441 lines
68 KiB
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441 lines
68 KiB
XML
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<mods:title id="3718DDDB3ED80BCDDDF24DC4051FA085">Contribution to the knowledge of genus Nalassus Mulsant, 1856 (Coleoptera Tenebrionidae) from the Eastern Caucasus, Russia</mods:title>
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<mods:namePart id="302F6ADBA3A97521156658361BEB4334">Nabozhenko, Maxim V.</mods:namePart>
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<mods:namePart id="FC96C09D0173E490AE05B9EA809DE70A">Gadaborsheva, Mariam A.</mods:namePart>
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<mods:affiliation id="D18C653E6838EB06D16CB720C04B14A4">Ingush State University, I. B. Zyazikova av., 7, Magas, Republic of Ingushetia 386001, Russia. & State Natural Reserve “ Erzi ”, Pobedy str., 3, Nazran, Republic of Ingushetia 386101, Russia.</mods:affiliation>
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<taxonomicName id="4C0B4D18FFFC2F5BDC90FDB3DBFCFD1D" authority="Nabozhenko" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[151,710,638,665]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. nov.">
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<emphasis id="B97FEA89FFFC2F5BDC90FDB3D83EFD1D" bold="true" box="[151,260,638,665]" italics="true" pageId="1" pageNumber="420">Nalassus</emphasis>
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(s. str.)
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<emphasis id="B97FEA89FFFC2F5BDD6FFDB3DB19FD1D" bold="true" box="[360,547,638,665]" italics="true" pageId="1" pageNumber="420">magomedrasuli</emphasis>
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Nabozhenko
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(
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<figureCitation id="13302A1EFFFC2F5BDC98FD6ED803FD39" box="[159,313,675,701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Figs 1A–C, F</figureCitation>
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,
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<figureCitation id="13302A1EFFFC2F5BDD43FD6ED8B0FD39" box="[324,394,675,701]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">2A–C</figureCitation>
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,
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<figureCitation id="13302A1EFFFC2F5BDD92FD6EDB21FD39" box="[405,539,675,701]" captionStart="FIGURE 3" captionStartId="4.[151,250,1838,1862]" captionTargetBox="[151,1432,373,1804]" captionTargetId="figure-122@4.[151,1436,368,1812]" captionTargetPageId="4" captionText="FIGURE 3. Nalassus spp., males, holotypes.A, C, D, E = N. magomedrasuli sp. n.; B, F, G, H = N. kalashiani; A, B = abdomen; C, G = protarsus; D, H = mesotarsus; E, F = middle of abdominal ventrites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.10152232" httpUri="https://zenodo.org/record/10152232/files/figure.png" pageId="1" pageNumber="420">3A, C, D, E</figureCitation>
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,
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<figureCitation id="13302A1EFFFC2F5BDE20FD6EDB57FD39" box="[551,621,675,701]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="1" pageNumber="420">4A–D</figureCitation>
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)
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<bibRefCitation id="EF9A4B6AFFFC2F5BDC90FD2AD8BDFD7B" author="Nabozhenko, M. V." box="[151,391,743,767]" pageId="1" pageNumber="420" pagination="909 - 942" refId="ref4250" refString="Nabozhenko, M. V. (2001) On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomologicheskoe obozrenie, 80 (3), 627 - 668. [in Russian; English translation: Entomological Review, 81 (8), 909 - 942]" type="journal article" year="2001">Nabozhenko 2001: 640</bibRefCitation>
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(
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<taxonomicName id="4C0B4D18FFFC2F5BDD90FD2ADB5FFD7B" authorityName="Nabozhenko" authorityYear="2001" box="[407,613,743,767]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani">
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<emphasis id="B97FEA89FFFC2F5BDD90FD2ADB5FFD7B" box="[407,613,743,767]" italics="true" pageId="1" pageNumber="420">Nalassus kalashiani</emphasis>
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, part, only
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(
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<taxonomicName id="4C0B4D18FFFC2F5BD91FFD2AD823FC9A" authorityName="Nabozhenko" authorityYear="2001" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani">
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<emphasis id="B97FEA89FFFC2F5BD91FFD2AD823FC9A" italics="true" pageId="1" pageNumber="420">Nalassus kalashiani</emphasis>
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</taxonomicName>
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, part, only
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<typeStatus id="54B08839FFFC2F5BDD89FCCBD8D0FC9A" box="[398,490,774,798]" pageId="1" pageNumber="420" type="paratype">paratype</typeStatus>
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<bibRefCitation id="EF9A4B6AFFFC2F5BDDFCFCCBDA9DFC9A" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[507,935,774,798]" pageId="1" pageNumber="420" pagination="6 - 14" refId="ref4073" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2009) Relic and endemic elements in the fauna of tenebrionid beetles (Coleoptera, Tenebrionidae) of the Big Caucasus. South of Russia: ecology, development, 4 (2), 6 - 14. [in Russian]. https: // doi. org / 10.18470 / 1992 - 1098 - 2009 - 2 - 8 - 17" type="journal article" year="2009">Abdurakhmanov & Nabozhenko 2009: 13</bibRefCitation>
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(
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<taxonomicName id="4C0B4D18FFFC2F5BDFB1FCCBDDB9FC9A" authorityName="Nabozhenko" authorityYear="2001" box="[950,1155,774,798]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani">
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<emphasis id="B97FEA89FFFC2F5BDFB1FCCBDDB9FC9A" box="[950,1155,774,798]" italics="true" pageId="1" pageNumber="420">Nalassus kalashiani</emphasis>
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</taxonomicName>
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, part, only population from Levashi,
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<collectingRegion id="49CFF879FFFC2F5BDD23FCE8D8BDFCB9" box="[292,391,805,829]" country="Russia" name="Dagestan" pageId="1" pageNumber="420">Dagestan</collectingRegion>
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);
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<bibRefCitation id="EF9A4B6AFFFC2F5BDD91FCE9DA71FCB9" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[406,843,804,829]" pageId="1" pageNumber="420" refId="ref4151" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov & Nabozhenko 2011: 292</bibRefCitation>
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(
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<taxonomicName id="4C0B4D18FFFC2F5BDF5EFCE9DD1FFCB8" authorityName="Nabozhenko" authorityYear="2001" box="[857,1061,804,828]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani">
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<emphasis id="B97FEA89FFFC2F5BDF5EFCE9DD1FFCB8" box="[857,1061,804,828]" italics="true" pageId="1" pageNumber="420">Nalassus kalashiani</emphasis>
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</taxonomicName>
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, part, only
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<typeStatus id="54B08839FFFC2F5BD890FCE8DDC9FCB9" box="[1175,1267,805,829]" pageId="1" pageNumber="420" type="paratype">paratype</typeStatus>
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);
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<bibRefCitation id="EF9A4B6AFFFC2F5BD903FCE9D861FCDF" author="Nabozhenko, M. V. & Gagarina, L. V. & Chigray, I. A." pageId="1" pageNumber="420" pagination="119 - 158" refId="ref4541" refString="Nabozhenko, M. V., Gagarina, L. V. & Chigray, I. A. (2022 a) A new Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) from Transcaucasia with a key to species from the Greater Caucasus and notes on the taxonomy, distribution, bionomics and trophic relations. Acta Zoologica Academiae Scientiarum Hungaricae, 68 (2), 119 - 158. https: // doi. org / 10.17109 / AZH. 68.2.119.2022" type="journal article" year="2022">
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Nabozhenko
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||
<emphasis id="B97FEA89FFFC2F5BD98DFCE8D9D8FCDF" italics="true" pageId="1" pageNumber="420">et al.</emphasis>
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2022a: 124
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</bibRefCitation>
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, fig. 4 (
|
||
<taxonomicName id="4C0B4D18FFFC2F5BDDACFC8EDBD0FCDF" authorityName="Nabozhenko" authorityYear="2001" box="[427,746,835,859]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani" subGenus="Nalassus">
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||
<emphasis id="B97FEA89FFFC2F5BDDACFC8EDB3DFCDF" box="[427,519,835,859]" italics="true" pageId="1" pageNumber="420">Nalassus</emphasis>
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||
(
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||
<emphasis id="B97FEA89FFFC2F5BDE11FC8EDB48FCDF" box="[534,626,835,859]" italics="true" pageId="1" pageNumber="420">Nalassus</emphasis>
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||
)
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<emphasis id="B97FEA89FFFC2F5BDE87FC8EDBD0FCDF" box="[640,746,835,859]" italics="true" pageId="1" pageNumber="420">kalashiani</emphasis>
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||
</taxonomicName>
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||
);
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||
<bibRefCitation id="EF9A4B6AFFFC2F5BDF07FC8EDD12FCDF" author="Nabozhenko, M. V. & Gagarina, L. V. & Chigray, I. A. & Nabozhenko, S. V." box="[768,1064,835,859]" pageId="1" pageNumber="420" pagination="16 - 34" refId="ref4627" refString="Nabozhenko, M. V., Gagarina, L. V., Chigray, I. A. & Nabozhenko, S. V. (2022 b) Trophic relations and ecological niches of darkling beetles of the genus Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) in the Caucasus and description of a new species from Abkhazia. South of Russia: ecology, development, 17 (3), 16 - 34. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2022 - 3 - 16 - 34" type="journal article" year="2022">
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||
Nabozhenko
|
||
<emphasis id="B97FEA89FFFC2F5BDF80FC89DA80FCDF" box="[903,954,835,859]" italics="true" pageId="1" pageNumber="420">et al.</emphasis>
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2022b: 31
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||
</bibRefCitation>
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||
, fig. 10B (
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<taxonomicName id="4C0B4D18FFFC2F5BD891FC8EDC58FCDF" authorityName="Nabozhenko" authorityYear="2001" box="[1174,1378,835,859]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFC2F5BD891FC8EDC58FCDF" box="[1174,1378,835,859]" italics="true" pageId="1" pageNumber="420">Nalassus kalashiani</emphasis>
|
||
</taxonomicName>
|
||
, part, only population from
|
||
