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<mods:title id="5438A6E0C142B1E16DEF8B98E9DBCBC2">Contribution to the knowledge of genus Nalassus Mulsant, 1856 (Coleoptera Tenebrionidae) from the Eastern Caucasus, Russia</mods:title>
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<mods:namePart id="1EBECF3317DBB0AC6DA33BA31DF684D1">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="187D1BDE47310C4F1A22F8EDD50A3344">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia. &amp; State Natural Reserve “ Erzi ”, Pobedy str., 3, Nazran, Republic of Ingushetia 386101, Russia.</mods:affiliation>
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<mods:affiliation id="44B1A7E4063B735D17FD44FB6943C938">Ingush State University, I. B. Zyazikova av., 7, Magas, Republic of Ingushetia 386001, Russia. &amp; State Natural Reserve “ Erzi ”, Pobedy str., 3, Nazran, Republic of Ingushetia 386101, Russia.</mods:affiliation>
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<emphasis id="B97FEA89FFF82F5FDC90F9FDD9C5F9CF" bold="true" box="[151,255,1584,1611]" italics="true" pageId="5" pageNumber="424">Nalassus</emphasis>
(s. str.) 
<emphasis id="B97FEA89FFF82F5FDD59F9FDD8EDF9CF" bold="true" box="[350,471,1584,1611]" italics="true" pageId="5" pageNumber="424">kalashiani</emphasis>
<bibRefCitation id="EF9A4B6AFFF82F5FDDD9F9FCDB80F9CF" author="Nabozhenko, M. V." box="[478,698,1585,1611]" pageId="5" pageNumber="424" pagination="909 - 942" refId="ref4250" refString="Nabozhenko, M. V. (2001) On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomologicheskoe obozrenie, 80 (3), 627 - 668. [in Russian; English translation: Entomological Review, 81 (8), 909 - 942]" type="journal article" year="2001">Nabozhenko, 2001</bibRefCitation>
</emphasis>
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(
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, D, E, G, 
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, 
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)
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, 658 (key), figs 17–19 (part, only 
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); 
<bibRefCitation id="EF9A4B6AFFF82F5FDF71F954DC1CF936" author="Nabozhenko, M. V. &amp; Abdurakhmanov, G. M." box="[886,1318,1689,1714]" pageId="5" pageNumber="424" pagination="187 - 191" refId="ref4474" refString="Nabozhenko, M. V. &amp; Abdurakhmanov, G. M. (2007) Review of the genus Nalassus Mulsant, 1854 (Coleoptera, Tenebrionidae) in Daghestan. Caucasian Entomological Bulletin, 3 (2), 187 - 191. https: // doi. org / 10.23885 / 1814 - 3326 - 2007 - 3 - 2 - 187 - 191" type="journal article" year="2007">Nabozhenko, Abdurakhmanov, 2007: 188</bibRefCitation>
(part, only 
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); 
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(part, only population from Itum-Kale, Chechen Republic); 
<bibRefCitation id="EF9A4B6AFFF82F5FD94FF975DB1FF96B" author="Abdurakhmanov, G. M. &amp; Nabozhenko, M. V." pageId="5" pageNumber="424" refId="ref4151" refString="Abdurakhmanov, G. M. &amp; Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov, Nabozhenko, 2011: 292</bibRefCitation>
, 139 (key), fig. 393 (part, only 
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); 
<bibRefCitation id="EF9A4B6AFFF82F5FDFD4F91ADC3BF96B" author="Nabozhenko, M. V. &amp; Gagarina, L. V. &amp; Chigray, I. A. &amp; Nabozhenko, S. V." box="[979,1281,1751,1775]" pageId="5" pageNumber="424" pagination="16 - 34" refId="ref4627" refString="Nabozhenko, M. V., Gagarina, L. V., Chigray, I. A. &amp; Nabozhenko, S. V. (2022 b) Trophic relations and ecological niches of darkling beetles of the genus Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) in the Caucasus and description of a new species from Abkhazia. South of Russia: ecology, development, 17 (3), 16 - 34. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2022 - 3 - 16 - 34" type="journal article" year="2022">
Nabozhenko 
<emphasis id="B97FEA89FFF82F5FD85DF915DDB7F96B" box="[1114,1165,1751,1775]" italics="true" pageId="5" pageNumber="424">et al.</emphasis>
, 2022b: 31
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, fig. 10B (part, only population from Chechen Republic).
</paragraph>
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<emphasis id="B97FEA89FFF82F5FDC90F8F1D87DF8D1" bold="true" box="[151,327,1851,1878]" pageId="5" pageNumber="424">Type material.</emphasis>
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<typeStatus id="54B08839FFF82F5FDD57F8F1D881F8D2" box="[336,443,1852,1878]" pageId="5" pageNumber="424" type="holotype">Holotype</typeStatus>
, ♁ (
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): “ 
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, 
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</date>
leg. 
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” (Cyrillic label), “
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<typeStatus id="54B08839FFF82F5FDCA6F8ADD822F8FE" box="[161,280,1888,1914]" pageId="5" pageNumber="424" type="holotype">Holotypus</typeStatus>
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<emphasis id="B97FEA89FFF82F5FDD18F8ADDB3AF8FE" box="[287,512,1888,1914]" italics="true" pageId="5" pageNumber="424">Nalassus kalashiani</emphasis>
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det. M. 
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. Nabozhenko” (red, handwritten)
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.
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<emphasis id="B97FEA89FFF82F5FDCC0F849D84AF819" bold="true" box="[199,368,1923,1950]" pageId="5" pageNumber="424">New material.</emphasis>
<materialsCitation id="3B633CC6FFF82F5FDD7FF849DCB9F846" collectingDate="2023-12-06" collectionCode="ZIN, V" collectorName="Nabozhenko, M. A" country="Russia" county="Dzheyrakh Distr." elevation="2200" latitude="42.831944" location="State" longLatPrecision="19" longitude="45.061943" municipality="Tsoreylam" pageId="5" pageNumber="424" specimenCount="1" specimenCount-female="1" stateProvince="Ingushetiya">
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(
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), 
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, 
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, 
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, 
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ridge, 
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natural reserve «Erzi» 
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, 
<geoCoordinate id="EE3F505CFFF82F5FDD2EF864D896F840" box="[297,428,1961,1988]" degrees="45" direction="east" minutes="03" orientation="longitude" pageId="5" pageNumber="424" precision="15" seconds="43" value="45.061943">45°03ʹ43ʺE</geoCoordinate>
, alpine zone, 
<quantity id="4CF39B7EFFF82F5FDE41F865DBA7F846" box="[582,669,1960,1986]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="424" unit="m" value="2200.0">
<elevation id="0026D1A8FFF82F5FDE41F865DBA7F846" box="[582,669,1960,1986]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="424" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, 
<date id="FFB5105BFFF82F5FDEAEF865DA1DF846" box="[681,807,1960,1986]" pageId="5" pageNumber="424" value="2023-12-06">
<collectingDate id="EFF1E9B3FFF82F5FDEAEF865DA1DF846" box="[681,807,1960,1986]" pageId="5" pageNumber="424" value="2023-12-06">12.06.2023</collectingDate>
</date>
(leg. M. 
<collectionCode id="ED1AAE5EFFF82F5FDF83F865DAA2F846" box="[900,920,1960,1986]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="424" type="Museum">V</collectionCode>
. &amp; S. 
<collectionCode id="ED1AAE5EFFF82F5FDFD1F865DAD7F846" box="[982,1005,1960,1986]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="424" type="Museum">V</collectionCode>
. 
