treatments-xml/data/03/A2/87/03A2878D6E23CA2EFFCDF9CBBB8B43FE.xml

<document id="3108C46A9119EE45EB7F11CBA6741A0D" ID-DOI="10.26107/RBZ-2021-0008" ID-ISSN="2345-7600" ID-Zenodo-Dep="13256870" ID-ZooBank="0EF4EEFD-D463-407A-8168-719354912D8A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723049820326" checkinUser="felipe" docAuthor="Dumrongrojwattana, Pongrat, , Sirilan Chuenit, Wongkamhaeng, &amp; Koraon" docDate="2021" docId="03A2878D6E23CA2EFFCDF9CBBB8B43FE" docLanguage="en" docName="RafflesBZool.69.102-108.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng 2021, new species" docType="treatment" docUuid="92696099-DD77-42A3-B0EF-09524B49CE8D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="106" masterDocId="FF9BFFF56E21CA2AFFB4FFF8B90B4603" masterDocTitle="A new species of the world’s smallest cave snail of the genus Angustopila Jochum, Slapnik &amp; Páll-Gergely in Jochum, et al., 2014 (Gastropoda: Hypselostomatidae) from eastern Thailand" masterLastPageNumber="108" masterPageNumber="102" pageNumber="104" updateTime="1723053786272" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B076D9C0C8A416D9CFDE04BCA0CA7753">A new species of the world’s smallest cave snail of the genus Angustopila Jochum, Slapnik &amp; Páll-Gergely in Jochum, et al., 2014 (Gastropoda: Hypselostomatidae) from eastern Thailand</mods:title>
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<mods:namePart id="FE86ECC8C95997E31E5B7B5C98795A0F">Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:namePart id="DB95001D90691289D1B5EA99668779B5">, Sirilan Chuenit</mods:namePart>
</mods:name>
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<mods:namePart id="B32B8DD6D5A46E041ED162C7A4EF9978">Wongkamhaeng,</mods:namePart>
</mods:name>
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<mods:namePart id="3EF3D42D4AF9EE6DA7FC59EA654AB5AD">Koraon</mods:namePart>
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<mods:title id="A752DFADAC5E03F9D49EEE841CD64CA5">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="77C130DBA02C0D55A8C04E80EEF4DD71">2021</mods:date>
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<mods:number id="54F4F25568AD3F33E9EC5761B9741C87">2021-03-15</mods:number>
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<mods:identifier id="2482367F503DC6DF138AD42C51AA44F7" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03A2878D6E23CA2EFFCDF9CBBB8B43FE" ID-Zenodo-Dep="13256922" LSID="urn:lsid:zoobank.org:act:92696099-DD77-42A3-B0EF-09524B49CE8D" httpUri="http://treatment.plazi.org/id/03A2878D6E23CA2EFFCDF9CBBB8B43FE" lastPageId="4" lastPageNumber="106" pageId="2" pageNumber="104">
<subSubSection id="C31165106E23CA28FFCDF9CBBB6C4068" pageId="2" pageNumber="104" type="nomenclature">
<paragraph id="8BB4369B6E23CA28FFCDF9CBBB6C4068" blockId="2.[121,743,1587,1675]" pageId="2" pageNumber="104">
<heading id="D0FC81F76E23CA28FFCDF9CBBB6C4068" bold="true" centered="true" fontSize="10" level="2" pageId="2" pageNumber="104" reason="2">
<emphasis id="B97FEA896E23CA28FFCDF9CBBB6C4068" bold="true" pageId="2" pageNumber="104">
<taxonomicName id="4C0B4D186E23CA28FFCDF9CBB8DF4068" ID-CoL="873PZ" authority="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityName="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityYear="2021" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="104" phylum="Mollusca" rank="species" species="pallgergelyi" status="sp. nov.">
<emphasis id="B97FEA896E23CA28FFCDF9CBB8834048" bold="true" box="[121,392,1587,1611]" italics="true" pageId="2" pageNumber="104">Angustopila pallgergelyi</emphasis>
Dumrongrojwattana, Chuenit &amp; Wongkamhaeng
</taxonomicName>
, 
<taxonomicNameLabel id="A24C57F26E23CA28FE56F9ABBB6C4068" box="[482,615,1619,1643]" pageId="2" pageNumber="104" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FEEDF98BBB0D4088" box="[345,518,1651,1675]" pageId="2" pageNumber="104" type="description">
<paragraph id="8BB4369B6E23CA28FEEDF98BBB0D4088" blockId="2.[121,743,1587,1675]" box="[345,518,1651,1675]" pageId="2" pageNumber="104">
(
<figureCitation id="13302A1E6E23CA28FED5F98BB8A54088" box="[353,430,1651,1675]" captionStart="Fig" captionStartId="3.[136,171,2064,2086]" captionTargetBox="[148,1462,189,2048]" captionTargetId="figure-7@3.[148,1462,189,2048]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species (ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm." pageId="2" pageNumber="104">Figs. 2</figureCitation>
, 
<figureCitation id="13302A1E6E23CA28FE08F98BB8F54088" box="[444,510,1651,1675]" captionStart="Fig" captionStartId="4.[813,848,1470,1492]" captionTargetBox="[813,1453,549,1455]" captionTargetId="figure-153@4.[813,1453,549,1455]" captionTargetPageId="4" captionText="Fig. 3. A, B, Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species, living snail at Phet Pho Thong Cave, Sa Kaeo, approximately 300 metres from the entrance. Photograph: Rattanwadee Tekavong." pageId="2" pageNumber="104">3A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FFC5F94BB82841E8" pageId="2" pageNumber="104" type="materials_examined">
<paragraph id="8BB4369B6E23CA28FFC5F94BB82841E8" blockId="2.[111,752,1715,2027]" pageId="2" pageNumber="104">
