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<mods:title id="031B67C52A5C23E32E582ACC4FC7B82C">New species and records of Coryphiini from China (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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Key to species of
<taxonomicName id="4C0B4D1E004EF70FDB3B1EE2FB56FD65" ID-CoL="8VR7T" authorityName="Jakobson" authorityYear="1908" box="[1117,1254,725,751]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="tribe" tribe="Coryphiini">Coryphiini</taxonomicName>
of
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<keyLead id="30FA95A8004EF70FDC5A1F32FA2CFC97" box="[828,1436,773,797]" pageId="6" pageNumber="597">1 Gular sutures parallel or diverging anteriad. .............. 2</keyLead>
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<keyLead id="30FA95A8004EF70FDC5A1F11FA2CFCB7" box="[828,1436,805,829]" pageId="6" pageNumber="597"> Gular sutures converging anteriad. .............................. 5</keyLead>
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2 Pronotum longer than wide. Body yellow-brown to reddish-brown. Aedeagus as in
<figureCitation id="13302A18004EF70FDBD11F52FAB4FCF7" box="[1207,1284,869,893]" captionStart="Figs 1114" captionStartId="3.[151,188,1479,1498]" captionTargetBox="[171,1416,196,1459]" captionTargetId="figure-13@3.[171,1417,189,1459]" captionTargetPageId="3" captionText="Figs 1114. Boreaphilus japonicus Sharp, 1874 (Figs 1113) and map of distribution of Coryphiini in China (Fig. 14). 11 habitus; 12 labrum, dorsal view.13 aedeagus, parameral view (internal sac is not shown); 14 map: Archaeoboreaphilus paradoxus sp. nov. (black circle), Boreaphilus japonicus (white circles), Caloboreaphilus hammondi Zerche, 1990 (black star), C. concisus sp. nov. (white star), Coryphium taibaiense Li, Li &amp; Zhao, 2007 (black square), C. tangi Li, Li &amp; Zhao, 2007 (white square), Haida argonautarum Smetana, 2003 (black diamond), H. satoi Smetana, 2003 (white diamond). Scale bars: 1.0 mm (Fig. 11); 0.1 mm (Figs 1213)." figureDoi="http://doi.org/10.5281/zenodo.5177786" httpUri="https://zenodo.org/record/5177786/files/figure.png" pageId="6" pageNumber="597">Fig. 13</figureCitation>
. Body length:
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. Habitus as in
<figureCitation id="13302A18004EF70FDBEF1FB2FB62FC17" box="[1161,1234,901,925]" captionStart="Figs 1114" captionStartId="3.[151,188,1479,1498]" captionTargetBox="[171,1416,196,1459]" captionTargetId="figure-13@3.[171,1417,189,1459]" captionTargetPageId="3" captionText="Figs 1114. Boreaphilus japonicus Sharp, 1874 (Figs 1113) and map of distribution of Coryphiini in China (Fig. 14). 11 habitus; 12 labrum, dorsal view.13 aedeagus, parameral view (internal sac is not shown); 14 map: Archaeoboreaphilus paradoxus sp. nov. (black circle), Boreaphilus japonicus (white circles), Caloboreaphilus hammondi Zerche, 1990 (black star), C. concisus sp. nov. (white star), Coryphium taibaiense Li, Li &amp; Zhao, 2007 (black square), C. tangi Li, Li &amp; Zhao, 2007 (white square), Haida argonautarum Smetana, 2003 (black diamond), H. satoi Smetana, 2003 (white diamond). Scale bars: 1.0 mm (Fig. 11); 0.1 mm (Figs 1213)." figureDoi="http://doi.org/10.5281/zenodo.5177786" httpUri="https://zenodo.org/record/5177786/files/figure.png" pageId="6" pageNumber="597">Fig. 11</figureCitation>
. Far Eastern
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,
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(
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,
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(Honshu, Kyushu). ...................... ............................
