227 lines
19 KiB
XML
227 lines
19 KiB
XML
<document ID-DOI="10.5252/geodiversitas2021v43a22" ID-ISSN="1638-9395" ID-Zenodo-Dep="5764379" ID-ZooBank="urn:lsid:zoobank.org:pub:6D1643AD-66A5-4678-9397-CE08E610D641" approvalRequired="387" approvalRequired_for_taxonomicNames="20" approvalRequired_for_textStreams="367" checkinTime="1638890191441" checkinUser="felipe" docAuthor="Vasseur, Raphaël & Lathuilière, Bernard" docDate="2021" docId="03A287A2484A313B9B45640DFA51FC3E" docLanguage="en" docName="Geodiversitas.43.22.1187-1291.pdf" docOrigin="Geodiversitas 43 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Allocoenia undetermined" docType="treatment" docVersion="6" lastPageNumber="1194" masterDocId="FF9BFFDA484331329A586119FFC1FFD0" masterDocTitle="Pliensbachian corals from the Western Tethys" masterLastPageNumber="1291" masterPageNumber="1187" pageNumber="1194" updateTime="1639152608618" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pliensbachian corals from the Western Tethys</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vasseur, Raphaël</mods:namePart>
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<mods:affiliation>Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<mods:nameIdentifier type="email">raphaelvasseur@wanadoo.fr</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lathuilière, Bernard</mods:namePart>
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<mods:affiliation>Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<mods:nameIdentifier type="email">bernard.lathuiliere@univ-lorraine.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Geodiversitas</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>43</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>22</mods:number>
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</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1187</mods:start>
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<mods:end>1291</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/geodiversitas2021v43a22</mods:identifier>
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<mods:identifier type="ISSN">1638-9395</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">5764379</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:6D1643AD-66A5-4678-9397-CE08E610D641</mods:identifier>
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||
</mods:mods>
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<subSection pageId="9" pageNumber="1194" type="multiple">
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5764986" ID-Zenodo-Dep="5764986" LSID="urn:lsid:plazi:treatment:03A287A2484A313B9B45640DFA51FC3E" httpUri="http://treatment.plazi.org/id/03A287A2484A313B9B45640DFA51FC3E" lastPageNumber="1194" pageId="9" pageNumber="1194">
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<subSubSection box="[285,622,1299,1326]" pageId="9" pageNumber="1194" type="nomenclature">
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<paragraph blockId="9.[285,622,1299,1326]" box="[285,622,1299,1326]" pageId="9" pageNumber="1194">
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<heading box="[285,622,1299,1326]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1194" reason="2">
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Genus
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<taxonomicName authority="Etallon, 1859" box="[362,622,1299,1326]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[362,468,1299,1325]" italics="true" pageId="9" pageNumber="1194">Allocoenia</emphasis>
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Etallon, 1859
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="1194" type="type_taxon">
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<paragraph blockId="9.[132,777,1356,1407]" pageId="9" pageNumber="1194">
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SPECIES
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<collectionCode box="[216,331,1362,1379]" pageId="9" pageNumber="1194">ORIGINALLY</collectionCode>
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INCLUDED. —
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<taxonomicName authority="Michelin, 1847" authorityName="Michelin" authorityYear="1847" class="Anthozoa" family="Merulinidae" genus="Astrea" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[491,672,1356,1380]" italics="true" pageId="9" pageNumber="1194">Astrea trochiformis</emphasis>
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Michelin, 1847
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</taxonomicName>
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,
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<taxonomicName authority="Etallon, 1859" authorityName="Etallon" authorityYear="1859" box="[196,507,1383,1407]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[196,364,1383,1407]" italics="true" pageId="9" pageNumber="1194">Allocoenia furcata</emphasis>
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Etallon, 1859
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="9.[132,775,1430,1481]" pageId="9" pageNumber="1194">
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<collectionCode box="[132,181,1431,1454]" pageId="9" pageNumber="1194">
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<typeStatus box="[132,181,1431,1454]" pageId="9" pageNumber="1194">TYPE</typeStatus>
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</collectionCode>
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SPECIES. —
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<taxonomicName authority="Etallon, 1859" authorityName="Etallon" authorityYear="1859" box="[301,611,1430,1455]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[301,469,1430,1455]" italics="true" pageId="9" pageNumber="1194">Allocoenia furcata</emphasis>
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Etallon, 1859
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</taxonomicName>
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, subsequent designation by
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<bibRefCitation author="WELLS J. W." box="[249,430,1457,1481]" pageId="9" pageNumber="1194" pagination="97 - 134" refId="ref55962" refString="WELLS J. W. 1936. - The nomenclature and type species of some genera of recent and fossil corals. American Journal of Science (5) 31: 97 - 134. https: // doi. org / 10.2475 / ajs. s 5 - 31.182.97" type="journal article" year="1936">Wells (1936: 100)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="1194" type="discussion">
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<paragraph blockId="9.[131,775,1523,2028]" box="[132,232,1523,1549]" pageId="9" pageNumber="1194">REMARKS</paragraph>
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<paragraph blockId="9.[131,775,1523,2028]" pageId="9" pageNumber="1194">
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<bibRefCitation author="ALLOITEAU J." box="[132,314,1554,1581]" pageId="9" pageNumber="1194" pagination="9 - 16" refId="ref48497" refString="ALLOITEAU J. 1952. - Note sur des polypiers du Senegal. Bulletin de la Direction des Mines 14: 9 - 16." type="journal article" year="1952">Alloiteau (1952)</bibRefCitation>
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and several subsequent authors indicated erroneously
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<taxonomicName authority="Michelin, 1847" authorityName="Michelin" authorityYear="1847" box="[264,593,1586,1613]" class="Anthozoa" family="Merulinidae" genus="Astrea" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[264,418,1586,1613]" italics="true" pageId="9" pageNumber="1194">A. trochiformis</emphasis>
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Michelin, 1847
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</taxonomicName>
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as
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<typeStatus box="[628,674,1587,1613]" pageId="9" pageNumber="1194">type</typeStatus>
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species.
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</paragraph>
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<paragraph blockId="9.[131,775,1523,2028]" pageId="9" pageNumber="1194">
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The differences between the genera
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<taxonomicName authority="d'Orbigny, 1849" authorityName="d'Orbigny" authorityYear="1849" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[533,649,1619,1645]" italics="true" pageId="9" pageNumber="1194">Actinastrea</emphasis>
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d’Orbigny, 1849
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</taxonomicName>
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,
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<taxonomicName authority="Etallon, 1859" authorityName="Etallon" authorityYear="1859" box="[201,463,1650,1677]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[201,308,1650,1676]" italics="true" pageId="9" pageNumber="1194">Allocoenia</emphasis>
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Etallon, 1859
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</taxonomicName>
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,
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<taxonomicName authority="Etallon, 1864" authorityName="Etallon" authorityYear="1864" box="[471,771,1650,1676]" class="Anthozoa" family="Actinastraeidae" genus="Stephanastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[471,613,1650,1676]" italics="true" pageId="9" pageNumber="1194">Stephanastrea</emphasis>
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Etallon, 1864
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</taxonomicName>
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,
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<taxonomicName authority="Tomes, 1893" authorityName="Tomes" authorityYear="1893" box="[131,402,1682,1709]" class="Anthozoa" family="Actinastreidae" genus="Stelidioseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[131,254,1682,1708]" italics="true" pageId="9" pageNumber="1194">Stelidioseris</emphasis>
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Tomes, 1893
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</taxonomicName>
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<taxonomicName authority="Alloiteau, 1956" authorityName="Alloiteau" authorityYear="1956" box="[405,728,1682,1709]" class="Anthozoa" family="Actinastreidae" genus="Allocoeniopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[405,549,1682,1708]" italics="true" pageId="9" pageNumber="1194">Allocoeniopsis</emphasis>
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Alloiteau, 1956
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</taxonomicName>
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and
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<emphasis box="[131,281,1714,1740]" italics="true" pageId="9" pageNumber="1194">Chondrocenia</emphasis>
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<bibRefCitation author="RONIEWICZ E." box="[292,483,1714,1740]" pageId="9" pageNumber="1194" pagination="1 - 152" refId="ref54266" refString="RONIEWICZ E. 1989. - Triassic scleractinian corals of the Zlambach Beds, Northern Calcareous Alps, Austria. Denkschriften-Osterreichische Akademie der Wissenschaften 126: 1 - 152." type="journal article" year="1989">Roniewicz, 1989</bibRefCitation>
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remain very difficult and we propose here an assignment to the genus
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<taxonomicName authorityName="Etallon" authorityYear="1859" box="[625,736,1746,1772]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[625,736,1746,1772]" italics="true" pageId="9" pageNumber="1194">Allocoenia</emphasis>
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</taxonomicName>
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for the following reasons:
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</paragraph>
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<paragraph blockId="9.[131,775,1523,2028]" lastBlockId="9.[811,1457,215,1007]" pageId="9" pageNumber="1194">
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The genus
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<taxonomicName authorityName="d'Orbigny" authorityYear="1849" box="[270,388,1810,1836]" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[270,388,1810,1836]" italics="true" pageId="9" pageNumber="1194">Actinastrea</emphasis>
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</taxonomicName>
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being the most anciently described was defined in the Late Cretaceous and was used in the most lumping practices (see for instance
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<bibRefCitation author="GEYER O. F." box="[589,720,1873,1900]" pageId="9" pageNumber="1194" pagination="317 - 356" refId="ref50901" refString="GEYER O. F. 1955. - Korallen-Faunen aus dem oberen Jura von Portugal. Senckenbergiana Lethaea 35 (5 - 6): 317 - 356." type="journal article" year="1955">Geyer 1955</bibRefCitation>
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who grouped in
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<taxonomicName authorityName="d'Orbigny" authorityYear="1849" box="[260,379,1906,1932]" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[260,379,1906,1932]" italics="true" pageId="9" pageNumber="1194">Actinastrea</emphasis>
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</taxonomicName>
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three species that are today considered as belonging to, at least, two different genera). Despite recent revisions (
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<bibRefCitation author="LOSER H." box="[311,430,1969,1996]" pageId="9" pageNumber="1194" pagination="15 - 22" refId="ref52685" refString="LOSER H. 2012. - Revision of Actinastrea, the most common Cretaceous coral genus. Palaontologische Zeitschrift 86: 15 - 22." type="journal article" year="2012">Löser 2012</bibRefCitation>
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), the status of the genus remains nomenclaturally problematic in the sense that the currently admitted type species
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<taxonomicName authority="d'Orbigny" authorityName="d'Orbigny" box="[1053,1297,215,242]" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[1053,1174,215,242]" italics="true" pageId="9" pageNumber="1194">A. goldfussi</emphasis>
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d’Orbigny
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</taxonomicName>
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was not a part of the originally included nominal species (opposition to article 67-2 ICZN). On a taxonomic point of view, it seems that the
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<typeStatus box="[978,1080,311,337]" pageId="9" pageNumber="1194">lectotype</typeStatus>
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of
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<taxonomicName box="[1116,1237,311,337]" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis box="[1116,1237,311,337]" italics="true" pageId="9" pageNumber="1194">A. goldfussi</emphasis>
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</taxonomicName>
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(if, after all, we use it as a reference) and also the
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<typeStatus box="[1142,1244,343,369]" pageId="9" pageNumber="1194">lectotype</typeStatus>
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of
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<taxonomicName authorityName="Tomes" authorityYear="1893" class="Anthozoa" family="Actinastreidae" genus="Stelidioseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="species">
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<emphasis italics="true" pageId="9" pageNumber="1194">Stelidioseris gibbosa</emphasis>
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</taxonomicName>
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(another probable species of Late Cretaceous) show septal strong granules near the wall. In
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<taxonomicName authorityName="Tomes" authorityYear="1893" box="[1259,1384,406,432]" class="Anthozoa" family="Actinastreidae" genus="Stelidioseris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[1259,1384,406,432]" italics="true" pageId="9" pageNumber="1194">Stelidioseris</emphasis>
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</taxonomicName>
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, these granules have been misinterpreted by
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<bibRefCitation author="LOSER H." box="[1252,1398,439,465]" pageId="9" pageNumber="1194" pagination="15 - 22" refId="ref52685" refString="LOSER H. 2012. - Revision of Actinastrea, the most common Cretaceous coral genus. Palaontologische Zeitschrift 86: 15 - 22." type="journal article" year="2012">Löser (2012)</bibRefCitation>
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who considered these septal parts, trabecular in nature and dark in thin section, as lacunae. We cannot assign the Liassic samples to these taxa and prefer to place temporarily our samples within
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<emphasis box="[984,1103,566,592]" italics="true" pageId="9" pageNumber="1194">
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<taxonomicName authorityName="Etallon" authorityYear="1859" box="[984,1098,566,592]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">Allocoenia</taxonomicName>
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,
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</emphasis>
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a genus that would also deserve a revision.
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</paragraph>
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<paragraph blockId="9.[811,1457,215,1007]" pageId="9" pageNumber="1194">
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The groupings proposed by
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<bibRefCitation author="GEYER O. F." box="[1120,1262,630,657]" pageId="9" pageNumber="1194" pagination="317 - 356" refId="ref50901" refString="GEYER O. F. 1955. - Korallen-Faunen aus dem oberen Jura von Portugal. Senckenbergiana Lethaea 35 (5 - 6): 317 - 356." type="journal article" year="1955">Geyer (1955)</bibRefCitation>
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that placed within
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<taxonomicName authorityName="d'Orbigny" authorityYear="1849" box="[811,925,662,688]" class="Anthozoa" family="Actinastreidae" genus="Actinastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[811,925,662,688]" italics="true" pageId="9" pageNumber="1194">Actinastrea</emphasis>
|
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</taxonomicName>
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the
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<typeStatus box="[976,1022,662,688]" pageId="9" pageNumber="1194">type</typeStatus>
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species of
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<taxonomicName authority="Etallon, 1862" authorityName="Etallon" authorityYear="1862" box="[1140,1407,662,688]" genus="Coenastrea" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[1140,1252,662,688]" italics="true" pageId="9" pageNumber="1194">Coenastrea</emphasis>
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Etallon, 1862
|
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</taxonomicName>
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and
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<taxonomicName authority="d'Orbigny, 1849" authorityName="d'Orbigny" authorityYear="1849" box="[813,1130,693,720]" class="Anthozoa" family="Actinastreidae" genus="Enallocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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||
<emphasis box="[813,945,693,719]" italics="true" pageId="9" pageNumber="1194">Enallocoenia</emphasis>
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d’Orbigny, 1849
|
||
</taxonomicName>
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seem immoderate. The occurrence of S2 with paliform inner edge, with S3 tending to join on S2 (the septa are free in both latter genera), and globally smaller calices are characteristic for the forms named
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<taxonomicName authorityName="Etallon" authorityYear="1859" box="[1350,1453,789,815]" class="Anthozoa" family="Actinastreidae" genus="Allocoenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[1350,1453,789,815]" italics="true" pageId="9" pageNumber="1194">Allocoenia</emphasis>
|
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</taxonomicName>
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in the present work.
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</paragraph>
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<paragraph blockId="9.[811,1457,215,1007]" pageId="9" pageNumber="1194">
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These forms are distinguished from
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<taxonomicName authority="Duncan, 1867" authorityName="Duncan" authorityYear="1867" class="Anthozoa" family="Pinacophyllidae" genus="Cyathocoenia" higherTaxonomySource="CoL" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
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<emphasis box="[1216,1353,853,879]" italics="true" pageId="9" pageNumber="1194">Cyathocoenia</emphasis>
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Duncan, 1867
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</taxonomicName>
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by the occurrence of a styliform columella.
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</paragraph>
|
||
<paragraph blockId="9.[811,1457,215,1007]" pageId="9" pageNumber="1194">
|
||
In the end, the genus differs from
|
||
<taxonomicName authority="Milne Edward & Haime, 1851" authorityName="Milne Edwards & Haime" authorityYear="1851" class="Anthozoa" family="Thecosmiliidae" genus="Isastrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="9" pageNumber="1194" phylum="Cnidaria" rank="genus">
|
||
<emphasis box="[1191,1266,918,943]" italics="true" pageId="9" pageNumber="1194">Isastrea</emphasis>
|
||
Milne Edward & Haime, 1851
|
||
</taxonomicName>
|
||
by its straight septa with an often marked angle between confluent septa and the presence of a columella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |