEAA6487731069E2D6664FB42F85E" bold="true" box="[1141,1155,1917,1934]" pageId="52" pageNumber="1237">C</emphasis>
: transverse section in the specimen CPUN DA2905E1.1 showing the true epitheca surrounding the septal apparatus and the stereome deposits on septa;
<emphasis id="B97FEAA6487731069E15668DFB9AF875" bold="true" box="[1101,1115,1940,1957]" pageId="52" pageNumber="1237">D</emphasis>
: detail of the ghosts of septal structure in transverse section (specimen CPUN 2110A2) and the stereome deposits in “ogives” or “glove fingers”;
<emphasis id="B97FEAA648773106999666B3FC1BF86B" bold="true" box="[974,986,1962,1979]" pageId="52" pageNumber="1237">E</emphasis>
: longitudinal section of the specimen MNHN.F.A57331-2 showing the endothecal organization in tabular dissepiments and the axial porosity of the pseudocolumella;
<emphasis id="B97FEAA6487731069E5D66D8FBD1F802" bold="true" box="[1029,1040,1985,2002]" pageId="52" pageNumber="1237">F</emphasis>
, detail of the parathecal wall in continuity with the tabular dissepiments (longitudinal section of the specimen CPUN 2110A2). Scale bars: A, B, E, F, 5 mm; C, 4 mm; D, 2 mm.
</paragraph>
</caption>
<paragraph id="8BB436B4487631079AC3632EFDF9FCDF" blockId="53.[130,777,215,848]" pageId="53" pageNumber="1238">In the main cases, hexameral symmetry can be established. The superimposed bilateral symmetry is evidenced by the spatial organization of the axial structure. Endotheca abundant, made of tabular dissepiments regularly arranged in the axial zone. Parathecal wall. Paratheca is continuous with the dissepiments. Synapticulae absent. A true epitheca with growth striae and wrinkles encloses the corallite.</paragraph>
<paragraph id="8BB436B4487631079AC3620CFCC6FC80" blockId="53.[130,777,215,848]" pageId="53" pageNumber="1238">
Calicular diameter:
<quantity id="4CF39B51487631079B28620CFE31FCE0" box="[368,496,789,816]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.9" metricValueMin="0.9" pageId="53" pageNumber="1238" unit="mm" value="19.0" valueMax="29.0" valueMin="9.0">9 to 29 mm</quantity>
Number of septa: 41 to 104 (mostly 96) Septal density: 5 to 10 S1 + S2 for
<quantity id="4CF39B514876310798E7622CFCC2FC80" box="[703,771,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="1238" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BB436B4487631079ADC626CFE13FC5D" blockId="53.[130,775,885,1007]" box="[132,466,885,911]" pageId="53" pageNumber="1238">SIMILARITIES AND DIFFERENCES</paragraph>
<paragraph id="8BB436B4487631079ADC628CFED4FC3F" blockId="53.[130,775,885,1007]" pageId="53" pageNumber="1238">
This taxon is the single case of post-paleozoic coral with a pseudocolumella
<taxonomicName id="4C0B4D37487631079B6462ACFD54FC1F" authority="Berkowski &amp; Weyer (2012)" authorityName="Berkowski &amp; Weyer" authorityYear="2012" box="[316,661,949,975]" pageId="53" pageNumber="1238" rank="species" species="sensu">
<emphasis id="B97FEAA6487631079B6462ACFEB1FC1E" box="[316,368,949,974]" italics="true" pageId="53" pageNumber="1238">sensu</emphasis>
<bibRefCitation id="EF9A4B45487631079B2C62ACFD54FC1F" author="BERKOWSKI B. &amp; WEYER D." box="[372,661,949,975]" pageId="53" pageNumber="1238" pagination="245 - 253" refId="ref49134" refString="BERKOWSKI B. &amp; WEYER D. 2012. - Hamaraxonia, a new pseudocolumellate genus of Middle Devonian deep-water Rugosa (Anthozoa) from Morocco, in DENAYER J., ARETZ M. &amp; POTY E. (eds), Proceedings of the XIth International Symposium on Fossil Cnidaria and Porifera, Geologica Belgica 15 (4): 245 - 253. https: // popups. uliege. be / 1374 - 8505 / index. php? id = 3753" type="journal article" year="2012">Berkowski &amp; Weyer (2012)</bibRefCitation>
</taxonomicName>
referenced up until now.
</paragraph>
</subSubSection>
</treatment>
</document>