<taxonomicName id="4C0B4D1E004EF70FDB411FD2FA2CFC77" ID-CoL="MJNS" authority="Sharp, 1874" authorityName="Sharp" authorityYear="1874" box="[1063,1436,997,1021]" class="Insecta" family="Staphylinidae" genus="Boreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B97FEA8F004EF70FDB411FD2FAA9FC77" bold="true" box="[1063,1305,997,1021]" italics="true" pageId="6" pageNumber="597">Boreaphilus japonicus</emphasis>
Sharp, 1874
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<keyLead id="30FA95A8004EF70FDC5A1831FA2CFB97" box="[828,1436,1029,1053]" pageId="6" pageNumber="597"> Pronotum transverse. ..................................................... 3</keyLead>
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3 Lateral margins of pronotum finely crenulated, rounded at widest point. Antennomeres VIX elongate. Aedeagus with narrow, rod-like median lobe (
<figureCitation id="13302A18004EF70FDA281852FA3FFBF7" box="[1358,1423,1125,1149]" captionStart="Figs 110" captionStartId="2.[151,188,1447,1466]" captionTargetBox="[208,1397,200,1416]" captionTargetId="figure-13@2.[180,1408,189,1431]" captionTargetPageId="2" captionText="Figs 110. Archaeoboreaphilus paradoxus sp. nov. 1 habitus (holotype); 2 labrum, dorsal view; 3 left mandible, dorsal view; 4 labium and mentum, ventral view; 5 left maxilla, ventral view; 6 aedeagus, parameral view; 7 aedeagus, lateral view; 8 abdominal tergite VIII, dorsal view; 9 abdominal sternite IX, ventral view; 10 abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 210)." figureDoi="http://doi.org/10.5281/zenodo.5177784" httpUri="https://zenodo.org/record/5177784/files/figure.png" pageId="6" pageNumber="597">Fig. 6</figureCitation>
). Body brown. Body smaller:
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. Habitus as in
<figureCitation id="13302A18004EF70FDA1218B2FCD8FB37" captionStart="Figs 110" captionStartId="2.[151,188,1447,1466]" captionTargetBox="[208,1397,200,1416]" captionTargetId="figure-13@2.[180,1408,189,1431]" captionTargetPageId="2" captionText="Figs 110. Archaeoboreaphilus paradoxus sp. nov. 1 habitus (holotype); 2 labrum, dorsal view; 3 left mandible, dorsal view; 4 labium and mentum, ventral view; 5 left maxilla, ventral view; 6 aedeagus, parameral view; 7 aedeagus, lateral view; 8 abdominal tergite VIII, dorsal view; 9 abdominal sternite IX, ventral view; 10 abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 210)." figureDoi="http://doi.org/10.5281/zenodo.5177784" httpUri="https://zenodo.org/record/5177784/files/figure.png" pageId="6" pageNumber="597">Fig. 1</figureCitation>
.
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(
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). ............................................................ ...................
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<taxonomicName id="4C0B4D1E004EF70FDC9718F2FAF3FB57" ID-CoL="8XV2B" authority="Shavrin &amp; Smetana, 2020" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[1009,1347,1221,1245]" class="Insecta" family="Staphylinidae" genus="Archaeoboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="paradoxus" status="sp. nov.">
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<keyLead id="30FA95A8004EF70FDC5A18D1FA2CFAF7" pageId="6" pageNumber="597"> Lateral margins of pronotum coarsely crenulated, sometimes angulate at widest part. Antennomeres VIX short. Aedeagus with median lobe different, not rodlike. Body reddish-brown, with darker abdomen. Body larger. ................................................................................ 4</keyLead>
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4 Head about as broad as pronotum. Lateral margins of pronotum at widest point angulate; crenulation of lateral margins moderately fine. Apical portion of medi- an lobe wide, asymmetrical; apical portion of internal sac with several very long spines (Fig.
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). Body length:
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. Habitus as in Fig.
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.
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). ................................... ................
<taxonomicName id="4C0B4D1E004EF70FDCBA1A52FA2CF9F7" ID-CoL="PWYV" authority="Zerche, 1990" authorityName="Zerche" authorityYear="1990" box="[988,1436,1637,1661]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="B97FEA8F004EF70FDCBA1A52FABEF9F7" bold="true" box="[988,1294,1637,1661]" italics="true" pageId="6" pageNumber="597">Caloboreaphilus hammondi</emphasis>
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Head slightly broader than pronotum. Lateral margins of pronotum at widest point not angulate; crenulation of lateral margins coarse. Apical portion of median lobe narrow, symmetrical; apical portion of internal sac with wide field of numerous moderately small spines (
<figureCitation id="13302A18004EF70FDCEA1B12FC69F8B7" box="[908,985,1829,1853]" captionStart="Figs 1527" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 1527. Caloboreaphilus concisus sp. nov. 15 habitus (paratype); 16 labrum, dorsal view; 17 labium and mentum, ventral view; 18 left mandible, dorsal view; 19 left maxilla, ventral view; 20 aedeagus, parameral view; 21 aedeagus, lateral view; 22 male abdominal tergite VIII, dorsal view; 23 male abdominal sternite IX, ventral view; 24 male abdominal sternite VIII, ventral view; 25 female abdominal tergite VIII, dorsal view; 26 female abdominal sternite IX; 27 female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 1627)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 20</figureCitation>
). Habitus as in
<figureCitation id="13302A18004EF70FDB181B12FB78F8B7" box="[1150,1224,1829,1853]" captionStart="Figs 1527" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 1527. Caloboreaphilus concisus sp. nov. 15 habitus (paratype); 16 labrum, dorsal view; 17 labium and mentum, ventral view; 18 left mandible, dorsal view; 19 left maxilla, ventral view; 20 aedeagus, parameral view; 21 aedeagus, lateral view; 22 male abdominal tergite VIII, dorsal view; 23 male abdominal sternite IX, ventral view; 24 male abdominal sternite VIII, ventral view; 25 female abdominal tergite VIII, dorsal view; 26 female abdominal sternite IX; 27 female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 1627)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 15</figureCitation>
. Body length:
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.
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(
<collectingRegion id="49CFF87F004EF70FDB751B72FBDEF8D7" box="[1043,1134,1861,1885]" country="China" name="Sichuan" pageId="6" pageNumber="597">Sichuan</collectingRegion>
). .............................................. ..............................
<emphasis id="B97FEA8F004EF70FDB561B52FA2DF8F7" bold="true" box="[1072,1437,1893,1917]" pageId="6" pageNumber="597">
<taxonomicName id="4C0B4D1E004EF70FDB561B52FAF4F8F7" ID-CoL="8XWDK" authority="Shavrin &amp; Smetana, 2020" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[1072,1348,1893,1917]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="concisus" status="sp. nov.">
<emphasis id="B97FEA8F004EF70FDB561B52FAF4F8F7" bold="true" box="[1072,1348,1893,1917]" italics="true" pageId="6" pageNumber="597">Caloboreaphilus concisus</emphasis>
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<taxonomicNameLabel id="A24C57F4004EF70FDA2F1B52FA2DF8F7" box="[1353,1437,1893,1917]" pageId="6" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
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5 Pronotum large, subangulately widened laterally around middle, lateral portions widely explanate. Elytra almost quadrate, with lateral portions explanate and longitudinally impressed in anterior two-thirds. .....................
<taxonomicName id="4C0B4D1E004EF70FDC9B1432FB0EF797" ID-CoL="8VRQ8" authority="Keen, 1897" authorityName="Keen" authorityYear="1897" box="[1021,1214,2053,2077]" class="Insecta" family="Staphylinidae" genus="Haida" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="B97FEA8F004EF70FDC9B1432FBF2F797" bold="true" box="[1021,1090,2053,2077]" italics="true" pageId="6" pageNumber="597">Haida</emphasis>
Keen, 1897
</taxonomicName>
(see
<bibRefCitation id="EF9A4B6C004EF70FDB921432FA25F797" author="SMETANA A." box="[1268,1429,2053,2077]" pageId="6" pageNumber="597" pagination="137 - 143" refId="ref5709" refString="SMETANA A. 2003: Haida argonautarum sp. nov. and Haida satoi sp. nov., the first representatives of the genus Haida Keen, 1897 (Coleoptera, Staphylinidae, Omaliinae, Coryphiini) in the Palaearctic Region. Special Bulletin of the Japanese Society of Coleopterology 6: 137 - 143." type="journal article" year="2003">SMETANA 2003</bibRefCitation>
)
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</paragraph>
<paragraph id="8BB4369D004FF70EDFF11C88FD48FEBC" blockId="7.[151,760,190,534]" pageId="7" pageNumber="598">
<keyLead id="30FA95A8004FF70EDFF11C88FD48FEBC" pageId="7" pageNumber="598"> Pronotum small, evenly rounded laterally, lateral portions not explanate. Elytra elongate, with lateral portions not explanate and not longitudinally impressed in anterior two-thirds. ............................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FF2E38004FF70EDFF11D09FD47FD9C" pageId="7" pageNumber="598">
<paragraph id="8BB4369D004FF70EDFF11D09FD47FE3C" blockId="7.[151,760,190,534]" pageId="7" pageNumber="598">
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6. Antennomeres IVX transverse. Aedeagus as in
<figureCitation id="13302A18004FF70EDDDA1D09FD48FEDC" box="[700,760,318,342]" captionStart="Figs 110" captionStartId="2.[151,188,1447,1466]" captionTargetBox="[208,1397,200,1416]" captionTargetId="figure-13@2.[180,1408,189,1431]" captionTargetPageId="2" captionText="Figs 110. Archaeoboreaphilus paradoxus sp. nov. 1 habitus (holotype); 2 labrum, dorsal view; 3 left mandible, dorsal view; 4 labium and mentum, ventral view; 5 left maxilla, ventral view; 6 aedeagus, parameral view; 7 aedeagus, lateral view; 8 abdominal tergite VIII, dorsal view; 9 abdominal sternite IX, ventral view; 10 abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 210)." figureDoi="http://doi.org/10.5281/zenodo.5177784" httpUri="https://zenodo.org/record/5177784/files/figure.png" pageId="7" pageNumber="598">
Fig.
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</figureCitation>
in
<bibRefCitation id="EF9A4B6C004FF70EDFB41D68FEC1FEFC" author="LI J. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[210,369,350,374]" pageId="7" pageNumber="598" pagination="89 - 93" refId="ref5452" refString="LI J. - W., LI L. - Z. &amp; ZHAO M. - J. 2007: A review on the genus Coryphium Stephens (Coleoptera, Staphylinidae) of China. Mitteilungen aus dem Museum fur Naturkunde Berlin. Deutsche Entomologische Zeitschrift, Neue Folge 54: 89 - 93." type="journal article" year="2007">LI et al. (2007)</bibRefCitation>
. Body smaller:
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. Habitus as in
<figureCitation id="13302A18004FF70EDE741D49FEFDFE1C" box="[274,333,382,406]" captionStart="Figs 110" captionStartId="2.[151,188,1447,1466]" captionTargetBox="[208,1397,200,1416]" captionTargetId="figure-13@2.[180,1408,189,1431]" captionTargetPageId="2" captionText="Figs 110. Archaeoboreaphilus paradoxus sp. nov. 1 habitus (holotype); 2 labrum, dorsal view; 3 left mandible, dorsal view; 4 labium and mentum, ventral view; 5 left maxilla, ventral view; 6 aedeagus, parameral view; 7 aedeagus, lateral view; 8 abdominal tergite VIII, dorsal view; 9 abdominal sternite IX, ventral view; 10 abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 210)." figureDoi="http://doi.org/10.5281/zenodo.5177784" httpUri="https://zenodo.org/record/5177784/files/figure.png" pageId="7" pageNumber="598">
Fig.
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</figureCitation>
in
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.
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(
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). .... ........................