<collectingRegion id="49CFF879FFFC2F5BDDA2FCAFDB32FCFE" box="[421,520,866,890]" country="Russia" name="Dagestan" pageId="1" pageNumber="420">Dagestan</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116510FFFC2F5BDC90FC65DD9AFB8F" pageId="1" pageNumber="420" type="materials_examined">
|
||
<paragraph id="8BB4369BFFFC2F5BDC90FC65DD9AFB8F" blockId="1.[151,1437,936,2043]" pageId="1" pageNumber="420">
|
||
<emphasis id="B97FEA89FFFC2F5BDC90FC65D87CFC46" bold="true" box="[151,326,936,962]" pageId="1" pageNumber="420">Type material.</emphasis>
|
||
<materialsCitation id="3B633CC6FFFC2F5BDD48FC65D861FB8E" collectingDate="2021-06-14" collectionCode="ZIN, PCMN, V" collectorName="Nabozhenko, I. A" country="Russia" elevation="1144" latitude="42.465336" location="Levashi" longLatPrecision="1" longitude="47.342182" pageId="1" pageNumber="420" specimenCount="3" specimenCount-female="3" stateProvince="Dagestan" typeStatus="holotype">
|
||
<typeStatus id="54B08839FFFC2F5BDD48FC65D880FC46" box="[335,442,936,962]" pageId="1" pageNumber="420" type="holotype">Holotype</typeStatus>
|
||
, 1♁ (
|
||
<collectionCode id="ED1AAE5EFFFC2F5BDDFEFC65DB11FC46" box="[505,555,936,962]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="420" type="Museum">ZIN</collectionCode>
|
||
) and
|
||
<typeStatus id="54B08839FFFC2F5BDE6BFC64DBE0FC47" box="[620,730,937,963]" pageId="1" pageNumber="420" type="paratype">paratypes</typeStatus>
|
||
, 2 ♁♁,
|
||
<specimenCount id="9D0DFD12FFFC2F5BDF31FC65DA4CFC47" box="[822,886,936,963]" count="3" pageId="1" pageNumber="420" type="female">3 ♀♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED1AAE5EFFFC2F5BDF80FC65DA80FC46" box="[903,954,936,962]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="420" type="Museum">ZIN</collectionCode>
|
||
,
|
||
<collectionCode id="ED1AAE5EFFFC2F5BDFC1FC65DD21FC46" box="[966,1051,936,962]" pageId="1" pageNumber="420">PCMN</collectionCode>
|
||
):
|
||
<collectingCountry id="F31C760BFFFC2F5BD828FC65DD47FC46" box="[1071,1149,936,962]" name="Russia" pageId="1" pageNumber="420">Russia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CFF879FFFC2F5BD88EFC64DDCEFC47" box="[1161,1268,937,963]" country="Russia" name="Dagestan" pageId="1" pageNumber="420">Dagestan</collectingRegion>
|
||
, N of
|
||
<location id="8ED46040FFFC2F5BD93AFC65DCA2FC46" LSID="urn:lsid:plazi:treatment:03A2878DFFFC2F5FDC90FDB3DD43FEE1:8ED46040FFFC2F5BD93AFC65DCA2FC46" box="[1341,1432,936,962]" country="Russia" latitude="42.465336" longLatPrecision="1" longitude="47.342182" name="Levashi" pageId="1" pageNumber="420" stateProvince="Dagestan">Levashi</location>
|
||
,
|
||
<geoCoordinate id="EE3F505CFFFC2F5BDC90FC00D87BFC6C" box="[151,321,973,1000]" degrees="42" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="420" precision="1" seconds="55.21" value="42.465336">42°27ʹ55.21ʺN</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3F505CFFFC2F5BDD4BFC00D8C9FC6C" box="[332,499,973,1000]" degrees="47" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="420" precision="1" seconds="31.86" value="47.342182">47°20ʹ31.86ʺE</geoCoordinate>
|
||
, limestone phryganoid mountain steppe,
|
||
<quantity id="4CF39B7EFFFC2F5BDFC6FC01DD2DFC63" box="[961,1047,972,999]" metricMagnitude="3" metricUnit="m" metricValue="1.144" pageId="1" pageNumber="420" unit="m" value="1144.0">
|
||
<elevation id="0026D1A8FFFC2F5BDFC6FC01DD2DFC63" box="[961,1047,972,999]" metricMagnitude="3" metricUnit="m" metricValue="1.144" pageId="1" pageNumber="420" unit="m" value="1144.0">1144 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB5105BFFFC2F5BD824FC01DD98FC62" box="[1059,1186,972,998]" pageId="1" pageNumber="420" value="2021-06-14">
|
||
<collectingDate id="EFF1E9B3FFFC2F5BD824FC01DD98FC62" box="[1059,1186,972,998]" pageId="1" pageNumber="420" value="2021-06-14">14.06.2021</collectingDate>
|
||
</date>
|
||
(leg. M.
|
||
<collectionCode id="ED1AAE5EFFFC2F5BD906FC00DC2DFC63" box="[1281,1303,973,999]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="420" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="26FE534DFFFC2F5BD926FC01D9D3FB8E" pageId="1" pageNumber="420">Nabozhenko, I.A</collectorName>
|
||
. Chigray)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B633CC6FFFC2F5BDD62FC3CDD9AFB8F" box="[357,1184,1008,1036]" collectingDate="1960-05-27" collectionCode="MSU" collectorName="D. Panfilov." country="Russia" location="Levashi" pageId="1" pageNumber="420" specimenCount="1" stateProvince="Dagestan" typeStatus="paratype">
|
||
<typeStatus id="54B08839FFFC2F5BDD62FC3CD8F0FB8F" box="[357,458,1009,1035]" pageId="1" pageNumber="420" type="paratype">Paratype</typeStatus>
|
||
, ♁ (ZM
|
||
<collectionCode id="ED1AAE5EFFFC2F5BDE2BFC3DDB54FB8E" box="[556,622,1008,1034]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="1" pageNumber="420" type="University or college">MSU</collectionCode>
|
||
):
|
||
<collectingRegion id="49CFF879FFFC2F5BDE87FC3CDBD1FB8F" box="[640,747,1009,1035]" country="Russia" name="Dagestan" pageId="1" pageNumber="420">Dagestan</collectingRegion>
|
||
,
|
||
<location id="8ED46040FFFC2F5BDEF0FC3DDA68FB8E" LSID="urn:lsid:plazi:treatment:03A2878DFFFC2F5FDC90FDB3DD43FEE1:8ED46040FFFC2F5BDEF0FC3DDA68FB8E" box="[759,850,1008,1034]" country="Russia" name="Levashi" pageId="1" pageNumber="420" stateProvince="Dagestan">Levashi</location>
|
||
,
|
||
<date id="FFB5105BFFFC2F5BDF5AFC3DDAE4FB8F" box="[861,990,1008,1035]" pageId="1" pageNumber="420" value="1960-05-27">
|
||
<collectingDate id="EFF1E9B3FFFC2F5BDF5AFC3DDAE4FB8F" box="[861,990,1008,1035]" pageId="1" pageNumber="420" value="1960-05-27">27.05.1960</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FE534DFFFC2F5BD81EFC3CDD9AFB8F" box="[1049,1184,1008,1035]" pageId="1" pageNumber="420">D. Panfilov.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116510FFFC2F5EDCC0FBD9DC2EFEE1" lastPageId="4" lastPageNumber="423" pageId="1" pageNumber="420" type="description">