<emphasis id="B97FEA896E23CA28FFC5F94BB82640C8" bold="true" box="[113,301,1715,1739]" pageId="2" pageNumber="104">Type material.</emphasis>
<materialsCitation id="3B633CC66E23CA28FE8EF94BB9F14148" collectingDate="2018-07-05" collectionCode="ZRCBUU" collectorName="S. Chuenit" country="Thailand" elevation="280" latitude="13.413722" location="Tham Phet Pho Thong" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" pageId="2" pageNumber="104" specimenCode="ZRCBUU 0699" specimenCount="1" stateProvince="Sa Kaeo" typeStatus="holotype">
<typeStatus id="54B088396E23CA28FE8EF94BB8A040C8" box="[314,427,1715,1739]" pageId="2" pageNumber="104" type="holotype">Holotype</typeStatus>
. 1 shell (
<specimenCode id="DBAD9EE06E23CA28FD95F94BBBE940C8" box="[545,738,1715,1739]" collectionCode="ZRCBUU" pageId="2" pageNumber="104">ZRCBUU 0699</specimenCode>
), 
<collectingCountry id="F31C760B6E23CA28FFC5F92BB80F40E8" box="[113,260,1747,1771]" name="Thailand" pageId="2" pageNumber="104">THAILAND</collectingCountry>
, 
<collectingRegion id="49CFF8796E23CA28FEA7F92BB8E240E8" box="[275,489,1747,1771]" country="Thailand" name="Sa Kaeo" pageId="2" pageNumber="104">Sa Kaeo Province</collectingRegion>
, 
<collectingMunicipality id="6BD0ACE16E23CA28FE4DF92BBBE040E8" box="[505,747,1747,1771]" pageId="2" pageNumber="104">Klong Haad District</collectingMunicipality>
, 
<location id="8ED460406E23CA28FFC5F90BB8664108" LSID="urn:lsid:plazi:treatment:03A2878D6E23CA2EFFCDF9CBBB8B43FE:8ED460406E23CA28FFC5F90BB8664108" box="[113,365,1779,1803]" country="Thailand" latitude="13.413722" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" name="Tham Phet Pho Thong" pageId="2" pageNumber="104" stateProvince="Sa Kaeo">Tham Phet Pho Thong</location>
(Cave), an isolated limestone hill; 
<geoCoordinate id="EE3F505C6E23CA28FFC5F8EAB80F4128" box="[113,260,1810,1835]" degrees="13" direction="north" minutes="24" orientation="latitude" pageId="2" pageNumber="104" precision="1" seconds="49. 4" value="13.413722">13°24′49.4″N</geoCoordinate>
<geoCoordinate id="EE3F505C6E23CA28FEBFF8EAB8A04128" box="[267,427,1810,1835]" degrees="102" direction="east" minutes="19" orientation="longitude" pageId="2" pageNumber="104" precision="1" seconds="38. 0" value="102.327225">102°19′38.0″E</geoCoordinate>
; 
<elevation id="0026D1A86E23CA28FE03F8EBBB3F4128" box="[439,564,1811,1835]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="104" unit="m" value="280.0">
<quantity id="4CF39B7E6E23CA28FE03F8EBB8F04128" box="[439,507,1811,1835]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="104" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
, coll. 
<collectorName id="26FE534D6E23CA28FDC3F8EBBBE04128" box="[631,747,1811,1835]" pageId="2" pageNumber="104">S. Chuenit</collectorName>
, 
<date id="FFB5105B6E23CA28FFC5F8CBB9F14148" box="[113,250,1843,1867]" pageId="2" pageNumber="104" value="2018-07-05">
<collectingDate id="EFF1E9B36E23CA28FFC5F8CBB9F14148" box="[113,250,1843,1867]" pageId="2" pageNumber="104" value="2018-07-05">5 July 2018</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B633CC66E23CA28FEBCF8CCBB864188" collectingDate="2019-09-15" collectionCode="ZRC" collectorName="P. Dumrongrojwattana" country="Thailand" elevation="280" latitude="13.413722" location="Tham Phet Pho Thong" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" pageId="2" pageNumber="104" specimenCode="ZRCBUU 0700" specimenCount="1" stateProvince="Sa Kaeo" typeStatus="paratype">
<typeStatus id="54B088396E23CA28FEBCF8CCB87D414F" box="[264,374,1844,1868]" pageId="2" pageNumber="104" type="paratype">Paratypes</typeStatus>
. 3 shells (
<specimenCode id="DBAD9EE06E23CA28FE58F8CBBB954148" box="[492,670,1843,1867]" collectionCode="ZRCBUU" pageId="2" pageNumber="104">ZRCBUU 0700</specimenCode>
) and 2 shells (
<collectionCode id="ED1AAE5E6E23CA28FF0EF8ABB9FA4168" box="[186,241,1875,1899]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="104">ZRC</collectionCode>
.MOL.020945) same collecting data as holotype, coll. 
<collectorName id="26FE534D6E23CA28FF1CF88CB8AF4188" box="[168,420,1907,1931]" pageId="2" pageNumber="104">P. Dumrongrojwattana</collectorName>
, 
<date id="FFB5105B6E23CA28FE06F88BBB864188" box="[434,653,1907,1931]" pageId="2" pageNumber="104" value="2019-09-15">
<collectingDate id="EFF1E9B36E23CA28FE06F88BBB864188" box="[434,653,1907,1931]" pageId="2" pageNumber="104" value="2019-09-15">15 September 2019</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B633CC66E23CA28FD2FF88CB82B41E8" collectingDate="2020-12-12" collectionCode="ZRCBUU" collectorName="P. Dumrongrojwattana" country="Thailand" elevation="280" latitude="13.413722" location="Tham Phet Pho Thong" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" pageId="2" pageNumber="104" specimenCode="ZRCBUU 0730" specimenCount="1" stateProvince="Sa Kaeo" typeStatus="paratype">
7 shells (
<specimenCode id="DBAD9EE06E23CA28FFCCF86BB82C41A8" box="[120,295,1939,1963]" collectionCode="ZRCBUU" pageId="2" pageNumber="104">ZRCBUU 0730</specimenCode>
) and 2 shells (THNHM-Iv-17645), same collecting data as holotype, coll. 
<collectorName id="26FE534D6E23CA28FE67F84CBBC141C8" box="[467,714,1971,1995]" pageId="2" pageNumber="104">P. Dumrongrojwattana</collectorName>
, 
<date id="FFB5105B6E23CA28FD61F84BB82B41E8" pageId="2" pageNumber="104" value="2020-12-12">
<collectingDate id="EFF1E9B36E23CA28FD61F84BB82B41E8" pageId="2" pageNumber="104" value="2020-12-12">12 December 2020</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FC99FC82BD994252" pageId="2" pageNumber="104" type="description">
<paragraph id="8BB4369B6E23CA28FC99FC82BD994252" blockId="2.[813,1452,890,1106]" pageId="2" pageNumber="104">
<emphasis id="B97FEA896E23CA28FC99FC82BAD04591" bold="true" box="[813,987,890,914]" pageId="2" pageNumber="104">Measurements.</emphasis>
<typeStatus id="54B088396E23CA28FC56FC82BD4E4591" box="[994,1093,890,914]" pageId="2" pageNumber="104" type="holotype">Holotype</typeStatus>
. SH = 
<quantity id="4CF39B7E6E23CA28FB27FC82BDFF4592" box="[1171,1268,890,914]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="104" unit="mm" value="0.64">0.64 mm</quantity>
, SW = 
<quantity id="4CF39B7E6E23CA28FAF1FC82BCAC4592" box="[1349,1447,890,914]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="104" unit="mm" value="0.76">0.76 mm</quantity>
, AH = 
<quantity id="4CF39B7E6E23CA28FCC0FC62BAD345B2" box="[884,984,922,946]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="104" unit="mm" value="0.27">0.27 mm</quantity>
, AW = 0.27, SW/SH = 1.19, and AW/AH = 1.01. 
<typeStatus id="54B088396E23CA28FC32FC42BAE445D1" box="[902,1007,954,978]" pageId="2" pageNumber="104" type="paratype">Paratypes</typeStatus>
. SH = 0.59– 
<quantity id="4CF39B7E6E23CA28FB37FC42BDEC45D2" box="[1155,1255,954,978]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="104" unit="mm" value="0.71">0.71 mm</quantity>
(0.66± 
<quantity id="4CF39B7E6E23CA28FA8CFC42BC9745D2" box="[1336,1436,954,978]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="104" unit="mm" value="0.03">0.03 mm</quantity>
), SW = 0.76– 
<quantity id="4CF39B7E6E23CA28FC19FC22BD0045F2" box="[941,1035,986,1010]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="104" unit="mm" value="0.82">0.82 mm</quantity>
(1.25± 
<quantity id="4CF39B7E6E23CA28FBEDFC22BDBC45F2" box="[1113,1207,986,1010]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="104" unit="mm" value="0.03">0.03 mm</quantity>
), AH = 0.26– 
<quantity id="4CF39B7E6E23CA28FAF9FC22BCA045F2" box="[1357,1451,986,1010]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="104" unit="mm" value="0.31">0.31 mm</quantity>
(0.28± 
<quantity id="4CF39B7E6E23CA28FCC0FC02BAD34212" box="[884,984,1018,1042]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
), AW = 0.25– 
<quantity id="4CF39B7E6E23CA28FB35FC02BDEE4212" box="[1153,1253,1018,1042]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="104" unit="mm" value="0.31">0.31 mm</quantity>
(0.28± 
<quantity id="4CF39B7E6E23CA28FA83FC02BC904212" box="[1335,1435,1018,1042]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
), SW/SH = 1.19–1.35 (1.25±0.08), and AW/AH = 0.90–1.08 (0.98±0.06). See 
<tableCitation id="C68903206E23CA28FC5DFBC2BD4D4251" box="[1001,1094,1082,1106]" captionStart="Table 1" captionStartId="2.[113,168,189,211]" captionText="Table 1. Shell measurements (mm) for Angustopila pallgergelyi, new species, from the type locality." pageId="2" pageNumber="104">Tables 1</tableCitation>
and 
<tableCitation id="C68903206E23CA28FBCAFBC2BD804251" box="[1150,1163,1082,1106]" captionStart="Table 2" captionStartId="4.[113,168,189,211]" captionText="Table 2. The average, minimum value (min), a maximum value (max), a standard deviation of a set of values (stdev) for Angustopila pallgergelyi, new species (n = 15)." pageId="2" pageNumber="104">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FC99FB82BAB542F1" pageId="2" pageNumber="104" type="materials_examined">
<paragraph id="8BB4369B6E23CA28FC99FB82BAB542F1" blockId="2.[811,1452,1146,1266]" pageId="2" pageNumber="104">
<materialsCitation id="3B633CC66E23CA28FC99FB82BAB242F1" country="Thailand" elevation="280" latitude="13.413722" location="Tham Phet Pho Thong" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" pageId="2" pageNumber="104" specimenCount="1" stateProvince="Sa Kaeo" typeStatus="holotype">
<emphasis id="B97FEA896E23CA28FC99FB82BAC64291" bold="true" box="[813,973,1146,1170]" pageId="2" pageNumber="104">
<typeStatus id="54B088396E23CA28FC99FB82BA634291" box="[813,872,1146,1170]" pageId="2" pageNumber="104">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31C760B6E23CA28FC6CFB82BD304291" box="[984,1083,1146,1170]" name="Thailand" pageId="2" pageNumber="104">Thailand</collectingCountry>
, 
<collectingRegion id="49CFF8796E23CA28FBFDFB82BC1E4291" box="[1097,1301,1146,1170]" country="Thailand" name="Sa Kaeo" pageId="2" pageNumber="104">Sa Kaeo Province</collectingRegion>
, 
<collectingMunicipality id="6BD0ACE16E23CA28FA90FB82BA7442B1" pageId="2" pageNumber="104">Klong Haad District</collectingMunicipality>
, 
<location id="8ED460406E23CA28FC3CFB62BD7F42B1" LSID="urn:lsid:plazi:treatment:03A2878D6E23CA2EFFCDF9CBBB8B43FE:8ED460406E23CA28FC3CFB62BD7F42B1" box="[904,1140,1178,1202]" country="Thailand" latitude="13.413722" longLatPrecision="1" longitude="102.327225" municipality="Klong Haad District" name="Tham Phet Pho Thong" pageId="2" pageNumber="104" stateProvince="Sa Kaeo">Tham Phet Pho Thong</location>
(Cave), an isolated limestone hill; 
<geoCoordinate id="EE3F505C6E23CA28FCEAFB40BAF942D2" box="[862,1010,1208,1233]" degrees="13" direction="north" minutes="24" orientation="latitude" pageId="2" pageNumber="104" precision="1" seconds="49. 4" value="13.413722">13°24′49.4″N</geoCoordinate>
<geoCoordinate id="EE3F505C6E23CA28FC4DFB40BD9142D2" box="[1017,1178,1208,1233]" degrees="102" direction="east" minutes="19" orientation="longitude" pageId="2" pageNumber="104" precision="1" seconds="38. 0" value="102.327225">102°19′38.0″E</geoCoordinate>
; 
<elevation id="0026D1A86E23CA28FB11FB42BC2842D2" box="[1189,1315,1210,1234]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="104" unit="m" value="280.0">
<quantity id="4CF39B7E6E23CA28FB11FB42BDE142D2" box="[1189,1258,1210,1234]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="104" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
(locality no. 
<quantity id="4CF39B7E6E23CA28FC99FB22BA6D42F1" box="[813,870,1242,1266]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="104" unit="in" value="10.0">10 in</quantity>
<figureCitation id="13302A1E6E23CA28FCDAFB22BAA542F1" box="[878,942,1242,1266]" captionStart="Fig" captionStartId="1.[136,171,2064,2086]" captionTargetBox="[139,1475,192,2048]" captionTargetId="figure-8@1.[122,1483,177,2067]" captionTargetPageId="1" captionText="Fig. 1. Distribution map of Angustopila species. 1, A. huoyani; 2, A. dominikae; 3, A. subelevata; 4, A. szekeresi; 5, A. fabella; 6, A. singuladentis; 7, A. tamlod; 8, A. elevata; 9, A. concava; 10, Angustopila pallgergelyi, new species." pageId="2" pageNumber="104">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FC99FAE2BCA14372" pageId="2" pageNumber="104" type="etymology">
<paragraph id="8BB4369B6E23CA28FC99FAE2BCA14372" blockId="2.[813,1452,1306,1394]" pageId="2" pageNumber="104">
<emphasis id="B97FEA896E23CA28FC99FAE2BAA74331" bold="true" box="[813,940,1306,1330]" pageId="2" pageNumber="104">Etymology.</emphasis>
This new species is named in honour of Dr.Barna Páll-Gergely, the Hungarian malacologist, in appreciation of his significant contributions to the study of microsnails.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA28FC99FA62BC7E40D2" pageId="2" pageNumber="104" type="description">
<paragraph id="8BB4369B6E23CA28FC99FA62BC7E40D2" blockId="2.[811,1452,1434,1746]" pageId="2" pageNumber="104">
<emphasis id="B97FEA896E23CA28FC99FA62BABD43B1" bold="true" box="[813,950,1434,1458]" pageId="2" pageNumber="104">Description.</emphasis>
Shell minute, lenticular, white, apex blunt, spire lenticular with deep sutures (
<figureCitation id="13302A1E6E23CA28FBDFFA42BDC043D1" box="[1131,1227,1466,1490]" captionStart="Fig" captionStartId="3.[136,171,2064,2086]" captionTargetBox="[148,1462,189,2048]" captionTargetId="figure-7@3.[148,1462,189,2048]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species (ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm." pageId="2" pageNumber="104">Figs. 2A</figureCitation>
, 
<figureCitation id="13302A1E6E23CA28FB6CFA42BC1143D2" box="[1240,1306,1466,1490]" captionStart="Fig" captionStartId="4.[813,848,1470,1492]" captionTargetBox="[813,1453,549,1455]" captionTargetId="figure-153@4.[813,1453,549,1455]" captionTargetPageId="4" captionText="Fig. 3. A, B, Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species, living snail at Phet Pho Thong Cave, Sa Kaeo, approximately 300 metres from the entrance. Photograph: Rattanwadee Tekavong." pageId="2" pageNumber="104">3A, B</figureCitation>
); protoconch about one whorl, surface finely reticulate (
<figureCitation id="13302A1E6E23CA28FAABFA22BC9143F1" box="[1311,1434,1498,1522]" captionStart="Fig" captionStartId="3.[136,171,2064,2086]" captionTargetBox="[148,1462,189,2048]" captionTargetId="figure-7@3.[148,1462,189,2048]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species (ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm." pageId="2" pageNumber="104">Fig. 2B, C</figureCitation>
); teleoconch 4–4¼ rounded whorls, irregularly spaced growth lines crossed by rows of fine spiral threads forming fine reticulate sculpture (
<figureCitation id="13302A1E6E23CA28FBAFF9C2BD784051" box="[1051,1139,1594,1618]" captionStart="Fig" captionStartId="3.[136,171,2064,2086]" captionTargetBox="[148,1462,189,2048]" captionTargetId="figure-7@3.[148,1462,189,2048]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species (ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm." pageId="2" pageNumber="104">Fig. 2D</figureCitation>
); aperture broadly kidney-shaped, peristome slightly thickened, slightly expanded, slightly prosocline, detached from the body whorl; apertural teeth well developed, one parietal and one palatal directly opposite (
<figureCitation id="13302A1E6E23CA28FC2DF942BAE240D1" box="[921,1001,1722,1746]" captionStart="Fig" captionStartId="3.[136,171,2064,2086]" captionTargetBox="[148,1462,189,2048]" captionTargetId="figure-7@3.[148,1462,189,2048]" captionTargetPageId="3" captionText="Fig. 2. Holotype of Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species (ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm." pageId="2" pageNumber="104">Fig. 2E</figureCitation>
); umbilicus deep, relatively narrow.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E23CA2EFC99F902B83D455F" lastPageId="4" lastPageNumber="106" pageId="2" pageNumber="104" type="diagnosis">
<paragraph id="8BB4369B6E23CA2EFC99F902B83D455F" blockId="2.[813,1452,1786,2034]" lastBlockId="4.[113,753,549,861]" lastPageId="4" lastPageNumber="106" pageId="2" pageNumber="104">
<emphasis id="B97FEA896E23CA28FC99F902BD214111" bold="true" box="[813,1066,1786,1810]" pageId="2" pageNumber="104">Differential diagnosis.</emphasis>
<taxonomicName id="4C0B4D186E23CA28FB86F902BC364111" authorityName="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityYear="2021" box="[1074,1341,1786,1810]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="104" phylum="Mollusca" rank="species" species="pallgergelyi">
<emphasis id="B97FEA896E23CA28FB86F902BC364111" box="[1074,1341,1786,1810]" italics="true" pageId="2" pageNumber="104">Angustopila pallgergelyi</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C0B4D186E23CA28FCFEF8E2BAF34131" authorityName="Inkhavilay &amp; Panha" authorityYear="2016" box="[842,1016,1818,1842]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="104" phylum="Mollusca" rank="species" species="singuladentis">
<emphasis id="B97FEA896E23CA28FCFEF8E2BAF34131" box="[842,1016,1818,1842]" italics="true" pageId="2" pageNumber="104">A. singuladentis</emphasis>
</taxonomicName>
by having a white shell, nearly smooth protoconch sculpture, and the teleoconch sculpture having irregular growth lines crossed with fine spiral threads, peristome slightly thickened and slightly expanded. The new species differs from 
<taxonomicName id="4C0B4D186E23CA28FB87F862BDD441B1" authorityName="Inkhavilay &amp; Panha" authorityYear="2016" box="[1075,1247,1946,1970]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="104" phylum="Mollusca" rank="species" species="singuladentis">
<emphasis id="B97FEA896E23CA28FB87F862BDD441B1" box="[1075,1247,1946,1970]" italics="true" pageId="2" pageNumber="104">A. singuladentis</emphasis>
</taxonomicName>
in some significant ways. The aperture is kidney-shaped (vs. circular in 
<taxonomicName id="4C0B4D186E23CA28FA20F842BAB041F1" authorityName="Inkhavilay &amp; Panha" authorityYear="2016" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="104" phylum="Mollusca" rank="species" species="singuladentis">
<emphasis id="B97FEA896E23CA28FA20F842BAB041F1" italics="true" pageId="2" pageNumber="104">A. singuladentis</emphasis>
</taxonomicName>
); apertural teeth consist of parietal and palatal teeth (vs. only parietal tooth in 
<taxonomicName id="4C0B4D186E25CA2EFE61FDDDBB8E443E" authorityName="Inkhavilay &amp; Panha" authorityYear="2016" box="[469,645,549,573]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="singuladentis">
<emphasis id="B97FEA896E25CA2EFE61FDDDBB8E443E" box="[469,645,549,573]" italics="true" pageId="4" pageNumber="106">A. singuladentis</emphasis>
</taxonomicName>
), and the whorl number ranges from 4–4.5 (vs. 
<quantity id="4CF39B7E6E25CA2EFE49FDBDBB33445E" box="[509,568,581,605]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="4" pageNumber="106" unit="in" value="3.5">3.5 in</quantity>
<taxonomicName id="4C0B4D186E25CA2EFD89FDBDBBEC445E" authorityName="Inkhavilay &amp; Panha" authorityYear="2016" box="[573,743,581,605]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="singuladentis">
<emphasis id="B97FEA896E25CA2EFD89FDBDBBEC445E" box="[573,743,581,605]" italics="true" pageId="4" pageNumber="106">A. singuladentis</emphasis>
</taxonomicName>
) (
<tableCitation id="C68903206E25CA2EFFCEFD9DB9C6447E" box="[122,205,613,637]" captionStart="Table 3" captionStartId="5.[79,134,471,494]" captionText="Table 3. Comparison of essential morphological traits and ecological information among Angustopila species." pageId="4" pageNumber="106">Table 3</tableCitation>
). The apertural teeth of 
<taxonomicName id="4C0B4D186E25CA2EFE69FD9DBB75447E" authorityName="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityYear="2021" box="[477,638,613,637]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="pallgergelyi">
<emphasis id="B97FEA896E25CA2EFE69FD9DBB75447E" box="[477,638,613,637]" italics="true" pageId="4" pageNumber="106">A. pallgergelyi</emphasis>
</taxonomicName>
consist of parietal and palatal teeth, and share this characteristic with only 
<taxonomicName id="4C0B4D186E25CA2EFF18FD5DB83444BE" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" box="[172,319,677,701]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="dominikae">
<emphasis id="B97FEA896E25CA2EFF18FD5DB83444BE" box="[172,319,677,701]" italics="true" pageId="4" pageNumber="106">A. dominikae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0B4D186E25CA2EFEE4FD5DB8C344BE" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[336,456,677,701]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="huoyani">
<emphasis id="B97FEA896E25CA2EFEE4FD5DB8C344BE" box="[336,456,677,701]" italics="true" pageId="4" pageNumber="106">A. huoyani</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C0B4D186E25CA2EFDBFFD5DBB7344BE" authorityName="Panha &amp; Burch" authorityYear="1999" box="[523,632,677,701]" class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="tamlod">
<emphasis id="B97FEA896E25CA2EFDBFFD5DBB7344BE" box="[523,632,677,701]" italics="true" pageId="4" pageNumber="106">A. tamlod</emphasis>
</taxonomicName>
. This new species can be distinguished from these three species by having teleoconch sculpture irregular growth lines crossed with fine spiral threads and the umbilicus deep and relatively narrow. In contrast, the other species have spiral threads and narrow umbilicus.
</paragraph>
</subSubSection>
<caption id="DF7466136E22CA29FF3CF7E8BD504E41" pageId="3" pageNumber="105" startId="3.[136,171,2064,2086]" targetBox="[148,1462,189,2048]" targetPageId="3" targetType="figure">
<paragraph id="8BB4369B6E22CA29FF3CF7E8BD504E41" blockId="3.[136,1475,2064,2115]" pageId="3" pageNumber="105">
Fig. 2. Holotype of 
<taxonomicName id="4C0B4D186E22CA29FEE6F7E8BD254E25" authority="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityName="Dumrongrojwattana, Chuenit &amp; Wongkamhaeng" authorityYear="2021" box="[338,1070,2064,2086]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="105" phylum="Mollusca" rank="species" species="pallgergelyi" status="sp. nov.">
<emphasis id="B97FEA896E22CA29FEE6F7E8BB484E26" box="[338,579,2064,2085]" italics="true" pageId="3" pageNumber="105">Angustopila pallgergelyi</emphasis>
Dumrongrojwattana, Chuenit &amp; Wongkamhaeng
</taxonomicName>
, 
<taxonomicNameLabel id="A24C57F26E22CA29FB8EF7E8BDBB4E25" box="[1082,1200,2064,2086]" pageId="3" pageNumber="105" rank="species">new species</taxonomicNameLabel>
(ZRCBUU 0699). A, front view with scale; B, C, protoconch; D, teleoconch sculpture; E, apertural teeth. Scale bar = 100 µm.
</paragraph>
</caption>
<caption id="DF7466136E25CA2EFFC5FF45B8C446F3" pageId="4" pageNumber="106" startId="4.[113,168,189,211]" targetType="table">
<paragraph id="8BB4369B6E25CA2EFFC5FF45B8C446F3" blockId="4.[113,1451,189,240]" pageId="4" pageNumber="106">
Table 2. The average, minimum value (min), a maximum value (max), a standard deviation of a set of values (stdev) for 
<taxonomicName id="4C0B4D186E25CA2EFA83FF45B9EE46EC" authorityName="Dumrongrojwattana &amp;  &amp; Wongkamhaeng &amp; Koraon" authorityYear="2021" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="pallgergelyi" status="sp. nov.">
<emphasis id="B97FEA896E25CA2EFA83FF45B9EE46EC" italics="true" pageId="4" pageNumber="106">Angustopila pallgergelyi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24C57F26E25CA2EFF47FF23B86446F3" box="[243,367,218,240]" pageId="4" pageNumber="106" rank="species">new species</taxonomicNameLabel>
(n = 15).
</paragraph>
</caption>
<paragraph id="8BB4369B6E25CA2EFF34FEDABC6A47D5" pageId="4" pageNumber="106">
<table id="F90BC43B6E2535D5FF34FED9BC7D47D5" box="[128,1398,289,470]" colsContinueFrom="2.[128,1402,261,811]" gridcols="7" gridrows="5" pageId="4" pageNumber="106">
<tr id="353B34D96E2535D5FF34FED9BC7D473A" box="[128,1398,289,313]" gridrow="0" pageId="4" pageNumber="106">
<th id="76EA5DA56E2535D5FF34FED9B9E8473A" box="[128,227,289,313]" gridcol="0" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFF34FEDAB9E8473B" bold="true" box="[128,227,290,312]" pageId="4" pageNumber="106">Specimen</emphasis>
</th>
<th id="76EA5DA56E2535D5FECFFED9B8AE473A" box="[379,421,289,313]" gridcol="1" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFE34FEDAB8AB473B" bold="true" box="[384,416,290,312]" pageId="4" pageNumber="106">SH</emphasis>
</th>
<th id="76EA5DA56E2535D5FD8EFED9BB6F473A" box="[570,612,289,313]" gridcol="2" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFD88FEDABB6A473B" bold="true" box="[572,609,290,312]" pageId="4" pageNumber="106">SW</emphasis>
</th>
<th id="76EA5DA56E2535D5FD5CFED9BA3E473A" box="[744,821,289,313]" gridcol="3" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFD5CFEDBBA06473A" bold="true" box="[744,781,291,313]" pageId="4" pageNumber="106">SW</emphasis>
<emphasis id="B97FEA896E25CA2EFCB9FED9BA1F473A" bold="true" box="[781,788,289,313]" pageId="4" pageNumber="106">/</emphasis>
<emphasis id="B97FEA896E25CA2EFCA1FEDBBA3E473A" bold="true" box="[789,821,291,313]" pageId="4" pageNumber="106">SH</emphasis>
</th>
<th id="76EA5DA56E2535D5FC0CFED9BAE8473A" box="[952,995,289,313]" gridcol="4" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFC08FEDABAEB4734" bold="true" box="[956,992,290,311]" pageId="4" pageNumber="106">AH</emphasis>
</th>
<th id="76EA5DA56E2535D5FBC3FED9BDA9473A" box="[1143,1186,289,313]" gridcol="5" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFBCCFEDABDAA4734" bold="true" box="[1144,1185,290,311]" pageId="4" pageNumber="106">AW</emphasis>
</th>
<th id="76EA5DA56E2535D5FA96FED9BC7D473A" box="[1314,1398,289,313]" gridcol="6" gridrow="0" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFA96FEDBBC40473B" bold="true" box="[1314,1355,291,312]" pageId="4" pageNumber="106">AW</emphasis>
<emphasis id="B97FEA896E25CA2EFAFFFED9BC59473A" bold="true" box="[1355,1362,289,313]" pageId="4" pageNumber="106">/</emphasis>
<emphasis id="B97FEA896E25CA2EFAE6FEDBBC7D473B" bold="true" box="[1362,1398,291,312]" pageId="4" pageNumber="106">AH</emphasis>
</th>
</tr>
<tr id="353B34D96E2535D5FF34FEA2BC7D4771" box="[128,1398,346,370]" gridrow="1" pageId="4" pageNumber="106">
<th id="76EA5DA56E2535D5FF34FEA2B9E84771" box="[128,227,346,370]" gridcol="0" gridrow="1" pageId="4" pageNumber="106">Min</th>
<td id="76EA5DA56E2535D5FECFFEA2B8AE4771" box="[379,421,346,370]" gridcol="1" gridrow="1" pageId="4" pageNumber="106">0.59</td>
<td id="76EA5DA56E2535D5FD8EFEA2BB6F4771" box="[570,612,346,370]" gridcol="2" gridrow="1" pageId="4" pageNumber="106">0.76</td>
<td id="76EA5DA56E2535D5FD5CFEA2BA3E4771" box="[744,821,346,370]" gridcol="3" gridrow="1" pageId="4" pageNumber="106">1.19</td>
<td id="76EA5DA56E2535D5FC0CFEA2BAE84771" box="[952,995,346,370]" gridcol="4" gridrow="1" pageId="4" pageNumber="106">0.26</td>
<td id="76EA5DA56E2535D5FBC3FEA2BDA94771" box="[1143,1186,346,370]" gridcol="5" gridrow="1" pageId="4" pageNumber="106">0.25</td>
<td id="76EA5DA56E2535D5FA96FEA2BC7D4771" box="[1314,1398,346,370]" gridcol="6" gridrow="1" pageId="4" pageNumber="106">0.90</td>
</tr>
<tr id="353B34D96E2535D5FF34FE84BC7D4790" box="[128,1398,380,403]" gridrow="2" pageId="4" pageNumber="106">
<th id="76EA5DA56E2535D5FF34FE84B9E84790" box="[128,227,380,403]" gridcol="0" gridrow="2" pageId="4" pageNumber="106">Max</th>
<td id="76EA5DA56E2535D5FECFFE84B8AE4790" box="[379,421,380,403]" gridcol="1" gridrow="2" pageId="4" pageNumber="106">0.71</td>
<td id="76EA5DA56E2535D5FD8EFE84BB6F4790" box="[570,612,380,403]" gridcol="2" gridrow="2" pageId="4" pageNumber="106">0.85</td>
<td id="76EA5DA56E2535D5FD5CFE84BA3E4790" box="[744,821,380,403]" gridcol="3" gridrow="2" pageId="4" pageNumber="106">1.35</td>
<td id="76EA5DA56E2535D5FC0CFE84BAE84790" box="[952,995,380,403]" gridcol="4" gridrow="2" pageId="4" pageNumber="106">0.31</td>
<td id="76EA5DA56E2535D5FBC3FE84BDA94790" box="[1143,1186,380,403]" gridcol="5" gridrow="2" pageId="4" pageNumber="106">0.31</td>
<td id="76EA5DA56E2535D5FA96FE84BC7D4790" box="[1314,1398,380,403]" gridcol="6" gridrow="2" pageId="4" pageNumber="106">1.08</td>
</tr>
<tr id="353B34D96E2535D5FF34FE65BC7D47B6" box="[128,1398,413,437]" gridrow="3" pageId="4" pageNumber="106">
<th id="76EA5DA56E2535D5FF34FE65B9E847B6" box="[128,227,413,437]" gridcol="0" gridrow="3" pageId="4" pageNumber="106">Average</th>
<td id="76EA5DA56E2535D5FECFFE65B8AE47B6" box="[379,421,413,437]" gridcol="1" gridrow="3" pageId="4" pageNumber="106">0.66</td>
<td id="76EA5DA56E2535D5FD8EFE65BB6F47B6" box="[570,612,413,437]" gridcol="2" gridrow="3" pageId="4" pageNumber="106">0.82</td>
<td id="76EA5DA56E2535D5FD5CFE65BA3E47B6" box="[744,821,413,437]" gridcol="3" gridrow="3" pageId="4" pageNumber="106">1.25</td>
<td id="76EA5DA56E2535D5FC0CFE65BAE847B6" box="[952,995,413,437]" gridcol="4" gridrow="3" pageId="4" pageNumber="106">0.28</td>
<td id="76EA5DA56E2535D5FBC3FE65BDA947B6" box="[1143,1186,413,437]" gridcol="5" gridrow="3" pageId="4" pageNumber="106">0.28</td>
<td id="76EA5DA56E2535D5FA96FE65BC7D47B6" box="[1314,1398,413,437]" gridcol="6" gridrow="3" pageId="4" pageNumber="106">0.98</td>
</tr>
<tr id="353B34D96E2535D5FF34FE47BC7D47D5" box="[128,1398,447,470]" gridrow="4" pageId="4" pageNumber="106">
<th id="76EA5DA56E2535D5FF34FE47B9E847D5" box="[128,227,447,470]" gridcol="0" gridrow="4" pageId="4" pageNumber="106">SD</th>
<td id="76EA5DA56E2535D5FECFFE47B8AE47D5" box="[379,421,447,470]" gridcol="1" gridrow="4" pageId="4" pageNumber="106">0.03</td>
<td id="76EA5DA56E2535D5FD8EFE47BB6F47D5" box="[570,612,447,470]" gridcol="2" gridrow="4" pageId="4" pageNumber="106">0.03</td>
<td id="76EA5DA56E2535D5FD5CFE47BA3E47D5" box="[744,821,447,470]" gridcol="3" gridrow="4" pageId="4" pageNumber="106">0.06</td>
<td id="76EA5DA56E2535D5FC0CFE47BAE847D5" box="[952,995,447,470]" gridcol="4" gridrow="4" pageId="4" pageNumber="106">0.02</td>
<td id="76EA5DA56E2535D5FBC3FE47BDA947D5" box="[1143,1186,447,470]" gridcol="5" gridrow="4" pageId="4" pageNumber="106">0.02</td>
<td id="76EA5DA56E2535D5FA96FE47BC7D47D5" box="[1314,1398,447,470]" gridcol="6" gridrow="4" pageId="4" pageNumber="106">0.06</td>
</tr>
</table>
</paragraph>
<subSubSection id="C31165106E25CA2EFFC6FC7DB8E0425F" pageId="4" pageNumber="106" type="biology_ecology">
<paragraph id="8BB4369B6E25CA2EFFC6FC7DB8E0425F" blockId="4.[114,753,901,1117]" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFFC6FC7DB9D8459E" bold="true" box="[114,211,901,925]" pageId="4" pageNumber="106">Ecology.</emphasis>
Living snails live in the dark and moisture zone of caves, gliding on the wall or hidden among coral-like stalactite (
<figureCitation id="13302A1E6E25CA2EFF57FC3DB85645DE" box="[227,349,965,989]" captionStart="Fig" captionStartId="4.[813,848,1470,1492]" captionTargetBox="[813,1453,549,1455]" captionTargetId="figure-153@4.[813,1453,549,1455]" captionTargetPageId="4" captionText="Fig. 3. A, B, Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species, living snail at Phet Pho Thong Cave, Sa Kaeo, approximately 300 metres from the entrance. Photograph: Rattanwadee Tekavong." pageId="4" pageNumber="106">Fig. 3A, B</figureCitation>
). All 
<typeStatus id="54B088396E25CA2EFE15FC3DB8C445DE" box="[417,463,965,989]" pageId="4" pageNumber="106">type</typeStatus>
specimens were collected from soil sampled from the dark zone of Phet Pho Thong Cave, 
<collectingRegion id="49CFF8796E25CA2EFF01FBFDB81A421F" box="[181,273,1029,1053]" country="Thailand" name="Sa Kaeo" pageId="4" pageNumber="106">Sa Kaeo</collectingRegion>
, approximately 300 metres from the entrance at the 
<typeStatus id="54B088396E25CA2EFF0DFBDDB9EC423E" box="[185,231,1061,1085]" pageId="4" pageNumber="106">type</typeStatus>
locality. We assume that this new species is a cave-dweller, similar to 
<taxonomicName id="4C0B4D186E25CA2EFECFFBBDB8EF425E" authorityName="Panha &amp; Burch" authorityYear="1999" box="[379,484,1093,1117]" class="Gastropoda" family="Gastrocoptidae" genus="Systenostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="tamlod">
<emphasis id="B97FEA896E25CA2EFECFFBBDB8EF425E" box="[379,484,1093,1117]" italics="true" pageId="4" pageNumber="106">A. tamlod</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E25CA2EFFC6FB7DB9F442BF" pageId="4" pageNumber="106" type="distribution">
<paragraph id="8BB4369B6E25CA2EFFC6FB7DB9F442BF" blockId="4.[114,751,1157,1213]" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFFC6FB7DB801429E" bold="true" box="[114,266,1157,1181]" pageId="4" pageNumber="106">Distribution.</emphasis>
The new species is only known from the 
<typeStatus id="54B088396E25CA2EFFC6FB5DB9AB42BE" box="[114,160,1189,1213]" pageId="4" pageNumber="106">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C31165106E25CA2EFFC6FB1DBB8B43FE" pageId="4" pageNumber="106" type="discussion">
<paragraph id="8BB4369B6E25CA2EFFC6FB1DBB8B43FE" blockId="4.[114,753,1253,1533]" pageId="4" pageNumber="106">
<emphasis id="B97FEA896E25CA2EFFC6FB1DB9EB42FE" bold="true" box="[114,224,1253,1277]" pageId="4" pageNumber="106">Remarks.</emphasis>
All specimens were collected inside the cave in 
<collectingRegion id="49CFF8796E25CA2EFFC6FAFDB83F431E" box="[114,308,1285,1309]" country="Thailand" name="Sa Kaeo" pageId="4" pageNumber="106">Sa Kaeo Province</collectingRegion>
, eastern 
<collectingCountry id="F31C760B6E25CA2EFE25FAFDB8F8431E" box="[401,499,1285,1309]" name="Thailand" pageId="4" pageNumber="106">Thailand</collectingCountry>
, about 300 metres from the entrance. Among the previously described species, 
<taxonomicName id="4C0B4D186E25CA2EFD6DFADDB9EA435E" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="dominikae">
<emphasis id="B97FEA896E25CA2EFD6DFADDB9EA435E" italics="true" pageId="4" pageNumber="106">A. dominikae</emphasis>
</taxonomicName>
was the smallest known land snail (
<bibRefCitation id="EF9A4B6A6E25CA2EFDD2FABDB9E0437E" author="Pall-Gergely B &amp; Hunyadi A &amp; Jochum A &amp; Asami T" pageId="4" pageNumber="106" pagination="31 - 64" refId="ref3661" refString="Pall-Gergely B, Hunyadi A, Jochum A &amp; Asami T (2015) Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys, 523: 31 - 64." type="journal article" year="2015">Pall-Gergely et al., 2015</bibRefCitation>
), with the shell height of the 
<typeStatus id="54B088396E25CA2EFDA9FA9DBB71437E" box="[541,634,1381,1405]" pageId="4" pageNumber="106" type="holotype">holotype</typeStatus>
being 
<quantity id="4CF39B7E6E25CA2EFD76FA9DB994439F" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="106" unit="mm" value="0.86">0.86 mm</quantity>
. 
<taxonomicName id="4C0B4D186E25CA2EFF1DFA7DB8B9439E" authorityName="Dumrongrojwattana &amp;  &amp; Wongkamhaeng &amp; Koraon" authorityYear="2021" box="[169,434,1413,1437]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="106" phylum="Mollusca" rank="species" species="pallgergelyi" status="sp. nov.">
<emphasis id="B97FEA896E25CA2EFF1DFA7DB8B9439E" box="[169,434,1413,1437]" italics="true" pageId="4" pageNumber="106">Angustopila pallgergelyi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24C57F26E25CA2EFE74FA7DBB4F439E" box="[448,580,1413,1437]" pageId="4" pageNumber="106" rank="species">new species</taxonomicNameLabel>
, is even smaller (shell height of the 
<typeStatus id="54B088396E25CA2EFEFFFA5DB8A343BE" box="[331,424,1445,1469]" pageId="4" pageNumber="106" type="holotype">holotype</typeStatus>
is 
<quantity id="4CF39B7E6E25CA2EFE79FA5DBB2543BF" box="[461,558,1445,1469]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="4" pageNumber="106" unit="mm" value="0.64">0.64 mm</quantity>
and shell heights of the 
<typeStatus id="54B088396E25CA2EFF08FA3DB82943DE" box="[188,290,1477,1501]" pageId="4" pageNumber="106" type="paratype">paratypes</typeStatus>
<quantity id="4CF39B7E6E25CA2EFE98FA3DB8C643DE" box="[300,461,1477,1501]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.1" metricValueMin="5.8999999999999995" pageId="4" pageNumber="106" unit="mm" value="0.6499999999999999" valueMax="0.71" valueMin="0.59">0.59–0.71 mm</quantity>
), and is therefore now the world’s smallest known terrestrial snail (
<figureCitation id="13302A1E6E25CA2EFD84FA1DBB7843FE" box="[560,627,1509,1533]" captionStart="Fig" captionStartId="6.[113,148,852,874]" captionTargetBox="[113,1451,189,837]" captionTargetId="figure-3@6.[110,1454,189,837]" captionTargetPageId="6" captionText="Fig. 4. Relative shell size of some Thai terrestrial microsnails. A, Boysidia phatangensis Dumrongrojwattana &amp; Assawawattagee, 2018; B, Anthroapiculus pendulus Panha &amp; Burch, 1999; C, Anauchen srakeoensis Panha &amp; Burch, 2004; D, Gyliotrachela tridentate Panha &amp; Burch, 2004; E, Gyliotrachela diamaidi Panha &amp; Burch, 2002; F, Anachen chedi (Panha, 1997); G, Paraboysidia tamphathai Panha &amp; Burch, 2000; H, Krobylos veruwan Panha &amp; Burch, 2004; I, Hypselostoma banmiensis Panha &amp; Burch, 2004; J, Aulacospira khaopratun Dumrongrojwattana &amp; Panha, 2005; K, Acinolaemus colpodon Thompson &amp; Upatham, 1997; L, Clostophis proboscidea (Panha &amp; Burch, 1999); M, Acinolaemus rhamphodon Thompson &amp; Upatham, 1997; N, Angustopila pallgergelyi Dumrongrojwattana, Chuenit &amp; Wongkamhaeng, new species." pageId="4" pageNumber="106">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>