<taxonomicName id="4C0B4D1E004FF70EDE0F1DA9FD47FE3C" ID-CoL="YSGP" authority="Li, Li &amp; Zhao, 2007" authorityName="Li, Li &amp; Zhao" authorityYear="2007" box="[361,759,414,438]" class="Insecta" family="Staphylinidae" genus="Coryphium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="tangi">
<emphasis id="B97FEA8F004FF70EDE0F1DA9FD96FE3C" bold="true" box="[361,550,414,438]" italics="true" pageId="7" pageNumber="598">Coryphium tangi</emphasis>
<bibRefCitation id="EF9A4B6C004FF70EDD4D1DA9FD47FE3C" author="LI J. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[555,759,414,438]" pageId="7" pageNumber="598" pagination="89 - 93" refId="ref5452" refString="LI J. - W., LI L. - Z. &amp; ZHAO M. - J. 2007: A review on the genus Coryphium Stephens (Coleoptera, Staphylinidae) of China. Mitteilungen aus dem Museum fur Naturkunde Berlin. Deutsche Entomologische Zeitschrift, Neue Folge 54: 89 - 93." type="journal article" year="2007">Li, Li &amp; Zhao, 2007</bibRefCitation>
</taxonomicName>
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<paragraph id="8BB4369D004FF70EDFF11D88FD47FD9C" blockId="7.[151,760,190,534]" pageId="7" pageNumber="598">
<keyLead id="30FA95A8004FF70EDFF11D88FD47FD9C" pageId="7" pageNumber="598">
Antennomeres IVX elongate. Aedeagus as in
<figureCitation id="13302A18004FF70EDDCF1D89FD44FE5C" box="[681,756,446,470]" captionStart="Figs 2837" captionStartId="5.[151,188,1593,1612]" captionTargetBox="[153,1431,199,1560]" captionTargetId="figure-13@5.[108,1439,156,1579]" captionTargetPageId="5" captionText="Figs 2837. Coryphium taibaiense Li, Li &amp; Zhao, 2007. 28 habitus (paratype); 29 labrum, dorsal view; 30 labium and mentum, ventral view; 31 left mandible, dorsal view; 32 left maxilla, ventral view; 33 aedeagus, parameral view; 34 aedeagus, lateral view; 35 male abdominal tergite VIII, dorsal view; 36 male abdominal sternite IX, ventral view; 37 male abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 28); 0.1 mm (Figs 2937)." figureDoi="http://doi.org/10.5281/zenodo.5177790" httpUri="https://zenodo.org/record/5177790/files/figure.png" pageId="7" pageNumber="598">Fig. 37</figureCitation>
. Body larger: 2.572.67. Habitus as in
<figureCitation id="13302A18004FF70EDD3C1DE9FD18FE7C" box="[602,680,478,502]" captionStart="Figs 2837" captionStartId="5.[151,188,1593,1612]" captionTargetBox="[153,1431,199,1560]" captionTargetId="figure-13@5.[108,1439,156,1579]" captionTargetPageId="5" captionText="Figs 2837. Coryphium taibaiense Li, Li &amp; Zhao, 2007. 28 habitus (paratype); 29 labrum, dorsal view; 30 labium and mentum, ventral view; 31 left mandible, dorsal view; 32 left maxilla, ventral view; 33 aedeagus, parameral view; 34 aedeagus, lateral view; 35 male abdominal tergite VIII, dorsal view; 36 male abdominal sternite IX, ventral view; 37 male abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 28); 0.1 mm (Figs 2937)." figureDoi="http://doi.org/10.5281/zenodo.5177790" httpUri="https://zenodo.org/record/5177790/files/figure.png" pageId="7" pageNumber="598">Fig. 28</figureCitation>
.
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(
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).
<taxonomicName id="4C0B4D1E004FF70EDE541DC9FD47FD9C" authority="Li, Li &amp; Zhao, 2007" authorityName="Li, Li &amp; Zhao" authorityYear="2007" box="[306,759,510,534]" class="Insecta" family="Staphylinidae" genus="Coryphium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="taibaiense">
<emphasis id="B97FEA8F004FF70EDE541DC9FD95FD9C" bold="true" box="[306,549,510,534]" italics="true" pageId="7" pageNumber="598">Coryphium taibaiense</emphasis>
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