|
||
<paragraph id="8BB4369BFFFC2F5BDCC0FBD9D8B7FA5A" blockId="1.[151,1437,936,2043]" pageId="1" pageNumber="420">
|
||
<emphasis id="B97FEA89FFFC2F5BDCC0FBD9D863FBAA" bold="true" box="[199,345,1044,1070]" pageId="1" pageNumber="420">Description.</emphasis>
|
||
Male. Body length
|
||
<quantity id="4CF39B7EFFFC2F5BDE3BFBD9DB8BFBAB" box="[572,689,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="1" pageNumber="420" unit="mm" value="6.9" valueMax="7.0" valueMin="6.8">6.8–7 mm</quantity>
|
||
, width
|
||
<quantity id="4CF39B7EFFFC2F5BDF02FBD9DAAAFBAB" box="[773,912,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="1" pageNumber="420" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.6–2.7 mm</quantity>
|
||
. Body elongate, robust, black-brown or black, dull (
|
||
<figureCitation id="13302A1EFFFC2F5BDCD4FBF5D820FBD6" box="[211,282,1080,1106]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Figs 1</figureCitation>
|
||
, A, C). Head widest at eye level. Eyes small dorsally, widely spaced, weakly convex (
|
||
<figureCitation id="13302A1EFFFC2F5BD8F7FBF5DC71FBD6" box="[1264,1355,1080,1106]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">Fig. 2A</figureCitation>
|
||
). Ratio of distance at widest part of head to distance between eyes 1.36–1.37. Anterior margin of epistoma straight, lateral margin of genae angulate, straight or widely emarginated in anterior two thirds. Lateral margin of head not situated between genae and epistoma. Puncturation (punctures round) of head coarse and moderately dense (
|
||
<figureCitation id="13302A1EFFFC2F5BD933FB69DCAAFB3A" box="[1332,1424,1188,1214]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">Fig. 2A</figureCitation>
|
||
): interpuncture distance subequal to puncture diameter; puncturation little sparser on epistoma. Each puncture with short recumbent seta, better visible on fronto-epistomal depression and on temples. Head ventrally with coarse puncturation and wrinkles ahead of gula, covered with short recumbent setae (
|
||
<figureCitation id="13302A1EFFFC2F5BDFFDFADDDD6BFAAE" box="[1018,1105,1296,1322]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Fig. 1C</figureCitation>
|
||
). Apical maxillary palpomere strongly widened, axe-shaped (
|
||
<figureCitation id="13302A1EFFFC2F5BDDF0FAF9DB68FACA" box="[503,594,1332,1358]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13302A1EFFFC2F5BDE67FAF9DBBCFACA" box="[608,646,1332,1358]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">2A</figureCitation>
|
||
). Antennae moderately short, with two distal antennomeres extending beyond base of pronotum. Antennomeres thinckened, widest antennomeres 2–8 and 10 (
|
||
<figureCitation id="13302A1EFFFC2F5BD880FA95DDE7FAF6" box="[1159,1245,1368,1394]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Fig. 1F</figureCitation>
|
||
); antennomere 2 transverse. Length ratio of antennomeres 2–11: 1.2, 3.5, 3, 3, 2.9, 2.9, 2.7, 2.5, 2.5, 3.4. Width ratio of antennomeres 2–11: 1.6, 1.7, 1.9, 2, 2, 2.1, 2, 1.8, 2, 1.8. Ratio of length / width of antennomeres: 0.75, 2.05, 1.57, 1.5, 1.45, 1.38, 1.35, 1.38, 1.25, 1.88.
|
||
</paragraph>
|
||
<paragraph id="8BB4369BFFFC2F5BDCC0FA25DA26F8A6" blockId="1.[151,1437,936,2043]" pageId="1" pageNumber="420">
|
||
Prothorax (
|
||
<figureCitation id="13302A1EFFFC2F5BDD43FA25D8A5F986" box="[324,415,1512,1538]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13302A1EFFFC2F5BDDABFA25D8E8F986" box="[428,466,1512,1538]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">2A</figureCitation>
|
||
). Pronotum transverse (1.48 times as wide as long), 1.67 times as wide as head, widest slightly behind middle (
|
||
<figureCitation id="13302A1EFFFC2F5BDDA5F9C1D8C7F9A2" box="[418,509,1548,1574]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="1" pageNumber="420">Fig. 2A</figureCitation>
|
||
). Lateral margins of pronotum almost straight at basal third and moderately evenly rounded after widest part to apical margin. Anterior margin almost straight, base widely weakly rounded. Angles of pronotum obtuse, widely rounded, anterior angles not projected. All margins beaded; bead of anterior margin wide and flattened (anterior bead can be interrupted in middle). Disc of pronotum moderately convex, lateral sides narrowly flattened. Puncturation of disc (punctures round) finer and sparser than on head: interpuncture distance 1.5–3 times as long as puncture diameter. Prosternum with sparse punctures in anterior half and denser smooth wrinkles puncturation in basal half. Prothoracic hypomera with longitudinal fine wrinkles and narrowly flattened outer margins. Prosternal process weakly convex at apex (
|
||
<figureCitation id="13302A1EFFFC2F5BDEB1F8C5DA34F8A6" box="[694,782,1800,1826]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4369BFFFC2F5BDCC0F8E1DAEBF87E" blockId="1.[151,1437,936,2043]" pageId="1" pageNumber="420">
|
||
Pterothorax (
|
||
<figureCitation id="13302A1EFFFC2F5BDD5EF8E1D8EDF8C2" box="[345,471,1836,1862]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="1" pageNumber="420">Figs 1A, C</figureCitation>
|
||
). Scutellar schield triangle, with slightly rounded margins and several very sparse fine punctures. Elytra elongate (1.55 times as long as wide), widest at middle, 1.9 times as wide as head, 1.14–1.15 times as wide and 2.6 times as long as pronotum. Strial punctures longitudinally elongate, connected by fine furrow. Interstriae flat, very sparsely and finely punctured (2–3 punctures in cross section). Mesoventrite pubescent by recumbent hairs, with coarse transverse wrinkles in anterior part and dense puncturation on mesocoxal process. Mesoventrite and mesepisterna with sparse and fine smooth puncturation.
|
||
</paragraph>
|
||
<caption id="DF746613FFFF2F58DC90F8CEDA5AF8E7" ID-DOI="http://doi.org/10.5281/zenodo.10152226" ID-Zenodo-Dep="10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="2" pageNumber="421" startId="2.[151,250,1795,1819]" targetBox="[155,1430,185,1765]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BB4369BFFFF2F58DC90F8CEDA5AF8E7" blockId="2.[151,1437,1795,1891]" pageId="2" pageNumber="421">
|
||
<emphasis id="B97FEA89FFFF2F58DC90F8CED829F89F" bold="true" box="[151,275,1795,1819]" pageId="2" pageNumber="421">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C0B4D18FFFF2F58DD1FF8CED89CF89E" box="[280,422,1795,1819]" class="Insecta" family="Tenebrionidae" genus="Nalassus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97FEA89FFFF2F58DD1FF8CED84EF89F" box="[280,372,1795,1819]" italics="true" pageId="2" pageNumber="421">Nalassus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, habitus and details of structure. A =
|
||
<taxonomicName id="4C0B4D18FFFF2F58DF1EF8C9DAEEF89F" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[793,980,1795,1819]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
|
||
<emphasis id="B97FEA89FFFF2F58DF1EF8C9DAEEF89F" box="[793,980,1795,1819]" italics="true" pageId="2" pageNumber="421">N. magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFFF2F58DFDEF8CEDD2BF89F" bold="true" box="[985,1041,1795,1819]" pageId="2" pageNumber="421">
|
||
<taxonomicNameLabel id="A24C57F2FFFF2F58DFDEF8CEDD2BF89F" box="[985,1041,1795,1819]" pageId="2" pageNumber="421" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D =
|
||
<taxonomicName id="4C0B4D18FFFF2F58DF0DF8E5DAABF8BB" authorityName="Nabozhenko" authorityYear="2001" box="[778,913,1831,1855]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFF2F58DF0DF8E5DAABF8BB" box="[778,913,1831,1855]" italics="true" pageId="2" pageNumber="421">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
, male (holotype) habitus dorsally; E = ditto, female; F =
|
||
<taxonomicName id="4C0B4D18FFFF2F58DCC7F881D846F8E7" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[192,380,1867,1891]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
|
||
<emphasis id="B97FEA89FFFF2F58DCC7F881D846F8E7" box="[192,380,1867,1891]" italics="true" pageId="2" pageNumber="421">N. magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFFF2F58DD85F886D881F8E7" bold="true" box="[386,443,1867,1891]" pageId="2" pageNumber="421">
|
||
<taxonomicNameLabel id="A24C57F2FFFF2F58DD85F886D881F8E7" box="[386,443,1867,1891]" pageId="2" pageNumber="421" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male antennae; G =
|
||
<taxonomicName id="4C0B4D18FFFF2F58DE93F881DA21F8E7" authorityName="Nabozhenko" authorityYear="2001" box="[660,795,1867,1891]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFF2F58DE93F881DA21F8E7" box="[660,795,1867,1891]" italics="true" pageId="2" pageNumber="421">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB4369BFFFF2F58DCC0F85FDDC0F84B" blockId="2.[151,1436,1937,2036]" pageId="2" pageNumber="421">
|
||
Legs comparatively thin. Trochanters with one long seta (
|
||
<figureCitation id="13302A1EFFFF2F58DF64F85CDA87F82F" box="[867,957,1937,1963]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="2" pageNumber="421">Fig. 1C</figureCitation>
|
||
). Tibiae straight. Pro- and mesotarsi very weakly widened, laterally and ventrally covered with moderately short yellowish hairs (
|
||
<figureCitation id="13302A1EFFFF2F58D869F878DDD4F84B" box="[1134,1262,1973,1999]" captionStart="FIGURE 3" captionStartId="4.[151,250,1838,1862]" captionTargetBox="[151,1432,373,1804]" captionTargetId="figure-122@4.[151,1436,368,1812]" captionTargetPageId="4" captionText="FIGURE 3. Nalassus spp., males, holotypes.A, C, D, E = N. magomedrasuli sp. n.; B, F, G, H = N. kalashiani; A, B = abdomen; C, G = protarsus; D, H = mesotarsus; E, F = middle of abdominal ventrites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.10152232" httpUri="https://zenodo.org/record/10152232/files/figure.png" pageId="2" pageNumber="421">Figs 3C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4369BFFFF2F59DCC0F814DD46FE0D" blockId="2.[151,1436,1937,2036]" lastBlockId="3.[151,1437,151,393]" lastPageId="3" lastPageNumber="422" pageId="2" pageNumber="421">
|
||
Abdomen (
|
||
<figureCitation id="13302A1EFFFF2F58DD41F814D8F2F877" box="[326,456,2009,2035]" captionStart="FIGURE 3" captionStartId="4.[151,250,1838,1862]" captionTargetBox="[151,1432,373,1804]" captionTargetId="figure-122@4.[151,1436,368,1812]" captionTargetPageId="4" captionText="FIGURE 3. Nalassus spp., males, holotypes.A, C, D, E = N. magomedrasuli sp. n.; B, F, G, H = N. kalashiani; A, B = abdomen; C, G = protarsus; D, H = mesotarsus; E, F = middle of abdominal ventrites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.10152232" httpUri="https://zenodo.org/record/10152232/files/figure.png" pageId="2" pageNumber="421">Figs 3A, E</figureCitation>
|
||
). Abdominal ventrites 1–4 finely and sparsely punctured; abdominal ventrite 5 denser punctured, completely beaded at apex. First abdominal ventrite with much coarser puncturation than on other surface and hair brush; metacoxal process of abdominal ventrite 1 has much sparser puncturation in apical part (
|
||
<figureCitation id="13302A1EFFFE2F59D968FF76D987FF7D" captionStart="FIGURE 3" captionStartId="4.[151,250,1838,1862]" captionTargetBox="[151,1432,373,1804]" captionTargetId="figure-122@4.[151,1436,368,1812]" captionTargetPageId="4" captionText="FIGURE 3. Nalassus spp., males, holotypes.A, C, D, E = N. magomedrasuli sp. n.; B, F, G, H = N. kalashiani; A, B = abdomen; C, G = protarsus; D, H = mesotarsus; E, F = middle of abdominal ventrites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.10152232" httpUri="https://zenodo.org/record/10152232/files/figure.png" pageId="3" pageNumber="422">Fig. 3A</figureCitation>
|
||
). Male genitalia (
|
||
<figureCitation id="13302A1EFFFE2F59DD85FF12DB3EFF7D" box="[386,516,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="3" pageNumber="422">Figs 4A–D</figureCitation>
|
||
). Apical piece of the aedeagus with slightly rounded lateral margins and narrowly rounded apex; median lobe of aedeagus with two straight wide alae (
|
||
<figureCitation id="13302A1EFFFE2F59DFBCFECEDD22FE99" box="[955,1048,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="3" pageNumber="422">Fig. 4A</figureCitation>
|
||
). Inner sternite 6 (true VIII) with rounded apices and deep rounded emargination in middle, lobes densely pubescent (
|
||
<figureCitation id="13302A1EFFFE2F59D858FEEADD80FEC5" box="[1119,1210,295,321]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="3" pageNumber="422">Fig. 4B</figureCitation>
|
||
). Spiculum gastrale with thin branches connected in long common stem and oval blades (
|
||
<figureCitation id="13302A1EFFFE2F59DF9BFE86DACCFEE1" box="[924,1014,331,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="3" pageNumber="422">Fig. 4D</figureCitation>
|
||
). IX tergite very small, evenly sclerotized (without median light line), with sparse punctures and long hairs apically (
|
||
<figureCitation id="13302A1EFFFE2F59D810FEA2DD4AFE0D" box="[1047,1136,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="3" pageNumber="422">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF746613FFFE2F59DC90F980D8E3F92A" ID-DOI="http://doi.org/10.5281/zenodo.10152228" ID-Zenodo-Dep="10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="3" pageNumber="422" startId="3.[151,250,1613,1637]" targetBox="[151,1436,404,1587]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BB4369BFFFE2F59DC90F980D8E3F92A" blockId="3.[151,1437,1613,1710]" pageId="3" pageNumber="422">
|
||
<emphasis id="B97FEA89FFFE2F59DC90F980D82EF9E1" bold="true" box="[151,276,1613,1637]" pageId="3" pageNumber="422">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C0B4D18FFFE2F59DD1BF983D891F9E1" box="[284,427,1614,1638]" class="Insecta" family="Tenebrionidae" genus="Nalassus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97FEA89FFFE2F59DD1BF983D842F9E2" box="[284,376,1614,1638]" italics="true" pageId="3" pageNumber="422">Nalassus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, details of structure. A =
|
||
<taxonomicName id="4C0B4D18FFFE2F59DEA8F983DA56F9E2" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[687,876,1614,1638]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
|
||
<emphasis id="B97FEA89FFFE2F59DEA8F983DA56F9E2" box="[687,876,1614,1638]" italics="true" pageId="3" pageNumber="422">N. magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFFE2F59DF74F983DA97F9E2" bold="true" box="[883,941,1614,1638]" pageId="3" pageNumber="422">
|
||
<taxonomicNameLabel id="A24C57F2FFFE2F59DF74F983DA97F9E2" box="[883,941,1614,1638]" pageId="3" pageNumber="422" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D =
|
||
<taxonomicName id="4C0B4D18FFFE2F59DED0F9BFDA65F90E" authorityName="Nabozhenko" authorityYear="2001" box="[727,863,1650,1674]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFE2F59DED0F9BFDA65F90E" box="[727,863,1650,1674]" italics="true" pageId="3" pageNumber="422">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB4369BFFFE2F59DCC0F911DAE7F8BA" blockId="3.[151,1437,1756,1998]" pageId="3" pageNumber="422">
|
||
Female (
|
||
<figureCitation id="13302A1EFFFE2F59DD2EF911D8BFF972" box="[297,389,1756,1782]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="3" pageNumber="422">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="13302A1EFFFE2F59DD94F911D88DF972" box="[403,439,1756,1782]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="3" pageNumber="422">2B</figureCitation>
|
||
). Body more robust, with shorter, not thickened antennae (three distal antennomeres are widest). Apical maxillary palpomere less widened than in male. Lateral margins of elytra more rounded. Pro- and mesotarsi narrower, not widened. Abdominal ventrite 1 without hair brush.
|
||
</paragraph>
|
||
<paragraph id="8BB4369BFFFE2F59DCC0F88AD844F84A" blockId="3.[151,1437,1756,1998]" pageId="3" pageNumber="422">
|
||
<emphasis id="B97FEA89FFFE2F59DCC0F88AD8DEF8E5" bold="true" box="[199,484,1863,1889]" pageId="3" pageNumber="422">Comparative diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="4C0B4D18FFFE2F59DF1DF884DA89F8E6" authorityName="Nabozhenko" authorityYear="2001" box="[794,947,1864,1890]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFE2F59DF1DF884DA89F8E6" box="[794,947,1864,1890]" italics="true" pageId="3" pageNumber="422">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
. Differences are given in the table 1. The similar species
|
||
<taxonomicName id="4C0B4D18FFFE2F59DD45F8A0D8D1F802" baseAuthorityName="Faldermann" baseAuthorityYear="1837" box="[322,491,1900,1926]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="faldermanni">
|
||
<emphasis id="B97FEA89FFFE2F59DD45F8A0D8D1F802" box="[322,491,1900,1926]" italics="true" pageId="3" pageNumber="422">N. faldermanni</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0B4D18FFFE2F59DE7BF8A0DA70F802" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[636,842,1900,1926]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
|
||
<emphasis id="B97FEA89FFFE2F59DE7BF8A0DA70F802" box="[636,842,1900,1926]" italics="true" pageId="3" pageNumber="422">N. magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFFE2F59DF56F8A1DAB5F802" bold="true" box="[849,911,1900,1926]" pageId="3" pageNumber="422">
|
||
<taxonomicNameLabel id="A24C57F2FFFE2F59DF56F8A1DAB5F802" box="[849,911,1900,1926]" pageId="3" pageNumber="422" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C0B4D18FFFE2F59DFC3F8A0DD62F802" authorityName="Nabozhenko" authorityYear="2001" box="[964,1112,1900,1926]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFFE2F59DFC3F8A0DD62F802" box="[964,1112,1900,1926]" italics="true" pageId="3" pageNumber="422">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
in the large convex narrower spaced eyes (Nabozhenko
|
||
<emphasis id="B97FEA89FFFE2F59DDB8F85CD8C2F82E" box="[447,504,1936,1962]" italics="true" pageId="3" pageNumber="422">at al.</emphasis>
|
||
2022) and very long acute apical piece of the aedeagus with straight lateral margins (
|
||
<bibRefCitation id="EF9A4B6AFFFE2F59DC98F879D84BF84A" author="Nabozhenko, M. V." box="[159,369,1972,1998]" pageId="3" pageNumber="422" pagination="909 - 942" refId="ref4250" refString="Nabozhenko, M. V. (2001) On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomologicheskoe obozrenie, 80 (3), 627 - 668. [in Russian; English translation: Entomological Review, 81 (8), 909 - 942]" type="journal article" year="2001">Nabozhenko 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB4369BFFF92F5EDCC0FF5ADC2EFEE1" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="423">
|
||
<emphasis id="B97FEA89FFF92F5EDCC0FF5AD82BFF35" bold="true" box="[199,273,151,177]" pageId="4" pageNumber="423">Notes.</emphasis>
|
||
<taxonomicName id="4C0B4D18FFF92F5EDD1FFF5ADB08FF35" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[280,562,151,177]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. nov.">
|
||
<emphasis id="B97FEA89FFF92F5EDD1FFF5ADB08FF35" box="[280,562,151,177]" italics="true" pageId="4" pageNumber="423">Nalassus magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFF92F5EDE3EFF5ADB42FF35" bold="true" box="[569,632,151,177]" pageId="4" pageNumber="423">
|
||
<taxonomicNameLabel id="A24C57F2FFF92F5EDE3EFF5ADB42FF35" box="[569,632,151,177]" pageId="4" pageNumber="423" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was originally described as
|
||
<collectingRegion id="49CFF879FFF92F5EDFB2FF5ADD27FF35" box="[949,1053,151,177]" country="Russia" name="Dagestan" pageId="4" pageNumber="423">Dagestan</collectingRegion>
|
||
population of
|
||
<taxonomicName id="4C0B4D18FFF92F5ED8C6FF5ADC6CFF35" authorityName="Nabozhenko" authorityYear="2001" box="[1217,1366,151,177]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFF92F5ED8C6FF5ADC6CFF35" box="[1217,1366,151,177]" italics="true" pageId="4" pageNumber="423">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
based on the structure of the aedeagus, which is similar in both species (compare the aedeagus on
|
||
<figureCitation id="13302A1EFFF92F5ED893FF76DDDBFF51" box="[1172,1249,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1371,1395]" captionTargetBox="[162,1436,368,1345]" captionTargetId="figure-99@5.[151,1436,368,1345]" captionTargetPageId="5" captionText="FIGURE 4. Nalassus spp., male genitalia and habitats. A–E = N. magomedrasuli sp. n.; F = N. kalashiani; A = aedeagus, B = inner sternite VIII; C = inner tergite IX; D = spiculum gastrale; E = limestone mountain phryganoid steppe, 1100 m (Levashi, Dagestan, Russia); F = alpine meadow, 2200 m (southern slope of Tsoreylam ridge, State Natural Reserve “Erzi”, Ingushetia, Russia)." figureDoi="http://doi.org/10.5281/zenodo.10152234" httpUri="https://zenodo.org/record/10152234/files/figure.png" pageId="4" pageNumber="423">Fig. 4a</figureCitation>
|
||
here and on Fig.
|
||
<quantity id="4CF39B7EFFF92F5EDC90FF12D9E8FF7D" box="[151,210,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="423" unit="in" value="18.0">18 in</quantity>
|
||
<bibRefCitation id="EF9A4B6AFFF92F5EDCDDFF12D8FDFF7D" author="Nabozhenko, M. V." box="[218,455,223,249]" pageId="4" pageNumber="423" pagination="909 - 942" refId="ref4250" refString="Nabozhenko, M. V. (2001) On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomologicheskoe obozrenie, 80 (3), 627 - 668. [in Russian; English translation: Entomological Review, 81 (8), 909 - 942]" type="journal article" year="2001">Nabozhenko (2001))</bibRefCitation>
|
||
. We studied an additional material from
|
||
<collectingRegion id="49CFF879FFF92F5EDF9FFF12DD3AFF7D" box="[920,1024,223,249]" country="Russia" name="Dagestan" pageId="4" pageNumber="423">Dagestan</collectingRegion>
|
||
and
|
||
<collectingRegion id="49CFF879FFF92F5ED83DFF12DD88FF7D" box="[1082,1202,223,249]" country="Russia" name="Ingushetiya" pageId="4" pageNumber="423">Ingushetia</collectingRegion>
|
||
, again compared the
|
||
<typeStatus id="54B08839FFF92F5EDC90FECED9C3FE99" box="[151,249,259,285]" pageId="4" pageNumber="423" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0B4D18FFF92F5EDD19FECED88EFE99" authorityName="Nabozhenko" authorityYear="2001" box="[286,436,259,285]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFF92F5EDD19FECED88EFE99" box="[286,436,259,285]" italics="true" pageId="4" pageNumber="423">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
with males from
|
||
<collectingRegion id="49CFF879FFF92F5EDE7FFECEDBDAFE99" box="[632,736,259,285]" country="Russia" name="Dagestan" pageId="4" pageNumber="423">Dagestan</collectingRegion>
|
||
and collected and compared females of both populations. As a result, we describe the new species based on several diagnostic characters, mountain isolation and different habitats. The similar structure of the aedeagus can be explained by allopatric distribution of these close species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF746613FFF92F5EDC90F8E3DADEF8EE" ID-DOI="http://doi.org/10.5281/zenodo.10152232" ID-Zenodo-Dep="10152232" httpUri="https://zenodo.org/record/10152232/files/figure.png" pageId="4" pageNumber="423" startId="4.[151,250,1838,1862]" targetBox="[151,1432,373,1804]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BB4369BFFF92F5EDC90F8E3DADEF8EE" blockId="4.[151,1436,1838,1898]" pageId="4" pageNumber="423">
|
||
<emphasis id="B97FEA89FFF92F5EDC90F8E3D836F8C2" bold="true" box="[151,268,1838,1862]" pageId="4" pageNumber="423">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C0B4D18FFF92F5EDD10F8E3D89EF8C1" box="[279,420,1838,1862]" class="Insecta" family="Tenebrionidae" genus="Nalassus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97FEA89FFF92F5EDD10F8E3D849F8C2" box="[279,371,1838,1862]" italics="true" pageId="4" pageNumber="423">Nalassus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, males, holotypes. A, C, D, E =
|
||
<taxonomicName id="4C0B4D18FFF92F5EDEE7F8E2DAA1F8C2" authority="Nabozhenko & Gadaborsheva, 2023" authorityName="Nabozhenko & Gadaborsheva" authorityYear="2023" box="[736,923,1838,1862]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
|
||
<emphasis id="B97FEA89FFF92F5EDEE7F8E2DAA1F8C2" box="[736,923,1838,1862]" italics="true" pageId="4" pageNumber="423">N. magomedrasuli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97FEA89FFF92F5EDF98F8E3DAEDF8C2" bold="true" box="[927,983,1838,1862]" pageId="4" pageNumber="423">
|
||
<taxonomicNameLabel id="A24C57F2FFF92F5EDF98F8E3DAEDF8C2" box="[927,983,1838,1862]" pageId="4" pageNumber="423" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; B, F, G, H =
|
||
<taxonomicName id="4C0B4D18FFF92F5ED867F8E2DDDDF8C2" authorityName="Nabozhenko" authorityYear="2001" box="[1120,1255,1838,1862]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="kalashiani">
|
||
<emphasis id="B97FEA89FFF92F5ED867F8E2DDDDF8C2" box="[1120,1255,1838,1862]" italics="true" pageId="4" pageNumber="423">N. kalashiani</emphasis>
|
||
</taxonomicName>
|
||
; A, B = abdomen; C, G = protarsus; D, H = mesotarsus; E, F = middle of abdominal ventrites 1 and 2.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3116510FFF92F5FDCC0F855D8C9FF51" lastPageId="5" lastPageNumber="424" pageId="4" pageNumber="423" type="etymology">
|
||
<paragraph id="8BB4369BFFF92F5FDCC0F855D8C9FF51" blockId="4.[151,1436,1944,2043]" lastBlockId="5.[151,1436,151,357]" lastPageId="5" lastPageNumber="424" pageId="4" pageNumber="423">
|
||
<emphasis id="B97FEA89FFF92F5EDCC0F855D874F836" bold="true" box="[199,334,1944,1970]" pageId="4" pageNumber="423">Etymology.</emphasis>
|
||
The new species in named in honour of Corresponding Member of the Russian Academy of Sciences Magomedrasul Dibirovich Magomedov (Precaspian Institute of Biological Resources of the
|
||
<collectingRegion id="49CFF879FFF92F5ED8ECF871DC5BF852" box="[1259,1377,1980,2006]" country="Russia" name="Dagestan" pageId="4" pageNumber="423">Daghestan</collectingRegion>
|
||
Federal Research Centre of the Russian Academy of Sciences,
|
||
<collectingRegion id="49CFF879FFF92F5EDF2CF82CDAACF87F" box="[811,918,2017,2043]" country="Russia" name="Dagestan" pageId="4" pageNumber="423">Dagestan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31C760BFFF92F5EDFA6F82DDACAF87E" box="[929,1008,2016,2042]" name="Russia" pageId="4" pageNumber="423">Russia</collectingCountry>
|
||
), who made a great contribution to the study of animals of the Caucasus in general and
|
||
<collectingRegion id="49CFF879FFF82F5FDEB0FF5ADA25FF35" box="[695,799,151,177]" country="Russia" name="Dagestan" pageId="5" pageNumber="424">Dagestan</collectingRegion>
|
||
in particular. He also organized the expedition, in which this new species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3116510FFF82F5FDCC0FF13DD43FEE1" pageId="5" pageNumber="424" type="distribution">
|
||
<paragraph id="8BB4369BFFF82F5FDCC0FF13DD43FEE1" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="424">
|
||
<emphasis id="B97FEA89FFF82F5FDCC0FF13D8D7FF7C" bold="true" box="[199,493,222,249]" pageId="5" pageNumber="424">Distribution and habitat.</emphasis>
|
||
<collectingCountry id="F31C760BFFF82F5FDDF5FF12DB7AFF7D" box="[498,576,223,249]" name="Russia" pageId="5" pageNumber="424">Russia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CFF879FFF82F5FDE4FFF12DB8EFF7D" box="[584,692,223,249]" country="Russia" name="Dagestan" pageId="5" pageNumber="424">Dagestan</collectingRegion>
|
||
. We collected this species in mountain steppe near Levashi (Levashi District) and one dry specimen (only elytra and abdomen) probably belonging to the new species in the similar habitat (limestone phryganoid steppe) near Verkhniy Gunib (Gunib District,
|
||
<collectingRegion id="49CFF879FFF82F5FDFF0FEEADD5EFEC5" box="[1015,1124,295,321]" country="Russia" name="Dagestan" pageId="5" pageNumber="424">Dagestan</collectingRegion>
|
||
). Adults feed on terricolous foliose lichen
|
||
<taxonomicName id="4C0B4D18FFF82F5FDD31FE86DB3FFEE1" box="[310,517,331,357]" class="Lecanoromycetes" family="Cladoniaceae" genus="Cladonia" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="424" phylum="Ascomycota" rank="species" species="pyxidata">
|
||
<emphasis id="B97FEA89FFF82F5FDD31FE86DB3FFEE1" box="[310,517,331,357]" italics="true" pageId="5" pageNumber="424">Cladonia pyxidata</emphasis>
|
||
</taxonomicName>
|
||
(L.) Hoffm. (
|
||
<taxonomicName id="4C0B4D18FFF82F5FDE99FE86DA02FEE1" authorityName="Zenker" authorityYear="1827" box="[670,824,331,357]" class="Lecanoromycetes" family="Cladoniaceae" kingdom="Fungi" order="Lecanorales" pageId="5" pageNumber="424" phylum="Ascomycota" rank="family">Cladoniaceae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EF9A4B6AFFF82F5FDF49FE86DD57FEE1" author="Nabozhenko, M. V. & Gagarina, L. V. & Chigray, I. A." box="[846,1133,331,357]" pageId="5" pageNumber="424" pagination="119 - 158" refId="ref4541" refString="Nabozhenko, M. V., Gagarina, L. V. & Chigray, I. A. (2022 a) A new Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) from Transcaucasia with a key to species from the Greater Caucasus and notes on the taxonomy, distribution, bionomics and trophic relations. Acta Zoologica Academiae Scientiarum Hungaricae, 68 (2), 119 - 158. https: // doi. org / 10.17109 / AZH. 68.2.119.2022" type="journal article" year="2022">
|
||
Nabozhenko
|
||
<emphasis id="B97FEA89FFF82F5FDFE2FE86DD27FEE1" box="[997,1053,331,357]" italics="true" pageId="5" pageNumber="424">et al.</emphasis>
|
||
2022a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |