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<document id="D053EDD2A88CBACB07BE163B30610DF3" ID-DOI="10.5252/geodiversitas2021v43a22" ID-ISSN="1638-9395" ID-Zenodo-Dep="5764379" ID-ZooBank="urn:lsid:zoobank.org:pub:6D1643AD-66A5-4678-9397-CE08E610D641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1638890191441" checkinUser="felipe" docAuthor="Vasseur, Raphaël &amp; Lathuilière, Bernard" docDate="2021" docId="03A287A24800317699FA640DFF3BF83B" docLanguage="en" docName="Geodiversitas.43.22.1187-1291.pdf" docOrigin="Geodiversitas 43 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Stylosmilia dresnayi Milne Edwards &amp; Haime 1848" docType="treatment" docVersion="9" lastPageNumber="1253" masterDocId="FF9BFFDA484331329A586119FFC1FFD0" masterDocTitle="Pliensbachian corals from the Western Tethys" masterLastPageNumber="1291" masterPageNumber="1187" pageNumber="1252" updateTime="1699441156090" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="DECD15350873F58E46C3B491DD0781BC">Pliensbachian corals from the Western Tethys</mods:title>
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<mods:namePart id="77528E7CBB35929D90A01748FBDDAD40">Vasseur, Raphaël</mods:namePart>
<mods:affiliation id="3215550A0121FA4B6A546CC2216EDE3E">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<mods:namePart id="8F3420BDB238D263FB109BFB4B23C630">Lathuilière, Bernard</mods:namePart>
<mods:affiliation id="2DAF0C40547E10D606183ADB96854576">Université de Lorraine, CNRS, lab. GeoRessources, UMR 7359, BP 70239, F- 54506 Vandoeuvre-lès-Nancy cedex (France)</mods:affiliation>
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<subSubSection id="C311653F4800317199FA640DFAFBFAFD" box="[930,1338,1299,1326]" pageId="67" pageNumber="1252" type="nomenclature">
<paragraph id="8BB436B44800317199FA640DFAFBFAFD" blockId="67.[930,1338,1299,1358]" box="[930,1338,1299,1326]" pageId="67" pageNumber="1252">
<heading id="D0FC81D84800317199FA640DFAFBFAFD" box="[930,1338,1299,1326]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="1252" reason="2">
<taxonomicName id="4C0B4D374800317199F5640AFAFBFAFD" ID-CoL="85V7Y" authority="Beauvais, 1986" authorityName="Milne Edwards &amp; Haime" authorityYear="1848" box="[941,1338,1299,1326]" class="Anthozoa" family="Stylinidae" genus="Stylosmilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="67" pageNumber="1252" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA64800317199F5640AFBE5FAFE" bold="true" box="[941,1060,1299,1326]" italics="true" pageId="67" pageNumber="1252">Stylosmilia</emphasis>
<emphasis id="B97FEAA6480031719E6E640AFB51FAFE" bold="true" box="[1078,1168,1299,1326]" italics="true" pageId="67" pageNumber="1252">dresnayi</emphasis>
<bibRefCitation id="EF9A4B45480031719ECF640AFAFBFAFD" author="BEAUVAIS L." box="[1175,1338,1299,1325]" pageId="67" pageNumber="1252" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311653F480031719E67642AFB5CFA9E" box="[1087,1181,1331,1358]" pageId="67" pageNumber="1252" type="description">
<paragraph id="8BB436B4480031719E67642AFB5CFA9E" blockId="67.[930,1338,1299,1358]" box="[1087,1181,1331,1358]" pageId="67" pageNumber="1252">
(
<figureCitation id="13302A31480031719E10642AFB55FA9E" box="[1096,1172,1331,1358]" captionStart="FIG" captionStartId="70.[132,143,1290,1307]" captionTargetBox="[132,1452,218,1246]" captionTargetId="figure-360@70.[132,1452,218,1247]" captionTargetPageId="70" captionText="FIG. 45. — “Stylosmilia” dresnayi Beauvais, 1986: A, distal view of the holotype MNHN.F.R11623; B, polished longitudinal section in one corallum of the holotype MNHN.F.R11623; C, transverse thin section in one corallum of the holotype MNHN.F.R11623; D, transverse thin section in the holotype MNHN.F.R11623; E, oblique thin section of septa in a corallum of the holotype MNHN.F.R11623. Scale bars: A, 1 mm; B, D, 2 mm; C, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5764503" httpUri="https://zenodo.org/record/5764503/files/figure.png" pageId="67" pageNumber="1252">Fig. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311653F480031719975647FFA8CFA78" pageId="67" pageNumber="1252" type="reference_group">
<paragraph id="8BB436B4480031719975647FFAE0FAAE" blockId="67.[813,1357,1382,1448]" box="[813,1313,1382,1406]" pageId="67" pageNumber="1252">
<treatmentCitationGroup id="AB1B119A480031719975647FFAE0FAAE" box="[813,1313,1382,1406]" pageId="67" pageNumber="1252">
<treatmentCitation id="0AAA10A5480031719975647FFB64FAAE" author="BEAUVAIS L." box="[813,1189,1382,1406]" page="41" pageId="67" pageNumber="1252" year="1986">
<taxonomicName id="4C0B4D37480031719975647FFB64FAAE" ID-CoL="85V7Y" authority="Beauvais, 1986: 41" authorityName="Beauvais" authorityPageNumber="41" authorityYear="1986" box="[813,1189,1382,1406]" class="Anthozoa" family="Stylinidae" genus="Stylosmilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="67" pageNumber="1252" phylum="Cnidaria" rank="species" species="dresnayi">
<emphasis id="B97FEAA6480031719975647FFC25FAAE" box="[813,996,1382,1406]" italics="true" pageId="67" pageNumber="1252">Stylosmilia dresnayi</emphasis>
<bibRefCitation id="EF9A4B454800317199B2647FFB64FAAE" author="BEAUVAIS L." box="[1002,1189,1382,1406]" pageId="67" pageNumber="1252" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais, 1986: 41</bibRefCitation>
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, pl. 8, fig. 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB436B44800317199756489FA8CFA78" blockId="67.[813,1357,1382,1448]" box="[813,1357,1424,1448]" pageId="67" pageNumber="1252">
<treatmentCitationGroup id="AB1B119A4800317199756489FA8CFA78" box="[813,1357,1424,1448]" pageId="67" pageNumber="1252">
<taxonomicName id="4C0B4D374800317199756489FC2DFA78" ID-CoL="85V7Y" authorityName="Milne Edwards &amp; Haime" authorityYear="1848" box="[813,1004,1424,1448]" class="Anthozoa" family="Stylinidae" genus="Stylosmilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="67" pageNumber="1252" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA64800317199756489FC53FA78" box="[813,914,1424,1448]" italics="true" pageId="67" pageNumber="1252">Stylosmilia</emphasis>
?
<emphasis id="B97FEAA64800317199F86489FC2DFA78" box="[928,1004,1424,1448]" italics="true" pageId="67" pageNumber="1252">dresnayi</emphasis>
</taxonomicName>
<treatmentCitation id="0AAA10A5480031719E5D6489FB2BFA78" author="VASSEUR R." box="[1029,1258,1424,1448]" page="338" pageId="67" pageNumber="1252" year="2018">
<bibRefCitation id="EF9A4B45480031719E5D6489FB2BFA78" author="VASSEUR R." box="[1029,1258,1424,1448]" pageId="67" pageNumber="1252" refId="ref55769" refString="VASSEUR R. 2018. - Extinctions et recouvrements de coraux au cours de la crise Pliensbachien-Toarcien. These de doctorat, Universite de Lorraine, France, 542 p." type="book" year="2018">Vasseur 2018: 338-339</bibRefCitation>
</treatmentCitation>
, fig. 3.93.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C311653F48003171997564A3FAC4F917" pageId="67" pageNumber="1252" type="materials_examined">
<paragraph id="8BB436B448003171997564A3FAC4F917" blockId="67.[813,1455,1465,1735]" pageId="67" pageNumber="1252">
<materialsCitation id="3B633CE948003171997564A3FAC4F917" accessionNumber="R11623" collectionCode="TYPE, MNHN" country="Morocco" location="Beauvais" municipality="Two" pageId="67" pageNumber="1252" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED1AAE7148003171997564A3FC9CFA1F" box="[813,861,1466,1489]" pageId="67" pageNumber="1252">TYPE</collectionCode>
MATERIAL. —
<typeStatus id="54B088164800317199B564A0FB8CFA01" box="[1005,1101,1465,1489]" pageId="67" pageNumber="1252" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED1AAE71480031719E0E64A3FB6AFA01" box="[1110,1195,1466,1489]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="67" pageNumber="1252">MNHN</collectionCode>
.F.R11623,
<bibRefCitation id="EF9A4B45480031719F4464A0FC89FA3C" author="BEAUVAIS L." pageId="67" pageNumber="1252" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">
<location id="8ED4606F480031719F4464A0FAB1FA01" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F480031719F4464A0FAB1FA01" box="[1308,1392,1465,1489]" country="Morocco" municipality="Two" name="Beauvais" pageId="67" pageNumber="1252">Beauvais</location>
1986: 41
</bibRefCitation>
, pl. 8, fig. 3.
<typeStatus id="54B0881648003171997564E7FC9CF9C3" box="[813,861,1534,1557]" pageId="67" pageNumber="1252">TYPE</typeStatus>
HORIZON. —
<location id="8ED4606F4800317199B464E4FBB7F9C5" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F4800317199B464E4FBB7F9C5" box="[1004,1142,1533,1557]" country="Morocco" municipality="Two" name="Pliensbachian" pageId="67" pageNumber="1252">Pliensbachian</location>
.
<typeStatus id="54B088164800317199756731FC9CF9ED" box="[813,861,1576,1599]" pageId="67" pageNumber="1252">TYPE</typeStatus>
LOCALITY. —
<location id="8ED4606F4800317199B3673EFB4EF9EF" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F4800317199B3673EFB4EF9EF" box="[1003,1167,1575,1599]" country="Morocco" municipality="Two" name="Jebel Bou Dahar" pageId="67" pageNumber="1252">Jebel Bou Dahar</location>
,
<collectingCountry id="F31C7624480031719EC06731FB32F9EF" box="[1176,1267,1576,1599]" name="Morocco" pageId="67" pageNumber="1252">Morocco</collectingCountry>
. STUDIED SAMPLES. —
<collectingMunicipality id="6BD0ACCE4800317199A56748FBE6F9B8" box="[1021,1063,1617,1640]" pageId="67" pageNumber="1252">Two</collectingMunicipality>
<specimenCount id="9D0DFD3D480031719E736748FB51F9B9" box="[1067,1168,1617,1641]" pageId="67" pageNumber="1252" type="generic">specimens</specimenCount>
:
<typeStatus id="54B08816480031719EC06748FB2CF9B9" box="[1176,1261,1617,1641]" pageId="67" pageNumber="1252" type="holotype">holotype</typeStatus>
<collectionCode id="ED1AAE71480031719EA86748FA85F9B8" box="[1264,1348,1617,1640]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="67" pageNumber="1252">MNHN</collectionCode>
.F.R11623; CPUN
<location id="8ED4606F4800317199216772FC5BF953" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F4800317199216772FC5BF953" box="[889,922,1643,1667]" country="Morocco" municipality="Two" name="Gu" pageId="67" pageNumber="1252">Gu</location>
1005E2. GEOGRAPHIC AND STRATIGRAPHIC RANGES. —
<location id="8ED4606F480031719F51678CFA52F97D" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F480031719F51678CFA52F97D" box="[1289,1427,1685,1709]" country="Morocco" municipality="Two" name="Pliensbachian" pageId="67" pageNumber="1252">Pliensbachian</location>
of
<location id="8ED4606F48003171997667B6FC10F917" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F48003171997667B6FC10F917" box="[814,977,1711,1735]" country="Morocco" municipality="Two" name="Jebel Bou Dahar" pageId="67" pageNumber="1252">Jebel Bou Dahar</location>
and
<location id="8ED4606F480031719E5967B6FB5DF917" LSID="urn:lsid:plazi:treatment:03A287A24800317699FA640DFF3BF83B:8ED4606F480031719E5967B6FB5DF917" box="[1025,1180,1711,1735]" country="Morocco" municipality="Two" name="Guigou Plateau" pageId="67" pageNumber="1252">Guigou Plateau</location>
,
<collectingCountry id="F31C7624480031719EFE67A9FAC0F917" box="[1190,1281,1712,1735]" name="Morocco" pageId="67" pageNumber="1252">Morocco</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311653F48003176997567EBFF3BF83B" lastPageId="68" lastPageNumber="1253" pageId="67" pageNumber="1252" type="description">
<paragraph id="8BB436B448003171997567EBFC7CF8DA" blockId="67.[811,1455,1778,2028]" box="[813,957,1778,1804]" pageId="67" pageNumber="1252">DESCRIPTION</paragraph>
<paragraph id="8BB436B4480031769975660BFE40F95C" blockId="67.[811,1455,1778,2028]" lastBlockId="68.[132,775,1586,1741]" lastPageId="68" lastPageNumber="1253" pageId="67" pageNumber="1252">
Phaceloid colony with cylindrical corallites multiplying by extracalicular budding. Radial elements are wavy, compact costosepta apparently free and organized in two distinct size-orders. External lateral faces of the corallites are ornamented with vertical striation made by the naked costae. Styliform columella according to
<bibRefCitation id="EF9A4B45480031719EC566A8FA91F81C" author="BEAUVAIS L." box="[1181,1360,1969,1996]" pageId="67" pageNumber="1252" pagination="1 - 68" refId="ref48913" refString="BEAUVAIS L. 1986. - Monographie des madreporaires du Jurassique inferieur du Maroc. Palaeontographica Abteilung A 194: 1 - 68." type="journal article" year="1986">Beauvais (1986)</bibRefCitation>
but that does not appear clearly in the sample. Endotheca made of rare dissepiments and tabulae that periodically divide the lumen vertically. Wall present, probably septothecal. No synapticulae observed.
</paragraph>
<caption id="DF74663C480731769ADC64A6FB6CFA36" ID-DOI="http://doi.org/10.5281/zenodo.5764497" ID-Zenodo-Dep="5764497" httpUri="https://zenodo.org/record/5764497/files/figure.png" pageId="68" pageNumber="1253" startId="68.[132,143,1471,1488]" targetBox="[133,1451,215,1427]" targetPageId="68">
<paragraph id="8BB436B4480731769ADC64A6FB6CFA36" blockId="68.[132,1455,1471,1510]" pageId="68" pageNumber="1253">
FIG. 43. —
<taxonomicName id="4C0B4D37480731769A8764A6FE96FA00" authorityName="Cuif" authorityYear="1975" box="[223,343,1471,1488]" class="Anthozoa" family="Reimaniphylliidae" genus="Distichophyllia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="68" pageNumber="1253" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6480731769A8764A6FE96FA00" box="[223,343,1471,1488]" italics="true" pageId="68" pageNumber="1253">Distichophyllia</emphasis>
</taxonomicName>
? sp.:
<emphasis id="B97FEAA6480731769BD364A6FE59FA00" bold="true" box="[395,408,1471,1488]" pageId="68" pageNumber="1253">A</emphasis>
, distal view of the specimen CPUN AM16177-5;
<emphasis id="B97FEAA648073176996A64A6FCFEFA00" bold="true" box="[818,831,1471,1488]" pageId="68" pageNumber="1253">B</emphasis>
, specimen pictured in the field (Dades Valley, Morocco);
<emphasis id="B97FEAA6480731769F4464A6FAEBFA00" bold="true" box="[1308,1322,1471,1488]" pageId="68" pageNumber="1253">C</emphasis>
, transverse thin section of the specimen CPUN AML8-1;
<emphasis id="B97FEAA6480731769B8064CCFE27FA36" bold="true" box="[472,486,1493,1510]" pageId="68" pageNumber="1253">D</emphasis>
, distal view of the specimen CPUN AM16177-9. Scale bars: A, 6 mm; B, C, D, 4 mm.
</paragraph>
</caption>
<paragraph id="8BB436B4480731769AC3678BFD89F91D" blockId="68.[132,775,1586,1741]" pageId="68" pageNumber="1253">
Calicular diameter:
<quantity id="4CF39B51480731769B29678BFDCCF97D" box="[369,525,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="68" pageNumber="1253" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5 to 2.5 mm</quantity>
Number of septa: 32 to 36 Septal density: 3 to 4 for
<quantity id="4CF39B51480731769BB267ABFD85F91D" box="[490,580,1714,1741]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="68" pageNumber="1253" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="8BB436B4480731769ADC67EBFE13F8DA" blockId="68.[131,776,1778,2027]" box="[132,466,1778,1804]" pageId="68" pageNumber="1253">SIMILARITIES AND DIFFERENCES</paragraph>
<paragraph id="8BB436B4480731769ADC660BFF3BF83B" blockId="68.[131,776,1778,2027]" pageId="68" pageNumber="1253">
Differs from the other taxa of the genus
<taxonomicName id="4C0B4D37480731769811660BFEEFF89C" authority="Milne Edwards, 1848" authorityName="Milne Edwards &amp; Haime" authorityYear="1848" class="Anthozoa" family="Stylinidae" genus="Stylosmilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="68" pageNumber="1253" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA6480731769811660BFD7CF8FC" box="[585,701,1810,1836]" italics="true" pageId="68" pageNumber="1253">Stylosmilia</emphasis>
<bibRefCitation id="EF9A4B4548073176989C660BFEEFF89C" author="MILNE EDWARDS H. &amp; HAIME J." pageId="68" pageNumber="1253" pagination="465 - 469" refId="ref53302" refString="MILNE EDWARDS H. &amp; HAIME J. 1848. - Observations sur les polypiers de la famille des astreides. Comptes rendus hebdomadaires des Seances de l'Academie des Sciences 27: 465 - 469. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 29840 / f 469. item" type="journal article" year="1848">Milne Edwards, 1848</bibRefCitation>
</taxonomicName>
by the lack of auriculae. With the doubt concerning the presence of a styliform columella we choose to maintain the species under the genus
<taxonomicName id="4C0B4D374807317698086668FD05F85B" authorityName="Milne Edwards &amp; Haime" authorityYear="1848" box="[592,708,1905,1931]" class="Anthozoa" family="Stylinidae" genus="Stylosmilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="68" pageNumber="1253" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA64807317698086668FD05F85B" box="[592,708,1905,1931]" italics="true" pageId="68" pageNumber="1253">Stylosmilia</emphasis>
</taxonomicName>
(with quotation marks) even if the genus
<taxonomicName id="4C0B4D374807317698426688FF3EF81B" authority="Roniewicz, 1976" authorityName="Roniewicz" authorityYear="1976" class="Anthozoa" family="Stylinidae" genus="Proaplophyllia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="68" pageNumber="1253" phylum="Cnidaria" rank="genus">
<emphasis id="B97FEAA64807317698426688FD75F87B" box="[538,692,1937,1963]" italics="true" pageId="68" pageNumber="1253">Proaplophyllia</emphasis>
Roniewicz, 1976
</taxonomicName>
would be more appropriate in case of lack of columella.
</paragraph>
</subSubSection>
</treatment>
</document>