<collectorName id="26FE534DFFF82F5FDFFFF865DDF0F846" box="[1016,1226,1960,1986]" pageId="5" pageNumber="424">Nabozhenko, M.A</collectorName>
. Gadaborsheva)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCC0FF5BD88FFE0C" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">
<emphasis id="B97FEA89FFFB2F5CDCC0FF5BDB36FF34" bold="true" box="[199,524,150,176]" pageId="6" pageNumber="425">Brief description of female.</emphasis>
Body length 
<quantity id="4CF39B7EFFFB2F5CDEA3FF5BDBC6FF35" box="[676,764,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="425" unit="mm" value="7.5">7.5 mm</quantity>
, width 
<quantity id="4CF39B7EFFFB2F5CDF49FF5BDA9DFF35" box="[846,935,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="425" unit="mm" value="3.4">3.4 mm</quantity>
. Body robust, convex, brown (elytra without weak reddish tinge), dull (
<figureCitation id="13302A1EFFFB2F5CDDC3FF77DB26FF50" box="[452,540,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="6" pageNumber="425">Fig. 1E</figureCitation>
). Head widest at eye level (
<figureCitation id="13302A1EFFFB2F5CDF59FF77DA8DFF50" box="[862,951,186,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="6" pageNumber="425">Fig. 2E</figureCitation>
). Eyes small dorsally, widely spaced, convex (
<figureCitation id="13302A1EFFFB2F5CDCC8FF13D81DFF7C" box="[207,295,222,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="6" pageNumber="425">Fig. 2E</figureCitation>
). Ratio of distance at widest part of head to distance between eyes 1.43 (the same in male, 
<typeStatus id="54B08839FFFB2F5CD92DFF13DCABFF7C" box="[1322,1425,222,248]" pageId="6" pageNumber="425" type="holotype">holotype</typeStatus>
). Puncturation of head dorsally moderately coarse and dense: interpuncture distance subequal to puncture diameter. Apical maxillary palpomere strongly widened, axe shaped, has the same width as in male (compare 
<figureCitation id="13302A1EFFFB2F5CD90DFEEBD982FEE0" captionStart="FIGURE 1" captionStartId="2.[151,250,1795,1819]" captionTargetBox="[155,1430,185,1765]" captionTargetId="figure-19@2.[151,1436,179,1769]" captionTargetPageId="2" captionText="FIGURE 1. Nalassus spp., habitus and details of structure.A = N. magomedrasuli sp. n., male (holotype), habitus dorsally; B = ditto (paratype), female; C = ditto, male habitus ventrally; D = N. kalashiani, male (holotype) habitus dorsally; E = ditto, female; F = N. magomedrasuli sp. n., male antennae; G = N. kalashiani, male." figureDoi="http://doi.org/10.5281/zenodo.10152226" httpUri="https://zenodo.org/record/10152226/files/figure.png" pageId="6" pageNumber="425">Figs 1D, and 1E</figureCitation>
). Antennae comparatively short, with only last distal antennomere extending beyond base of pronotum. Antennomeres 9–11 are widest.
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCC0FE5EDA26FDB8" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">
Pronotum transverse (1.55 times as wide as long), widest at basal third, 1.74 as wide as head (
<figureCitation id="13302A1EFFFB2F5CD8E5FE5FDC00FE28" box="[1250,1338,402,428]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="6" pageNumber="425">Fig. 2E</figureCitation>
). Lateral margins of pronotum very weakly sinuate near base and evenly rounded from widest part to apex; apical margin straight at middle and weakly emarginated near angles; base bisinuate, straight along scutellar shield. Lateral margins and base beaded, bead of anterior margin widely interrupted at middle. Disc evenly convex, not flattened along lateral margins. Puncturation of disc the same as on head.
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCC0FD8BDDB6FD48" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">
Scutellar shield helmet-like, with generally rounded lateral margins, but slightly emarginated near apex; densely finely punctured (
<figureCitation id="13302A1EFFFB2F5CDD7DFDA7D8E8FD00" box="[378,466,618,644]" captionStart="FIGURE 2" captionStartId="3.[151,250,1613,1637]" captionTargetBox="[151,1436,404,1587]" captionTargetId="figure-137@3.[151,1436,404,1588]" captionTargetPageId="3" captionText="FIGURE 2. Nalassus spp., details of structure. A = N. magomedrasuli sp. n., male (holotype), head and pronotum; B = ditto (paratype), female; C = ditto, male elytral intervals; D = N. kalashiani, male (holotype), head and pronotum; E = ditto, female; F = ditto, male elytral intervals." figureDoi="http://doi.org/10.5281/zenodo.10152228" httpUri="https://zenodo.org/record/10152228/files/figure.png" pageId="6" pageNumber="425">Fig. 2E</figureCitation>
). Elytra elongate-oval (1.4 times as long as wide), widest at middle, 2.07 times as wide as head, 11.8 times as wide and 2.6 times as long as pronotum. Elongate strial punctures connected by fine furrow. Intervals flat, intervals 2–4 wider, than other, contains 3–4 punctures in transverse section.
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCC0FD1ADDA4FC91" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">Pro and mesotarsi not widened, with comparatively short yellowish hairs on lateral and ventral sides. Abdominal ventrite 1 without hair brush at middle; abdominal ventrite 5 completely beaded at apex.</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCC0FCD3D836FC05" blockId="6.[151,1437,150,897]" pageId="6" pageNumber="425">
<emphasis id="B97FEA89FFFB2F5CDCC0FCD3D82BFCBC" bold="true" box="[199,273,798,824]" pageId="6" pageNumber="425">Notes.</emphasis>
This female specimen clearly belongs to 
<taxonomicName id="4C0B4D18FFFB2F5CDEE4FCD2DA42FCBC" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[739,888,798,824]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFB2F5CDEE4FCD2DA42FCBC" box="[739,888,798,824]" italics="true" pageId="6" pageNumber="425">N. kalashiani</emphasis>
</taxonomicName>
, which was known by 
<specimenCount id="9D0DFD12FFFB2F5CD87DFCD2DDD9FCBC" box="[1146,1251,798,825]" count="1" pageId="6" pageNumber="425" type="male">one male</specimenCount>
, by the structure of eyes having the same ratio of distance at widest part of head to distance between eyes and identical shape of pronotum.
</paragraph>
</subSubSection>
<caption id="DF746613FFFB2F5CDC9FFC7DDA7BF800" pageId="6" pageNumber="425" startId="6.[152,244,944,968]" targetType="table">
<paragraph id="8BB4369BFFFB2F5CDC9FFC7DDD2AFC43" blockId="6.[152,1420,944,1924]" box="[152,1040,944,968]" pageId="6" pageNumber="425">
<emphasis id="B97FEA89FFFB2F5CDC9FFC7DD835FC4C" bold="true" box="[152,271,944,968]" pageId="6" pageNumber="425">TABLE 1.</emphasis>
Differences between 
<taxonomicName id="4C0B4D18FFFB2F5CDDFFFC7DDA3FFC4C" authority="Nabozhenko &amp; Gadaborsheva, 2023" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[504,773,944,968]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. nov.">
<emphasis id="B97FEA89FFFB2F5CDDFFFC7DDA3FFC4C" box="[504,773,944,968]" italics="true" pageId="6" pageNumber="425">Nalassus magomedrasuli</emphasis>
</taxonomicName>
<emphasis id="B97FEA89FFFB2F5CDF0CFC7DDA7DFC4C" bold="true" box="[779,839,944,968]" pageId="6" pageNumber="425">
<taxonomicNameLabel id="A24C57F2FFFB2F5CDF0CFC7DDA7DFC4C" box="[779,839,944,968]" pageId="6" pageNumber="425" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C0B4D18FFFB2F5CDF7CFC7CDD33FC4C" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[891,1033,944,968]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFB2F5CDF7CFC7CDD33FC4C" box="[891,1033,944,968]" italics="true" pageId="6" pageNumber="425">N. kalashiani</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FC14DA9CFC74" blockId="6.[152,1420,944,1924]" box="[163,934,984,1008]" pageId="6" pageNumber="425">
<taxonomicName id="4C0B4D18FFFB2F5CDCA4FC14D864FC74" authority="Nabozhenko &amp; Gadaborsheva, 2023" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[163,350,984,1008]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="magomedrasuli" status="sp. n.">
<emphasis id="B97FEA89FFFB2F5CDCA4FC14D864FC74" box="[163,350,984,1008]" italics="true" pageId="6" pageNumber="425">N. magomedrasuli</emphasis>
</taxonomicName>
<emphasis id="B97FEA89FFFB2F5CDD63FC15D8A7FC74" bold="true" box="[356,413,984,1008]" pageId="6" pageNumber="425">
<taxonomicNameLabel id="A24C57F2FFFB2F5CDD63FC15D8A7FC74" box="[356,413,984,1008]" pageId="6" pageNumber="425" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C0B4D18FFFB2F5CDF19FC14DA9CFC74" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[798,934,984,1008]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFB2F5CDF19FC14DA9CFC74" box="[798,934,984,1008]" italics="true" pageId="6" pageNumber="425">N. kalashiani</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BB4369BFFFB2F5CDD5EFBCDDB62FB9C" blockId="6.[152,1420,944,1924]" box="[345,600,1024,1048]" pageId="6" pageNumber="425">Morphological structures</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FBE4DD8CFBDA" blockId="6.[152,1420,944,1924]" pageId="6" pageNumber="425">Eyes less convex. Ratio of distance at widest part of head to Eyes more convex. Ratio of distance at widest part of head to distance between eyes 1.36–1.37 (Figs 2A, B) distance between eyes 1.43 (Figs 2D, E)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FBA3DC27FB20" blockId="6.[152,1420,944,1924]" pageId="6" pageNumber="425">Sexual dimorphism in the structure of apical maxillary No sexual dimorphism in the structure of apical maxillary palpomere (much wider in male) (Figs 2A, B) palpomere (wide in male and female) (Figs 1D, E)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FB79DC23FB48" blockId="6.[152,1420,944,1924]" box="[163,1305,1204,1228]" pageId="6" pageNumber="425">Male antennomeres 2–7 less thickened (Fig. 1F) Male antennomeres 2–7 more thickened (Fig. 1G)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FB10DD6EFB70" blockId="6.[152,1420,944,1924]" box="[163,1108,1244,1268]" pageId="6" pageNumber="425">Pronotum widest slightly behind middle Pronotum widest at basal third</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FAC9DDD4FA99" blockId="6.[152,1420,944,1924]" box="[163,1262,1284,1309]" pageId="6" pageNumber="425">Disc of pronotum slightly narrowly flattened along lateral Disc of pronotum evenly convex (Figs 2D, E)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FAEFD840FABE" blockId="6.[152,1420,944,1924]" box="[163,378,1314,1338]" pageId="6" pageNumber="425">margins (Figs 2A, B)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FA87DA5EFA19" blockId="6.[152,1420,944,1924]" pageId="6" pageNumber="425">Male pro- and mesotarsi slightly widened, with Male pro- and mesotarsi little wider, with very dense and comparatively short yellowish hairs on lateral margins and long pubescence on lateral margins and ventral side (Figs ventral side (Figs 3C, D) 3G, H)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FA60DDBAFA41" blockId="6.[152,1420,944,1924]" box="[163,1152,1453,1477]" pageId="6" pageNumber="425">Male tibiae much thinner (Fig 1A) Male tibiae wider, robust (Fig. 1D)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FA18DC51FA69" blockId="6.[152,1420,944,1924]" box="[163,1387,1493,1518]" pageId="6" pageNumber="425">Scutellar shield with several very sparse punctures or Scutellar shield densely and finely punctured (Figs 2D, E)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4FA3FD8C6F98F" blockId="6.[152,1420,944,1924]" box="[163,508,1522,1547]" pageId="6" pageNumber="425">without puncturation (Figs 2A, B)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F9D6DC77F9B7" blockId="6.[152,1420,944,1924]" box="[163,1357,1563,1587]" pageId="6" pageNumber="425">Male elytral interstriae finely and sparsely punctured (Fig. Male elytral interstriae with slightly coarser and denser</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F9F5DAC4F9D4" blockId="6.[152,1420,944,1924]" box="[163,1022,1592,1616]" pageId="6" pageNumber="425">2C) puncturation (Fig. 2F)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F9ADDC5FF9FC" blockId="6.[152,1420,944,1924]" box="[163,1381,1632,1656]" pageId="6" pageNumber="425">Male abdomen only with one hair brush on ventrite 1 (Fig. Male abdomen with two hair brushes: larger on ventrite 1</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F9B3DD47F912" blockId="6.[152,1420,944,1924]" box="[163,1149,1662,1686]" pageId="6" pageNumber="425">3A) and smaller on ventrite 2 (Fig. 3B)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F96BDC47F97C" blockId="6.[152,1420,944,1924]" pageId="6" pageNumber="425">First abdominal ventrite with much coarser puncturation First abdominal ventrite with the same puncture size as on than on other surface; metacoxal process of abdominal other surface and together with metacoxal process much ventrite 1 with much sparser puncturation in apical part denser and finer (in in comparative species) punctured (Fig.</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F933DA78F892" blockId="6.[152,1420,944,1924]" box="[163,834,1790,1814]" pageId="6" pageNumber="425">(Fig. 3E) 3F)</paragraph>
<paragraph id="8BB4369BFFFB2F5CDDA8F8EBDB39F8BA" blockId="6.[152,1420,944,1924]" box="[431,515,1830,1854]" pageId="6" pageNumber="425">Habitats</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F883DC41F8E2" blockId="6.[152,1420,944,1924]" box="[163,1403,1870,1894]" pageId="6" pageNumber="425">Limestone montaine steppes with phryganoid vegetation Alpine meadows et least in Ingushetia (2100–2200 m) (Fig.</paragraph>
<paragraph id="8BB4369BFFFB2F5CDCA4F8A6DA7BF800" blockId="6.[152,1420,944,1924]" box="[163,833,1899,1924]" pageId="6" pageNumber="425">(1100–1500 m) (Fig. 4E) 4F)</paragraph>
</caption>
<subSubSection id="C3116510FFFB2F5DDCC0F87EDA67FE0D" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425" type="distribution">
<paragraph id="8BB4369BFFFB2F5DDCC0F87EDA67FE0D" blockId="6.[151,1437,1971,2034]" lastBlockId="7.[151,1436,151,393]" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425">
<emphasis id="B97FEA89FFFB2F5CDCC0F87ED8C8F849" bold="true" box="[199,498,1971,1997]" pageId="6" pageNumber="425">Distribution and habitat.</emphasis>
<collectingCountry id="F31C760BFFFB2F5CDDFEF87EDB7DF849" box="[505,583,1971,1997]" name="Russia" pageId="6" pageNumber="425">Russia</collectingCountry>
, Chechen Republic and the Republic of 
<collectingRegion id="49CFF879FFFB2F5CD80AF87EDDBFF849" box="[1037,1157,1971,1997]" country="Russia" name="Ingushetiya" pageId="6" pageNumber="425">Ingushetia</collectingRegion>
. We don’t known, where the 
<typeStatus id="54B08839FFFB2F5CDCC6F81AD819F875" box="[193,291,2007,2033]" pageId="6" pageNumber="425" type="holotype">holotype</typeStatus>
was collected. The collector, Dr Mark Kalashian also does not remember exactly locality, but he collected beetles in the mountains around Itum-Kale (they reach 
<quantity id="4CF39B7EFFFA2F5DDF40FF5ADA9AFF35" box="[839,928,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="426" unit="m" value="2200.0">2200 m</quantity>
). We collected only 
<specimenCount id="9D0DFD12FFFA2F5DD885FF5ADDC7FF35" box="[1154,1277,151,177]" count="1" pageId="7" pageNumber="426" type="female">one female</specimenCount>
of 
<taxonomicName id="4C0B4D18FFFA2F5DD925FF5AD981FF50" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFA2F5DD925FF5AD981FF50" italics="true" pageId="7" pageNumber="426">N. kalashiani</emphasis>
</taxonomicName>
under stone in alpine meadow (
<quantity id="4CF39B7EFFFA2F5DDE23FF76DB45FF51" box="[548,639,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="426" unit="m" value="2200.0">2200 m</quantity>
). This is the second species of the genus from 
<collectingRegion id="49CFF879FFFA2F5DD88BFF76DC3EFF51" box="[1164,1284,187,213]" country="Russia" name="Ingushetiya" pageId="7" pageNumber="426">Ingushetia</collectingRegion>
. Before, only 
<taxonomicName id="4C0B4D18FFFA2F5DDC90FF12DB96FF7D" authority="(Allard, 1876)" baseAuthorityName="Allard" baseAuthorityYear="1876" box="[151,684,223,249]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="diteras" subGenus="Caucasonotus">
<emphasis id="B97FEA89FFFA2F5DDC90FF12D9C6FF7D" box="[151,252,223,249]" italics="true" pageId="7" pageNumber="426">Nalassus</emphasis>
(
<emphasis id="B97FEA89FFFA2F5DDD0CFF12D890FF7D" box="[267,426,223,249]" italics="true" pageId="7" pageNumber="426">Caucasonotus</emphasis>
) 
<emphasis id="B97FEA89FFFA2F5DDDBEFF12DB3DFF7D" box="[441,519,223,249]" italics="true" pageId="7" pageNumber="426">diteras</emphasis>
(Allard, 1876)
</taxonomicName>
was known from this republic (
<bibRefCitation id="EF9A4B6AFFFA2F5DD80FFF12DDE2FF7D" author="Nabozhenko, M. V." box="[1032,1240,223,249]" pageId="7" pageNumber="426" pagination="909 - 942" refId="ref4250" refString="Nabozhenko, M. V. (2001) On the classification of the tenebrionid tribe Helopini, with a review of the genera Nalassus Mulsant and Odocnemis Allard (Coleoptera, Tenebrionidae) of the European part of CIS and the Caucasus. Entomologicheskoe obozrenie, 80 (3), 627 - 668. [in Russian; English translation: Entomological Review, 81 (8), 909 - 942]" type="journal article" year="2001">Nabozhenko 2001</bibRefCitation>
, 
<bibRefCitation id="EF9A4B6AFFFA2F5DD8E7FF12D82CFE99" author="Abdurakhmanov, G. M. &amp; Tochieva, F. T. &amp; Tochiev, T. Yu." pageId="7" pageNumber="426" refId="ref4207" refString="Abdurakhmanov, G. M., Tochieva, F. T. &amp; Tochiev, T. Yu. (2005) Darkling beetles of Dzheyrakh and Targim basins. Yupiter press, Makhachkala, 120 pp. [in Russian]" type="book" year="2005">
Abdurakhmanov 
<emphasis id="B97FEA89FFFA2F5DDC90FECED9EBFE99" box="[151,209,259,285]" italics="true" pageId="7" pageNumber="426">et al.</emphasis>
2005
</bibRefCitation>
). We collected the latter species lower than the type locality of 
<taxonomicName id="4C0B4D18FFFA2F5DDFECFECEDDB8FE99" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[1003,1154,259,285]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFA2F5DDFECFECEDDB8FE99" box="[1003,1154,259,285]" italics="true" pageId="7" pageNumber="426">N. kalashiani</emphasis>
</taxonomicName>
, at altitudes from 
<quantity id="4CF39B7EFFFA2F5DD953FECED9CBFEC5" metricMagnitude="2" metricUnit="m" metricValue="9.9" metricValueMax="13.5" metricValueMin="6.3" pageId="7" pageNumber="426" unit="m" value="990.0" valueMax="1350.0" valueMin="630.0">630 to 1350 m</quantity>
. Thus, these two species are probably allopatric. Considering the exceptional rarity of 
<taxonomicName id="4C0B4D18FFFA2F5DD8D4FEEADC51FEC5" authorityName="Nabozhenko &amp; Gadaborsheva" authorityYear="2023" box="[1235,1387,295,321]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="kalashiani">
<emphasis id="B97FEA89FFFA2F5DD8D4FEEADC51FEC5" box="[1235,1387,295,321]" italics="true" pageId="7" pageNumber="426">N. kalashiani</emphasis>
</taxonomicName>
and sometimes intensive grazing of livestock in its habitats, we propose this species for inclusion in the regional red books of the Chechen Republic and the Republic of 
<collectingRegion id="49CFF879FFFA2F5DDEE6FEA2DA63FE0D" box="[737,857,367,393]" country="Russia" name="Ingushetiya" pageId="7" pageNumber="426">Ingushetia</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>